MUSSELpdb | tribe Lamellidentini Modell, 1942 cladomics

	Home \| About \| Database \| Mussel of the Month \| Publications & Presentations


MUSSELpdb
	Introduction Browse d'basing


Taxonomy
	all genera Unionidae Margaritiferidae Hyriidae Etheriidae Mycetopodidae Iridinidae Species & Genus Richness by Taxonomy


Geography
	Summary North America Central America South America Afrotropics North Eurasia East Asia Australasia Species Richness & Endemism by Geography

Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe LAMELLIDENTINI Modell, 1942: cladomics
4 genera

Cladomics Summary

number of species = 21 (23 total)
number of subtaxa (rank: genus) = 4
analyzed in 55 cladograms in 34 publications
maximum number of terminals in an analysis = 17 (mean = 5.31, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.64, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 33
maximum number of included terminals = 17, (mean = 8.18 median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.06, median = 2)
proportion of cladograms that support basic monophyly = 0.91

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 30
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 17 (mean = 8.47, median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.1, median = 2)
proportion of cladograms that support rigorous monophyly = 0.93

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
17	86	91	3	615	67	4	0

  +-------------------------------- Lamellidens **          
  |                                                      
--+                             +-- Balwantia **            
  |  +--------------------------+                        
  |  |                          +-- Trapezidens **          
  +--+                                                   
     |  +-------------------------- Leoparreysiini       
     |  |                                                
     +--+  +----------------------- Pseudomulleria       
        |  |                                             
        |  |                    +-- Parreysiini          
        +--+  +-----------------+                        
           |  |                 +-- Coelaturini          
           |  |                                          
           +--+                 +-- Pseudoscabies        
              |  +--------------+                        
              |  |              +-- Indonaia             
              +--+                                       
                 |  +-------------- Indochinella         
                 |  |                                    
                 +--+  +----------- Radiatula            
                    |  |                                 
                    +--+        +-- Pseudodontini        
                       |  +-----+                        
                       |  |     +-- Unionetta_fabagina__2
                       +--+                              
                          |  +----- Harmandia            
                          +--+                           
                             |  +-- Unionetta_fabagina__1
                             +--+                        
                                +-- Scabies

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
17	17	15	3	46	45	4	96

  +-------------- Parreysia     
  |                             
  |        +----- Assamnaia     
--+  +-----+                    
  |  |     |  +-- Coelaturini   
  |  |     +--+                 
  +--+        +-- Indochinellini
     |                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Lamellidens **   
        +--+                    
           |  +-- Balwantia **     
           +--+                 
              +-- Trapezidens **

Bolotov et al., 2024 [Publ 4369], fig. 2

tT	bi	riS	oT	roT	roS	iD
17	17	15	3	45	45	4	102

     +----- Lamellidens **   
  +--+                    
  |  |  +-- Balwantia **     
  |  +--+                 
  |     +-- Trapezidens **   
--+                       
  |     +-- Leoparreysiini
  |  +--+                 
  |  |  +-- Parreysiini   
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
16	16	14	2	45	41	4	99

  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
16	16	12	2	45	41	4	103

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
14	14	12	2	44	40	4	95

  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
14	14	11	2	44	40	4	94

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
14	14	12	2	44	40	4	103

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
11	11	6	2	189	23	4	102

  +-------- Parreysiini   
  |                       
  |     +-- Trapezidens **   
--+-----+                 
  |     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
11	11	7	2	187	23	4	99

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	10	2	189	23	4	105

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	38	28	2	113	27	4	105

           +-------------- Trapezidens **             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus **  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus **
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus **   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi ** 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus ** 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis **  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis ** 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1 **
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2 **

Bolotov et al., 2022 [Publ 4250], fig. S4

tT	bi	riS	oT	roT	roS	iD
9	9	6	2	0	0	0	101

  +-- Lamellidens **
--+              
  +-- Trapezidens **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	40	15	3	98

        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Konopleva et al., 2020 [Publ 4072], fig. 2

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	3	3	1	102

  +----- Leoparreysiini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Trapezidens **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
8	8	5	2	130	21	4	100

     +----- Parreysiini   
  +--+                    
  |  |  +-- Trapezidens **   
  |  +--+                 
--+     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	103	32	4	101

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	130	21	4	101

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	109	33	4	101

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	76	18	4	101

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	26	6	3	102

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 5

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	0	0	0	102

  +-- Trapezidens **
--+              
  +-- Lamellidens **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	9	3	103

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	9	3	103

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	38	6	3	--

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Sonowal et al., 2021 [Publ 4221], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	10	0	1	14	0	0	112

  +-------------------------- Lamellidens_mainwaringi **
--+                                                  
  |  +----------------------- Lamprotula             
  +--+                                               
     |  +-------------------- Potomida               
     +--+                                            
        |  +----------------- Lampsilini             
        +--+                                         
           |  +-------------- Pleurobemini           
           +--+                                      
              |  +----------- Aculamprotulina        
              +--+                                   
                 |  +-------- Unio_pictorum          
                 +--+                                
                    |  +----- Anodontini             
                    +--+                             
                       |  +-- Unio_ravoisieri        
                       +--+                          
                          +-- Lamellidens_corrianus **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	54	5	3	--

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	21	4	--

           +-- Lamellidens **   
  +--------+                 
  |        +-- Trapezidens **   
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	55	8	3	--

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	55	8	3	--

        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	26	25	4	--

           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Dai et al., 2024 [Publ 4385], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4385], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Lamellidens_jenkinsianus **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	0	0	--

-- Lamellidens_jenkinsianus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	3	3	--

  +-------- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lamellidens_marginalis **

Dai et al., 2026 [Publ 4757], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

  +-- Indochinellini
--+                 
  +-- Trapezidens **

Imsong & Murali, 2023 [Publ 4370], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	2	2	--

  +----- Lamellidens **   
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Parreysiini

Choi et al., 2020 [Publ 4169], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	2	2	--

  +----- Indochinellini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Leoparreysiini

Sonowal et al., 2021 [Publ 4221], fig. 3a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Lamellidens_corrianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	4	3	--

  +-------- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Lamellidens_jenkinsianus **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Lamellidens_corrianus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Lamellidens_jenkinsianus **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lamellidens **
--+              
  +-- Coelaturini

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	2	1	--

  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Sonowal et al., 2021 [Publ 4221], fig. 3b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Lamellidens_mainwaringi **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	0	0	--

-- Lamellidens_jenkinsianus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	140	0	0	--

-- Lamellidens_jenkinsianus **

Araujo et al., 2017 [Publ 3801], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	1	1	--

  +-- Lamellidens **
--+              
  +-- Coelaturini

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	2	1	--

  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Lamellidens_jenkinsianus **

"Making the world a better place, one mollusk at a time."
	The MUSSEL Project — Home Page https://musselpdb.org/. Site developed and maintained by Dan Graf & Kevin Cummings. Funded by The National Science Foundation.