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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae

tribe LAMPROTULINI Modell, 1942: cladomics
     4 genera

Cladomics Summary

number of species = 22 (41 total)
number of subtaxa (rank: genus) = 4
analyzed in 209 cladograms in 127 publications
maximum number of terminals in an analysis = 8 (mean = 2.5, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.97, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 140
maximum number of included terminals = 8, (mean = 3.24 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.44, median = 3)
proportion of cladograms that support basic monophyly = 0.9

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 115
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 8 (mean = 3.45, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.65, median = 3)
proportion of cladograms that support rigorous monophyly = 0.91


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
887362412760
     +----------------------------- Indopseudodontina         
     |                                                        
     |                          +-- Lannanaia                 
  +--+  +-----------------------+                             
  |  |  |                       +-- Pilsbryoconcha            
  |  |  |                                                     
  |  +--+  +----------------------- Songkhlanaia              
  |     |  |                                                  
  |     |  |                    +-- Nyeinchanconcha           
  |     +--+  +-----------------+                             
  |        |  |                 +-- Sundadontina              
  |        |  |                                               
  |        |  |                 +-- Isannaia                  
  |        +--+              +--+                             
  |           |              |  +-- Namkongnaia               
  |           |  +-----------+                                
  |           |  |           |  +-- Thaiconcha                
  |           |  |           +--+                             
  |           +--+              +-- Bineurus                  
  |              |                                            
  |              |  +-------------- Pseudodon_walpolei        
  |              |  |                                         
  |              +--+  +----------- Pseudodon_vagulus         
  |                 |  |                                      
  |                 |  |     +----- Pseudodon_vondembuschianus
  |                 +--+  +--+                                
--+                    |  |  |  +-- Pseudodon_mekongi         
  |                    |  |  +--+                             
  |                    +--+     +-- Schepmaniini              
  |                       |                                   
  |                       |  +----- Pseudodon_lenyanensis__2  
  |                       +--+                                
  |                          |  +-- Pseudodon_inoscularis     
  |                          +--+                             
  |                             +-- Pseudodon_lenyanensis__1  
  |                                                           
  |           +-------------------- Chamberlainiini           
  |           |                                               
  |        +--+  +----------------- Parvasolenaia             
  |        |  |  |                                            
  |        |  +--+  +-------------- Pseudopostolata           
  |        |     |  |                                         
  |        |     +--+  +----------- Koreosolenaia             
  |        |        |  |                                      
  |        |        +--+        +-- Cosmopseudodon            
  |        |           |  +-----+                             
  |        |           |  |     +-- Guiunio                   
  |        |           +--+                                   
  |        |              |  +----- Ptychorhynchus            
  +--------+              +--+                                
           |                 |  +-- Obovalis                  
           |                 +--+                             
           |                    +-- Postolata                 
           |                                                  
           |              +-------- Caudiculatus              
           |     +--------+                                   
           |     |        |  +----- Lamprotula **                
           |     |        +--+                                
           |     |           |  +-- Pronodularia **              
           |     |           +--+                             
           +-----+              +-- Potomida **                  
                 |                                            
                 |  +-------------- Sinosolenaia_carinata     
                 |  |                                         
                 |  |        +----- Gonidea                   
                 +--+  +-----+                                
                    |  |     |  +-- Microcondylaea            
                    |  |     +--+                             
                    +--+        +-- Leguminaia                
                       |                                      
                       |  +-------- Inversidens               
                       +--+                                   
                          |  +----- Sinosolenaia_recognita    
                          +--+                                
                             |  +-- Sinosolenaia_iridinea     
                             +--+                             
                                +-- Sinosolenaia_oleivora     

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
6653892246
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
66521342046
  +----------- Chamberlainiini
  |                           
--+  +-------- Rectidentini   
  |  |                        
  +--+     +-- Potomida **       
     |  +--+                  
     |  |  +-- Lamprotula **     
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini      

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55431172846
           +-- Pseudodontini  
  +--------+                  
  |        +-- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
5533151346
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
57332218
  +----- Potomida **      
  |                    
--+  +-- Pronodularia **  
  |  |                 
  |  +-- Lamprotula **    
  +--+                 
     +-- Leguminaia    
     |                 
     +-- Microcondylaea

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5533861646
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5160311266420
  +-------- Rectidentini             
  |                                  
  |  +----- Pseudodontini            
  |  |                               
  |  |  +-- Sinosolenaia_iridinea    
--+  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  |  |  +-- Leguminaia               
  +--+  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii **         
        |                            
        +-- Lamprotula **               
        |                            
        +-- Lamprotula_caveata **       
        |                            
        +-- Potomida **                 
        |                            
        +-- Pronodularia **             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora    

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5533861646
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5533901546
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5533901546
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
55331944944
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
55331965044
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
5533151346
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
5533222046
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
55031066648
  +-------------- Rectidentini      
  |                                 
--+  +----------- Pseudodontini     
  |  |                              
  +--+  +-------- Gonideini         
     |  |                           
     +--+  +----- Chamberlainiini   
        |  |                        
        |  |  +-- Lamprotula_leaii **  
        +--+  |                     
           |  +-- Lamprotula **        
           |  |                     
           +--+-- Lamprotula_caveata **
              |                     
              +-- Potomida **          
              |                     
              +-- Pronodularia **      

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
554326647
  +-------------- Chamberlainiini
  |                              
--+           +-- Gonideini      
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Pronodularia **   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
5533481746
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
55331965044
  +----- Lamprotula **  
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4423691037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
442340937
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
4423201535
  +-------------------- Chamberlainiini
  |                                    
  |  +----------------- Pseudodontini  
--+  |                                 
  |  |        +-------- Sinosolenaia   
  |  |  +-----+                        
  +--+  |     |  +----- Lamprotula **     
     |  |     +--+                     
     |  |        |  +-- Potomida **       
     |  |        +--+                  
     +--+           +-- Pronodularia **   
        |                              
        |           +-- Ptychorhynchus 
        |  +--------+                  
        |  |        +-- Parvasolenaia  
        +--+                           
           |  +-------- Inversidens    
           +--+                        
              |  +----- Gonidea        
              +--+                     
                 |  +-- Microcondylaea 
                 +--+                  
                    +-- Leguminaia     

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
4423413947
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |        +----------- Inversidens    
  |  |  +-----+                           
  +--+  |     |  +-------- Chamberlainiini
     |  |     +--+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     |  |           |  +-- Pronodularia **   
     +--+           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
4423811447
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4423441347
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4423691037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4423772445
        +-------- Chamberlainiini
  +-----+                        
  |     |  +----- Lamprotula **     
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Pronodularia **   
  |                              
  |           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia     

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4423691037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
4423433857
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442352627
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
4423423755
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Lamprotula **     
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Potomida **       
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
44231354445
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
442340937
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442352927
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4130312146416
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida **              
  |                         
  +-- Pronodularia **          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4423872547
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4423872547
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
443226426
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
442334527
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442210115
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
442338427
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
442337637
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4423831647
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
4423491547
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
44324215
  +-------- Chamberlainia
--+                      
  |  +----- Sinohyriopsis
  +--+                   
     |  +-- Pronodularia ** 
     +--+                
        +-- Lamprotula **   

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
44231354445
  +-------------- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Pronodularia **   

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
4423443957
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
442350835
  +-------------- Chamberlainiini
--+                              
  |  +----------- Pseudodontini  
  +--+                           
     |  +-------- Gonideini      
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
4423413947
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |  +----------------- Inversidens    
  |  |  |                                 
  +--+  |        +-------- Chamberlainiini
     |  +--------+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     +--+           |  +-- Pronodularia **   
        |           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3313565658
           +-- Chamberlainiini
  +--------+                  
  |        +-- Rectidentini   
  |                           
--+     +----- Pronodularia **   
  |  +--+                     
  |  |  |  +-- Lamprotula **     
  |  |  +--+                  
  +--+     +-- Potomida **       
     |                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmaniini   
        +--+                  
           +-- Pseudodontini  

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
331228526
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3313313148
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Pronodularia **   
              +--+                  
                 +-- Lamprotula **     

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
33139428
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
3422333157
  +----------------------------- Schepmaniini   
  |                                             
--+  +-------------------------- Pseudodontini  
  |  |                                          
  +--+  +----------------------- Rectidentini   
     |  |                                       
     +--+  +-------------------- Inversidens    
        |  |                                    
        |  |              +----- Lamprotula **     
        +--+  +-----------+                     
           |  |           |  +-- Chamberlainiini
           |  |           +--+                  
           +--+              +-- Potomida **       
              |                                 
              |  +-------------- Sinosolenaia   
              |  |                              
              +--+           +-- Gonidea        
                 |  +--------+                  
                 |  |        +-- Microcondylaea 
                 +--+                           
                    |  +-------- Koreosolenaia  
                    +--+                        
                       |  +----- Parvasolenaia  
                       +--+                     
                          |  +-- Postolata      
                          +--+                  
                             +-- Ptychorhynchus 

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
3312333156
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
331338118
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
331250626
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
3313474738
           +-- Schepmaniini 
  +--------+                
  |        +-- Pseudodontini
--+                         
  |  +-------- Gonideini    
  +--+                      
     |  +----- Potomida **     
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Pronodularia ** 

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3313732846
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotula **     
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Pronodularia **   

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
3312333156
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Potomida **       
     |  +-----+                  
     |  |     +-- Lamprotula **     
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3313732848
  +-------- Chamberlainiini
--+                        
  |  +----- Potomida **       
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
331220006
  +-- Potomida **  
--+             
  +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
331143927
  +-- Lamprotula **
--+             
  +-- Gonideini 

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
331320428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
331233336
  +-- Pronodularia **
--+               
  +-- Potomida **    

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
331231116
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
331343528
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
331122316
  +-- Potomida ** 
--+            
  +-- Gonideini

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
331366526
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Ptychorhynchus 
     |  +-----+                  
     |  |     +-- Obovalis       
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3313323248
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Lamprotula **     
              +--+                  
                 +-- Pronodularia **   

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
331310228
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
342118217
  +----------- Chamberlainiini        
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata **     
     +--+                             
        |  +-- Gibbosula              
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
331345528
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
331336938
  +-------------- Chamberlainiini
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Pronodularia **   

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
3312441246
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331115006
1

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
331338118
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
331366526
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
3312333156
  +-------------------------- Chamberlainiini
  |                                          
--+  +----------------------- Rectidentini   
  |  |                                       
  |  |                    +-- Schepmaniini   
  +--+  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+              +----- Inversidens    
        |  +-----------+                     
        |  |           |  +-- Potomida **       
        |  |           +--+                  
        +--+              +-- Lamprotula **     
           |                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus 

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3313121236
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Potomida **       

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
313037931317
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana   
  |                 
  +-- Lampsilis     
  |                 
  +-- Cambarunio    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini    

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
331366526
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Obovalis       
     |  +-----+                  
     |  |     +-- Ptychorhynchus 
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
331319118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
331328428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
331231116
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3312651036
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Lamprotula **     
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
331319118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
3312333156
              +-- Schepmaniini   
  +-----------+                  
  |           +-- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Chamberlainiini
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
331314118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
331311218
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
331329428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
331310226
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
331350838
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
331233336
  +-- Pronodularia **
--+               
  +-- Potomida **    

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
331228526
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
332144927
  +----------------- Chamberlainiini           
  |                                            
  |           +----- Parvasolenaia_rivularis   
--+  +--------+                                
  |  |        |  +-- Parvasolenaia_triangularis
  |  |        +--+                             
  |  |           +-- Ptychorhynchus            
  +--+                                         
     |           +-- Sinosolenaia_carinata     
     |  +--------+                             
     |  |        +-- Lamprotula **                
     +--+                                      
        |  +-------- Sinosolenaia_recognita    
        |  |                                   
        +--+  +----- Sinosolenaia_oleivora__2  
           |  |                                
           +--+  +-- Sinosolenaia_oleivora__1  
              |  |                             
              +--+-- Sinosolenaia_iridinea     
                 |                             
                 +-- Sinosolenaia_oleivora__3  

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
331317228
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3313371138
  +-------------- Rectidentini 
--+                            
  |  +----------- Pseudodontini
  +--+                         
     |  +-------- Gonideini    
     +--+                      
        |  +----- Potomida **     
        +--+                   
           |  +-- Pronodularia ** 
           +--+                
              +-- Lamprotula **   

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3603333110
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
--+                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
3422333157
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
--+                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
  +--+  +----------------------- Rectidentini              
     |  |                                                  
     +--+  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula **                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida **                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus            

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
2202711--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
22012942--
  +----- Lamprotula **     
--+                     
  |  +-- Chamberlainiini
  +--+                  
     +-- Gonideini      

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
2202811--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
22023262--
  +-------- Gonideini      
--+                        
  |  +----- Chamberlainiini
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
22024200--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2201661--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2902130014
  +-- Aculamprotulina
  |                  
  +-- Lamprotula **     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida **       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini 
  |                  
  +-- Pleurobemini   

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
220110101--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
226022424232
  +-- Lamprotula **   
  |                
  +-- Leguminaia   
  |                
--+-- Potomida **     
  |                
  +-- Gonidea      
  |                
  +-- Pseudodontini

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
22012942--
1

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
22011300--
1

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
22021942--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22023593--
  +-------- Chamberlainiini
  |                        
--+     +-- Lamprotula **     
  |  +--+                  
  |  |  +-- Potomida **       
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
22021300--
  +-- Lamprotula **
--+             
  +-- Potomida **  

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
220280663--
  +----------- Pseudodontini
--+                         
  |  +-------- Rectidentini 
  +--+                      
     |  +----- Gonideini    
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Potomida **     

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
220234124--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Potomida **       

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
241024332647
     +----------- Rectidentini   
     |                           
  +--+        +-- Schepmaniini   
  |  |  +-----+                  
  |  |  |     +-- Pseudodontini  
  |  +--+                        
  |     |  +----- Caudiculatus   
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Gonideini      
  |                              
  |     +-------- Lamellidentini 
  |  +--+                        
  |  |  |  +----- Leoparreysiini 
  |  |  +--+                     
  |  |     |  +-- Indochinellini 
  |  |     +--+                  
  +--+        +-- Coelaturini    
     |                           
     |        +-- Anodontini     
     |     +--+                  
     |     |  +-- Unionini       
     +-----+                     
           |  +-- Chamberlainiini
           +--+                  
              +-- Discomya **       

Wu et al., 2026 [Publ 4825], fig. 5

iTtTbiriSoTroTroSiD
22021700--
  +-- Lamprotula **
--+             
  +-- Potomida **  

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
2202611--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2902140014
  +----------------------- Lamprotula **     
--+                                       
  |  +-------------------- Potomida **       
  +--+                                    
     |  +----------------- Lampsilini     
     +--+                                 
        |  +-------------- Pleurobemini   
        +--+                              
           |  +----------- Aculamprotulina
           +--+                           
              |  +-------- Unio_pictorum  
              +--+                        
                 |  +----- Anodontini     
                 +--+                     
                    |  +-- Unio_ravoisieri
                    +--+                  
                       +-- Lamellidentini 

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2201421--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
22024200--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
22023852--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Potomida **       

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
22022042--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
27024721312
  +-- Gibbosula    
  |                
  +-- Margaritifera
  |                
  +-- Sinosolenaia 
  |                
--+-- Leguminaia   
  |                
  +-- Gonidea      
  |                
  +-- Lamprotula **   
  |                
  +-- Potomida **     

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
22023162--
           +-- Obovalis       
  +--------+                  
  |        +-- Ptychorhynchus 
  |                           
--+        +-- Inversidens    
  |  +-----+                  
  |  |     +-- Sinosolenaia   
  +--+                        
     |  +----- Chamberlainiini
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22011452--
  +----- Chamberlainiini
--+                     
  |  +-- Lamprotula **     
  +--+                  
     +-- Gonideini      

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
22021842--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022700--
  +-- Potomida **  
--+             
  +-- Lamprotula **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
23401329339
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontini           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Oxynaiina               
  |              +-----+                              
  |              |     |  +-- Lepidodesmini           
--+              |     +--+                           
  |              |        +-- Aculamprotulina         
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |              +-----+                           
     |              |     +-- Potamilus_alatus        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis           
     +--+              +--+                           
        |                 +-- Gonideini               
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulini              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
220234124--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Potomida **       

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
220221104--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011231--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013711--
-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
11011511--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
11011532--
  +----- Chamberlainiini
--+                     
  |  +-- Gonideini      
  +--+                  
     +-- Pronodularia **   

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Potomida_littoralis **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012500--
-- Potomida_littoralis **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011422--
-- Pronodularia_japanensis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Potomida_littoralis **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011200--
-- Potomida_littoralis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
110156164--
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Lamprotula **     

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011500--
-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Potomida_littoralis **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
110128281--
  +-- Lamprotula **  
--+               
  +-- Rectidentini

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1101661--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011721--
-- Pronodularia_japanensis **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011811--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1101911--
  +-- Gonideini
--+            
  +-- Potomida ** 

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
110130304--
  +----------- Chamberlainiini
--+                           
  |  +-------- Lamprotula **     
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Rectidentini   

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101911--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1101511--
-- Potomida **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
11011311--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101721--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
11011611--
  +-- Potomida ** 
--+            
  +-- Gonideini

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1101221--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012411--
-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
110130265--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Potomida **       
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1101811--
-- Potomida_littoralis **

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011132--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Chamberlainiini
     +--+                  
        +-- Pronodularia **   

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101311--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1101100--
-- Pronodularia_japanensis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101500--
-- Potomida_littoralis **

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Pronodularia_japanensis **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101311--
  +-- Chamberlainiini
--+                  
  +-- Pronodularia **   

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1101300--
-- Potomida_littoralis **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1101921--
  +----- Inversidens 
--+                  
  |  +-- Obovalis    
  +--+               
     +-- Pronodularia **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
11011111--
  +-- Potomida ** 
--+            
  +-- Gonideini

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
110156164--
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Pseudodontini  

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012032--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Chamberlainiini

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012211--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1101511--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101611--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011922--
  +----- Chamberlainiini
--+                     
  |  +-- Rectidentini   
  +--+                  
     +-- Lamprotula **     

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **
 
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