MUSSELpdb | tribe Indochinellini Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe INDOCHINELLINI Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018: cladomics
8 genera

Cladomics Summary

number of species = 33 (33 total)
number of subtaxa (rank: genus) = 8
analyzed in 65 cladograms in 42 publications
maximum number of terminals in an analysis = 29 (mean = 8.02, median = 3)
maximum number of subtaxa (genus) = 8 (mean = 3.14, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 38
maximum number of included terminals = 29, (mean = 13 median = 12)
maximum number of subtaxa (genus) = 8 (mean = 4.66, median = 4)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 34
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 29 (mean = 13.65, median = 12)
maximum number of subtaxa (genus) = 8 (mean = 4.82, median = 4)
proportion of cladograms that support rigorous monophyly = 0.94

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
29	29	28	8	603	56	4	0

                    +-- Parreysiini          
  +-----------------+                        
  |                 +-- Coelaturini          
  |                                          
--+                 +-- Pseudoscabies **        
  |  +--------------+                        
  |  |              +-- Indonaia **             
  +--+                                       
     |  +-------------- Indochinella **         
     |  |                                    
     +--+  +----------- Radiatula **            
        |  |                                 
        +--+        +-- Scabiellus **           
           |  +-----+                        
           |  |     +-- Unionetta_fabagina__2 **
           +--+                              
              |  +----- Harmandia **            
              +--+                           
                 |  +-- Unionetta_fabagina__1 **
                 +--+                        
                    +-- Scabies **

Bolotov et al., 2024 [Publ 4369], fig. 2

tT	bi	riS	oT	roT	roS	iD
29	29	27	8	33	33	4	21

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Leoparreysiini
  |  +--------------------+                 
  |  |                    +-- Parreysiini   
  +--+                                      
     |  +-------------------- Coelaturini   
     |  |                                   
     +--+                 +-- Pseudoscabies ** 
        |  +--------------+                 
        |  |              +-- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Harmandia **     
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Scabies **

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
29	29	27	8	34	33	4	9

  +----------------------------- Parreysia     
  |                                            
--+                          +-- Leoparreysiini
  |  +-----------------------+                 
  |  |                       +-- Lamellidentini
  +--+                                         
     |  +----------------------- Assamnaia     
     |  |                                      
     +--+  +-------------------- Coelaturini   
        |  |                                   
        +--+                 +-- Pseudoscabies ** 
           |  +--------------+                 
           |  |              +-- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
27	27	25	7	34	30	4	23

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Parreysiini   
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
27	27	25	7	34	30	4	13

  +----------------------------- Parreysiini   
--+                                            
  |  +-------------------------- Lamellidentini
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
26	26	24	7	32	28	4	14

  +----------------------------- Parreysiini   
--+                                            
  |  +-------------------------- Lamellidentini
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
26	26	24	7	32	28	4	12

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Parreysiini   
  |  +--------------------+                 
  |  |                    +-- Leoparreysiini
  +--+                                      
     |  +-------------------- Coelaturini   
     +--+                                   
        |  +----------------- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Harmandia **     
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
26	26	23	7	32	28	4	21

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Parreysiini   
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
21	21	19	7	96	20	4	25

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Parreysiini   
  |  +--------------------+                 
  |  |                    +-- Leoparreysiini
  +--+                                      
     |  +-------------------- Coelaturini   
     +--+                                   
        |  +----------------- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Scabies **       
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Harmandia **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
20	20	17	7	91	20	4	25

  +----------------------------- Lamellidentini         
  |                                                     
--+                          +-- Parreysiini            
  |  +-----------------------+                          
  |  |                       +-- Leoparreysiini         
  +--+                                                  
     |  +----------------------- Coelaturini            
     |  |                                               
     +--+  +-------------------- Indonaia **               
        |  |                                            
        +--+  +----------------- Indochinella **           
           |  |                                         
           |  |           +----- Radiatula_mouhoti **      
           +--+  +--------+                             
              |  |        |  +-- Radiatula_myitkyinae **   
              |  |        +--+                          
              +--+           +-- Radiatula_chaudhurii **   
                 |                                      
                 |  +----------- Radiatula_crispisulcata **
                 +--+                                   
                    |  +-------- Scabiellus **             
                    +--+                                
                       |  +----- Harmandia **              
                       +--+                             
                          |  +-- Unionetta **              
                          +--+                          
                             +-- Scabies **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

tT	bi	riS	oT	roT	roS	iD
18	18	16	7	5	5	4	20

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Leoparreysiini
  +--+                                         
     |  +----------------------- Parreysiini   
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

tT	bi	riS	oT	roT	roS	iD
17	17	15	7	11	10	4	24

  +-------------------------------- Lamellidentini       
--+                                                      
  |  +----------------------------- Parreysiini          
  +--+                                                   
     |  +-------------------------- Leoparreysiini       
     +--+                                                
        |  +----------------------- Coelaturini          
        +--+                                             
           |  +-------------------- Indonaia **             
           +--+                                          
              |  +----------------- Indochinella **         
              +--+                                       
                 |  +-------------- Radiatula **            
                 +--+                                    
                    |  +----------- Scabiellus **           
                    +--+                                 
                       |  +-------- Unionetta_fabagina__2 **
                       +--+                              
                          |  +----- Harmandia **            
                          +--+                           
                             |  +-- Unionetta_fabagina__1 **
                             +--+                        
                                +-- Scabies **

Chen et al., 2026 [Publ 4833], fig. 1

tT	bi	riS	oT	roT	roS	iD
14	15	14	6	13	10	4	29

  +-------------------------------- Lamellidens                 
  |                                                             
--+  +----------------------------- Trapezidens                 
  |  |                                                          
  +--+  +-------------------------- Parreysiini                 
     |  |                                                       
     +--+  +----------------------- Leoparreysiini              
        |  |                                                    
        +--+  +-------------------- Coelaturini                 
           |  |                                                 
           +--+                 +-- Indochinella **                
              |  +--------------+                               
              |  |              +-- Indonaia **                    
              +--+                                              
                 |  +-------------- Radiatula **                   
                 |  |                                           
                 +--+  +----------- Pseudodontini               
                    |  |                                        
                    +--+  +-------- Harmandia **                   
                       |  |                                     
                       +--+     +-- Unionetta_xishuangbannaensis **
                          |  +--+                               
                          |  |  +-- Unionetta_fabagina__2 **       
                          +--+                                  
                             |  +-- Unionetta_fabagina__1 **       
                             +--+                               
                                +-- Scabies **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
14	14	10	7	68	0	0	26

  +----------------- Indonaia **    
--+                              
  |  +-------------- Indochinella **
  +--+                           
     |  +----------- Radiatula **   
     +--+                        
        |  +-------- Scabiellus **  
        +--+                     
           |  +----- Scabies **     
           +--+                  
              |  +-- Unionetta **   
              +--+               
                 +-- Harmandia **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	187	22	4	25

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Scabies **       
                 +--+                 
                    +-- Radiatula **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	189	22	4	27

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Scabies **       
                 +--+                 
                    +-- Radiatula **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	39	0	0	23

  +-------- Indonaia **    
--+                     
  |  +----- Indochinella **
  +--+                  
     |  +-- Scabies **     
     +--+               
        +-- Radiatula **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	13	9	4	113	26	4	25

           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella **            
        +--+  +-----+                              
           |  |     |  +-- Scabies **                 
           |  |     +--+                           
           +--+        +-- Radiatula **               
              |                                    
              |  +-------- Indonaia_shurtleffiana **  
              |  |                                 
              +--+  +----- Indonaia_rugosa **         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana **   
                 +--+--+                           
                    |  +-- Indonaia_caerulea **       
                    |                              
                    |  +-- Indonaia_bonneaudii **     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Bolotov et al., 2022 [Publ 4250], fig. S2

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	3	3	2	26

        +-- Parreysiini   
  +-----+                 
  |     +-- Leoparreysiini
--+                       
  |  +----- Indonaia **      
  +--+                    
     |  +-- Indochinella **  
     +--+                 
        +-- Radiatula **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	189	22	4	23

  +----------------- Parreysiini   
  |                                
--+----------------- Lamellidentini
  |                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Radiatula **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	128	18	4	25

  +-------------------------- Lamellidentini    
  |                                             
--+  +----------------------- Parreysiini       
  |  |                                          
  +--+  +-------------------- Leoparreysiini    
     |  |                                       
     +--+  +----------------- Coelaturini       
        |  |                                    
        +--+  +-------------- Indonaia **          
           |  |                                 
           +--+           +-- Indochinella **      
              |  +--------+                     
              |  |        +-- Scabies_crispata **  
              +--+                              
                 |  +-------- Radiatula **         
                 +--+                           
                    |  +----- Scabies_mandarinus **
                    +--+                        
                       |  +-- Scabies_humilis **   
                       +--+                     
                          +-- Scabies_phaselus **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	71	13	4	25

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Radiatula **     
                 +--+                 
                    +-- Scabies **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	128	18	4	25

                       +-- Parreysiini       
  +--------------------+                     
  |                    +-- Lamellidentini    
--+                                          
  |  +-------------------- Leoparreysiini    
  |  |                                       
  +--+  +----------------- Coelaturini       
     |  |                                    
     +--+  +-------------- Indonaia **          
        |  |                                 
        +--+           +-- Indochinella **      
           |  +--------+                     
           |  |        +-- Scabies_crispata **  
           +--+                              
              |  +-------- Radiatula **         
              +--+                           
                 |  +----- Scabies_mandarinus **
                 +--+                        
                    |  +-- Scabies_humilis **   
                    +--+                     
                       +-- Scabies_phaselus **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
9	9	7	3	40	15	3	23

  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Indochinella **  
           +--+                 
              +-- Radiatula **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

tT	bi	riS	oT	roT	roS	iD
8	8	3	4	28	28	2	25

  +-------------- Leoparreysiini      
  |                                   
--+  +----------- Coelaturini         
  |  |                                
  +--+  +-------- Indonaia **            
     |  |                             
     +--+  +----- Indochinella **        
        |  |                          
        +--+  +-- Radiatula_mouhoti **   
           |  |                       
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **
              |                       
              +-- Scabies **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	1	4	28	28	2	27

  +-------- Leoparreysiini      
  |                             
--+  +----- Coelaturini         
  |  |                          
  |  |  +-- Radiatula_mouhoti **   
  +--+  |                       
     |  +-- Radiatula_myitkyinae **
     |  |                       
     |  +-- Radiatula_chaudhurii **
     +--+                       
        +-- Indochinella **        
        |                       
        +-- Indonaia **            
        |                       
        +-- Scabies **

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	12	7	2	26

  +-------- Leoparreysiini
--+                       
  |  +----- Coelaturini   
  +--+                    
     |  +-- Indochinella **  
     +--+                 
        +-- Radiatula **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	86	8	3	29

  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Leoparreysiini      
  +--+                                      
     |  +-------------- Coelaturini         
     +--+                                   
        |  +----------- Indonaia **            
        +--+                                
           |  +-------- Radiatula_myitkyinae **
           +--+                             
              |  +----- Scabies **             
              +--+                          
                 |  +-- Radiatula_mouhoti **   
                 +--+                       
                    +-- Radiatula_chaudhurii **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	86	8	3	27

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Scabies **       
           +--+                 
              +-- Radiatula **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	6	3	26

  +----------------- Lamellidentini
--+                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Harmandia **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	6	3	26

  +----------------- Lamellidentini
--+                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Harmandia **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	4	3	27

  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Indochinella **  
           +--+                 
              +-- Radiatula **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	28	8	3	27

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indochinella **  
        +--+                    
           |  +-- Radiatula **     
           +--+                 
              +-- Scabies **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	37	5	3	27

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indochinella **  
        +--+                    
           |  +-- Radiatula **     
           +--+                 
              +-- Scabies **

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	20	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Coelaturini   
     +--+                    
        |  +-- Radiatula **     
        +--+                 
           +-- Indochinella **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Coelaturini   
     +--+                    
        |  +-- Radiatula **     
        +--+                 
           +-- Indochinella **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	20	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Scabies_crispata **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	43	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

Imsong & Murali, 2023 [Publ 4370], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	2	2	--

  +----- Lamellidentini
--+                    
  |  +-- Indonaia **      
  +--+                 
     +-- Parreysiini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	2	2	--

  +-- Coelaturini   
  |                 
--+-- Leoparreysiini
  |                 
  +-- Radiatula **

Combosch et al., 2017 [Publ 3806], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Scabies_crispata **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Scabies_crispata **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	4	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Unionetta **     
     +--+                 
        +-- Coelaturini

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Scabies_crispata **

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Scabies_crispata **

Dai et al., 2024 [Publ 4386], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Scabies_scobinatus **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

Xiang et al., 2025 [Publ 4433], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	0	0	--

-- Scabies_crispata **

Combosch et al., 2017 [Publ 3806], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Scabies_crispata **

Choi et al., 2020 [Publ 4169], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	2	2	--

  +----- Radiatula **     
--+                    
  |  +-- Lamellidentini
  +--+                 
     +-- Leoparreysiini

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	43	2	2	--

  +-- Coelaturini
  |              
--+-- Indonaia **   
  |              
  +-- Parreysiini

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Scabies_crispata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	80	3	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinella **

Dai et al., 2024 [Publ 4385], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Scabies_crispata **

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +-- Coelaturini
  |              
--+-- Indonaia **   
  |              
  +-- Parreysiini

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Scabies_phaselus **

Dai et al., 2026 [Publ 4757], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

  +-- Scabies **       
--+                 
  +-- Lamellidentini

Dai et al., 2024 [Publ 4385], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	0	0	--

-- Scabies_crispata **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	0	0	--

-- Indonaia_andersoniana **

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