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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe LAMPROTULINI Modell, 1942: cladomics
     4 genera

Cladomics Summary

number of species = 22 (41 total)
number of subtaxa (rank: genus) = 4
analyzed in 195 cladograms in 118 publications
maximum number of terminals in an analysis = 5 (mean = 2.44, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.96, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 127
maximum number of included terminals = 5, (mean = 3.21 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.47, median = 3)
proportion of cladograms that support basic monophyly = 0.89

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 107
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.38, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.66, median = 3)
proportion of cladograms that support rigorous monophyly = 0.91


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Dai et al., 2025 [Publ 4441], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4423443970
           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
  |           +-- Contradentini  
--+                              
  |  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
242024433845
        +-------- Lamellidentini 
     +--+                        
     |  |  +----- Leoparreysiini 
     |  +--+                     
     |     |  +-- Indochinellini 
     |     +--+                  
  +--+        +-- Coelaturini    
  |  |                           
  |  |        +-- Anodontini     
  |  |     +--+                  
  |  |     |  +-- Unionini       
  |  +-----+                     
  |        |  +-- Chamberlainiini
--+        +--+                  
  |           +-- Discomya **       
  |                              
  |        +----- Ctenodesmini   
  |  +-----+                     
  |  |     |  +-- Rectidentini   
  |  |     +--+                  
  |  |        +-- Contradentini  
  +--+                           
     |        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Caudiculatus   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Gonideini      

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
5533151351
  +-------------- Chamberlainiini
  |                              
  |           +-- Contradentini  
--+  +--------+                  
  |  |        +-- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
55331945154
  +----- Lamprotula **  
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
55331945154
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
5533491852
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5533901552
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5533861652
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5160311367515
        +-- Rectidentini             
  +-----+                            
  |     +-- Contradentini            
  |                                  
  |  +----- Pseudodontini            
  |  |                               
  |  |  +-- Sinosolenaia_iridinea    
--+  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  |  |  +-- Leguminaia               
  +--+  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii **         
        |                            
        +-- Lamprotula **               
        |                            
        +-- Lamprotula_caveata **       
        |                            
        +-- Potomida **                 
        |                            
        +-- Pronodularia **             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora    

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
55331925054
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
55031076753
              +-- Rectidentini      
  +-----------+                     
  |           +-- Contradentini     
--+                                 
  |  +----------- Pseudodontini     
  |  |                              
  +--+  +-------- Gonideini         
     |  |                           
     +--+  +----- Chamberlainiini   
        |  |                        
        |  |  +-- Lamprotula_leaii **  
        +--+  |                     
           |  +-- Lamprotula **        
           |  |                     
           +--+-- Lamprotula_caveata **
              |                     
              +-- Potomida **          
              |                     
              +-- Pronodularia **      

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
57332215
  +----- Potomida **      
  |                    
--+  +-- Pronodularia **  
  |  |                 
  |  +-- Lamprotula **    
  +--+                 
     +-- Leguminaia    
     |                 
     +-- Microcondylaea

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
5533151352
  +-------------- Chamberlainiini
  |                              
  |           +-- Contradentini  
--+  +--------+                  
  |  |        +-- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55431162856
           +-- Pseudodontini  
  +--------+                  
  |        +-- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
554326646
  +-------------- Chamberlainiini
  |                              
--+           +-- Gonideini      
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Contradentini  
     +--+                        
        |  +----- Pronodularia **   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5533861652
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
5533232152
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5533901552
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
44231354553
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
442340931
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4423872553
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
442334521
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
4423433872
           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
  |           +-- Contradentini  
--+                              
  |  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Lamprotula **     
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Potomida **       
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
4423201533
                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina
  |  +-----------+                       
  |  |           |  +-- Pseudodon        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha   
  |  |                                   
  +--+        +-------- Sinosolenaia     
     |  +-----+                          
     |  |     |  +----- Lamprotula **       
     |  |     +--+                       
     |  |        |  +-- Potomida **         
     |  |        +--+                    
     +--+           +-- Pronodularia **     
        |                                
        |           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia       

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
4423491551
  +-------------- Chamberlainiini
  |                              
  |           +-- Rectidentini   
--+  +--------+                  
  |  |        +-- Contradentini  
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
442338421
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
44231354553
  +-------------- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Pronodularia **   

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4423691031
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
442336631
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442352621
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
443225423
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
4423413950
  +----------------------- Pseudodontini  
  |                                       
  |                    +-- Rectidentini   
  |  +-----------------+                  
--+  |                 +-- Contradentini  
  |  |                                    
  |  |        +----------- Inversidens    
  |  |  +-----+                           
  +--+  |     |  +-------- Chamberlainiini
     |  |     +--+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     |  |           |  +-- Pronodularia **   
     +--+           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4423691031
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4130312147512
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida **              
  |                         
  +-- Pronodularia **          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
4423413950
  +----------------------- Pseudodontini  
  |                                       
  |                    +-- Rectidentini   
--+  +-----------------+                  
  |  |                 +-- Contradentini  
  |  |                                    
  |  |  +----------------- Inversidens    
  +--+  |                                 
     |  |        +-------- Chamberlainiini
     |  +--------+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     +--+           |  +-- Pronodularia **   
        |           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4423691031
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442210113
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442352920
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4423762353
        +-------- Chamberlainiini
  +-----+                        
  |     |  +----- Lamprotula **     
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Pronodularia **   
  |                              
  |           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia     

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4423872551
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4423841550
  +----- Potomida **    
--+                  
  |  +-- Pronodularia **
  +--+               
     +-- Lamprotula **  

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4423441351
  +-------------- Chamberlainiini
  |                              
  |           +-- Contradentini  
--+  +--------+                  
  |  |        +-- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
442340931
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
44324213
  +-------- Chamberlainia
--+                      
  |  +----- Sinohyriopsis
  +--+                   
     |  +-- Pronodularia ** 
     +--+                
        +-- Lamprotula **   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
442349833
  +-------------- Chamberlainiini
--+                              
  |  +----------- Pseudodontini  
  +--+                           
     |  +-------- Gonideini      
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
331228522
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
331320422
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
331346522
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
331365524
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3313333351
                 +-- Contradentini  
  +--------------+                  
  |              +-- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Lamprotula **     
              +--+                  
                 +-- Pronodularia **   

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
331317222
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
331250622
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
331343522
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
3422343273
  +----------------------------- Schepmaniini       
  |                                                 
  |  +-------------------------- Pseudodontini      
--+  |                                              
  |  |              +----------- Hyriopsis_bialata  
  |  |              |                               
  |  |  +-----------+        +-- Rectidens          
  +--+  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula **         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida **           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus     

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3313732853
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotula **     
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Pronodularia **   

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
331319112
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
331231112
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
331365524
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Obovalis       
     |  +-----+                  
     |  |     +-- Ptychorhynchus 
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3313555573
     +-------- Chamberlainiini
  +--+                        
  |  |  +----- Ctenodesmini   
  |  +--+                     
  |     |  +-- Rectidentini   
  |     +--+                  
--+        +-- Contradentini  
  |                           
  |     +----- Pronodularia **   
  |  +--+                     
  |  |  |  +-- Lamprotula **     
  |  |  +--+                  
  +--+     +-- Potomida **       
     |                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmaniini   
        +--+                  
           +-- Pseudodontini  

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
331365524
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Ptychorhynchus 
     |  +-----+                  
     |  |     +-- Obovalis       
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3312651032
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Lamprotula **     
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3313333351
                 +-- Contradentini  
  +--------------+                  
  |              +-- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Pronodularia **   
              +--+                  
                 +-- Lamprotula **     

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
331233333
  +-- Pronodularia **
--+               
  +-- Potomida **    

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
331231112
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331115003
1

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
3312343272
  +-------------------------- Chamberlainiini
  |                                          
  |                    +----- Ctenodesmini   
--+  +-----------------+                     
  |  |                 |  +-- Rectidentini   
  |  |                 +--+                  
  |  |                    +-- Contradentini  
  +--+                                       
     |                    +-- Schepmaniini   
     |  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+              +----- Inversidens    
        |  +-----------+                     
        |  |           |  +-- Potomida **       
        |  |           +--+                  
        +--+              +-- Lamprotula **     
           |                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus 

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
33139422
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
3312353372
  +-------------- Chamberlainiini
  |                              
  |        +----- Ctenodesmini   
--+  +-----+                     
  |  |     |  +-- Rectidentini   
  |  |     +--+                  
  |  |        +-- Contradentini  
  +--+                           
     |        +-- Potomida **       
     |  +-----+                  
     |  |     +-- Lamprotula **     
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
342118214
  +----------- Chamberlainiini        
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata **     
     +--+                             
        |  +-- Gibbosula              
        +--+                          
           +-- Lamprotula_cornuumlunae **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
331338114
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3313732851
  +-------- Chamberlainiini
--+                        
  |  +----- Potomida **       
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
331338114
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
331310224
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
3313474731
           +-- Schepmaniini 
  +--------+                
  |        +-- Pseudodontini
--+                         
  |  +-------- Gonideini    
  +--+                      
     |  +----- Potomida **     
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Pronodularia ** 

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
313038032413
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana   
  |                 
  +-- Lampsilis     
  |                 
  +-- Cambarunio    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini    

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
331233333
  +-- Pronodularia **
--+               
  +-- Potomida **    

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
331140924
  +-- Lamprotula **
--+             
  +-- Gonideini 

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
331311212
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
331314112
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
331122315
  +-- Potomida ** 
--+            
  +-- Gonideini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
331329422
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3313371142
              +-- Rectidentini 
  +-----------+                
  |           +-- Contradentini
--+                            
  |  +----------- Pseudodontini
  +--+                         
     |  +-------- Gonideini    
     +--+                      
        |  +----- Potomida **     
        +--+                   
           |  +-- Pronodularia ** 
           +--+                
              +-- Lamprotula **   

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
332142925
  +----------------- Chamberlainiini           
  |                                            
  |           +----- Parvasolenaia_rivularis   
--+  +--------+                                
  |  |        |  +-- Parvasolenaia_triangularis
  |  |        +--+                             
  |  |           +-- Ptychorhynchus            
  +--+                                         
     |           +-- Sinosolenaia_carinata     
     |  +--------+                             
     |  |        +-- Lamprotula **                
     +--+                                      
        |  +-------- Sinosolenaia_recognita    
        |  |                                   
        +--+  +----- Sinosolenaia_oleivora__2  
           |  |                                
           +--+  +-- Sinosolenaia_oleivora__1  
              |  |                             
              +--+-- Sinosolenaia_iridinea     
                 |                             
                 +-- Sinosolenaia_oleivora__3  

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
331310222
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
331219002
  +-- Potomida **  
--+             
  +-- Lamprotula **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
331349832
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3313121233
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Potomida **       

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
3312343272
              +-- Schepmaniini   
  +-----------+                  
  |           +-- Pseudodontini  
  |                              
--+  +----------- Gonideini      
  |  |                           
  |  |     +----- Chamberlainiini
  +--+  +--+                     
     |  |  |  +-- Potomida **       
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Ctenodesmini   
        +--+                     
           |  +-- Rectidentini   
           +--+                  
              +-- Contradentini  

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
331228522
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
331328422
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
3422343273
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |  +-------------------------- Pseudodontini             
--+  |                                                     
  |  |                       +-- Ensidens                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi       
  |  |                       |                             
  +--+                       +-- Rectidens                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula **                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida **                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus            

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
3312343272
  +-------------- Chamberlainiini
  |                              
  |        +----- Ctenodesmini   
--+  +-----+                     
  |  |     |  +-- Rectidentini   
  |  |     +--+                  
  |  |        +-- Contradentini  
  +--+                           
     |        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
331319112
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
331335932
  +-------------- Chamberlainiini
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Pronodularia **   

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
360333316
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
--+                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
220221105--
  +-------------- Chamberlainiini
  |                              
--+           +-- Contradentini  
  |  +--------+                  
  |  |        +-- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
220110101--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
22024300--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
22024300--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22023493--
  +-------- Chamberlainiini
  |                        
--+     +-- Lamprotula **     
  |  +--+                  
  |  |  +-- Potomida **       
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
2202711--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
220280664--
  +----------- Pseudodontini
  |                         
--+        +-- Rectidentini 
  |  +-----+                
  |  |     +-- Contradentini
  +--+                      
     |  +----- Gonideini    
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Potomida **     

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
22023852--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Potomida **       

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2201661--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
22023800--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
22023800--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
22021842--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
23401329334
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontini           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesmini           
--+              |     +--+                           
  |              |        +-- Aculamprotula           
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |              +-----+                           
     |              |     +-- Potamilus_alatus        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis           
     +--+              +--+                           
        |                 +-- Gonideini               
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulini              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2902130010
  +-- Aculamprotula  
  |                  
  +-- Lamprotula **     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida **       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini 
  |                  
  +-- Pleurobemini   

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2201421--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2902140010
  +----------------------- Lamprotula **     
--+                                       
  |  +-------------------- Potomida **       
  +--+                                    
     |  +----------------- Lampsilini     
     +--+                                 
        |  +-------------- Pleurobemini   
        +--+                              
           |  +----------- Aculamprotula  
           +--+                           
              |  +-------- Unio_pictorum  
              +--+                        
                 |  +----- Anodontini     
                 +--+                     
                    |  +-- Unio_ravoisieri
                    +--+                  
                       +-- Lamellidentini 

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
22011300--
1

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
22023262--
  +-------- Gonideini      
--+                        
  |  +----- Chamberlainiini
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
2202811--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
2702472137
  +-- Gibbosula    
  |                
  +-- Margaritifera
  |                
  +-- Sinosolenaia 
  |                
--+-- Leguminaia   
  |                
  +-- Gonidea      
  |                
  +-- Lamprotula **   
  |                
  +-- Potomida **     

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
227022525228
  +-- Lamprotula **   
  |                
  +-- Leguminaia   
  |                
--+-- Potomida **     
  |                
  +-- Gonidea      
  |                
  +-- Pseudodontini

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
22021942--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022700--
  +-- Potomida **  
--+             
  +-- Lamprotula **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
2202611--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
22022042--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
22021300--
  +-- Lamprotula **
--+             
  +-- Potomida **  

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
22023162--
           +-- Obovalis       
  +--------+                  
  |        +-- Ptychorhynchus 
  |                           
--+        +-- Inversidens    
  |  +-----+                  
  |  |     +-- Sinosolenaia   
  +--+                        
     |  +----- Chamberlainiini
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22011452--
  +----- Chamberlainiini
--+                     
  |  +-- Lamprotula **     
  +--+                  
     +-- Gonideini      

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
11011511--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1101661--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Potomida_littoralis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011422--
-- Pronodularia_japanensis **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1101511--
-- Potomida **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
110130306--
  +----------------- Chamberlainiini
--+                                 
  |  +-------------- Lamprotula **     
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Pseudodontini  
        +--+                        
           |  +----- Contradentini  
           +--+                     
              |  +-- Ctenodesmini   
              +--+                  
                 +-- Rectidentini   

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101311--
  +-- Chamberlainiini
--+                  
  +-- Pronodularia **   

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1101921--
  +----- Inversidens 
--+                  
  |  +-- Obovalis    
  +--+               
     +-- Pronodularia **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101721--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
110128282--
  +----- Lamprotula **   
--+                   
  |  +-- Rectidentini 
  +--+                
     +-- Contradentini

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1101221--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011922--
  +----- Chamberlainiini
--+                     
  |  +-- Contradentini  
  +--+                  
     +-- Lamprotula **     

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
110156165--
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Pseudodontini  

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1101300--
-- Potomida_littoralis **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011231--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
11011111--
  +-- Potomida ** 
--+            
  +-- Gonideini

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011200--
-- Potomida_littoralis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011721--
-- Pronodularia_japanensis **

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
110156165--
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Lamprotula **     

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012211--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013611--
-- Pronodularia_japanensis **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1101511--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011811--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101611--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Potomida_littoralis **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101500--
-- Potomida_littoralis **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
11011311--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
11011611--
  +-- Potomida ** 
--+            
  +-- Gonideini

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
110131277--
  +-------------- Chamberlainiini
  |                              
--+  +----------- Potomida **       
  |  |                           
  |  |     +----- Gonideini      
  +--+  +--+                     
     |  |  |  +-- Schepmaniini   
     |  |  +--+                  
     +--+     +-- Pseudodontini  
        |                        
        |  +----- Ctenodesmini   
        +--+                     
           |  +-- Rectidentini   
           +--+                  
              +-- Contradentini  

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011500--
-- Lamprotula_leaii **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101311--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101911--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1101911--
  +-- Gonideini
--+            
  +-- Potomida ** 

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012032--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Chamberlainiini

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1101811--
-- Potomida_littoralis **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011132--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Chamberlainiini
     +--+                  
        +-- Pronodularia **   

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
11011532--
  +----- Chamberlainiini
--+                     
  |  +-- Gonideini      
  +--+                  
     +-- Pronodularia **   

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012411--
-- Pronodularia_japanensis **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1101100--
-- Pronodularia_japanensis **

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Lamprotula_leaii **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     
 
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