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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae

tribe LAMPROTULINI Modell, 1942: cladomics
     4 genera

Cladomics Summary

number of species = 22 (41 total)
number of subtaxa (rank: genus) = 4
analyzed in 206 cladograms in 125 publications
maximum number of terminals in an analysis = 8 (mean = 2.5, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.97, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 137
maximum number of included terminals = 8, (mean = 3.26 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.45, median = 3)
proportion of cladograms that support basic monophyly = 0.9

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 113
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 8 (mean = 3.48, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.66, median = 3)
proportion of cladograms that support rigorous monophyly = 0.91


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
887362312760
     +----------------------------- Indopseudodontina         
     |                                                        
     |                          +-- Lannanaia                 
  +--+  +-----------------------+                             
  |  |  |                       +-- Pilsbryoconcha            
  |  |  |                                                     
  |  +--+  +----------------------- Songkhlanaia              
  |     |  |                                                  
  |     |  |                    +-- Nyeinchanconcha           
  |     +--+  +-----------------+                             
  |        |  |                 +-- Sundadontina              
  |        |  |                                               
  |        |  |                 +-- Isannaia                  
  |        +--+              +--+                             
  |           |              |  +-- Namkongnaia               
  |           |  +-----------+                                
  |           |  |           |  +-- Thaiconcha                
  |           |  |           +--+                             
  |           +--+              +-- Bineurus                  
  |              |                                            
  |              |  +-------------- Pseudodon_walpolei        
  |              |  |                                         
  |              +--+  +----------- Pseudodon_vagulus         
  |                 |  |                                      
  |                 |  |     +----- Pseudodon_vondembuschianus
  |                 +--+  +--+                                
--+                    |  |  |  +-- Pseudodon_mekongi         
  |                    |  |  +--+                             
  |                    +--+     +-- Schepmaniini              
  |                       |                                   
  |                       |  +----- Pseudodon_lenyanensis__2  
  |                       +--+                                
  |                          |  +-- Pseudodon_inoscularis     
  |                          +--+                             
  |                             +-- Pseudodon_lenyanensis__1  
  |                                                           
  |           +-------------------- Chamberlainiini           
  |           |                                               
  |        +--+  +----------------- Parvasolenaia             
  |        |  |  |                                            
  |        |  +--+  +-------------- Pseudopostolata           
  |        |     |  |                                         
  |        |     +--+  +----------- Koreosolenaia             
  |        |        |  |                                      
  |        |        +--+        +-- Cosmopseudodon            
  |        |           |  +-----+                             
  |        |           |  |     +-- Guiunio                   
  |        |           +--+                                   
  |        |              |  +----- Ptychorhynchus            
  +--------+              +--+                                
           |                 |  +-- Obovalis                  
           |                 +--+                             
           |                    +-- Postolata                 
           |                                                  
           |              +-------- Caudiculatus              
           |     +--------+                                   
           |     |        |  +----- Lamprotula **                
           |     |        +--+                                
           |     |           |  +-- Pronodularia **              
           |     |           +--+                             
           +-----+              +-- Potomida **                  
                 |                                            
                 |  +-------------- Sinosolenaia_carinata     
                 |  |                                         
                 |  |        +----- Gonidea                   
                 +--+  +-----+                                
                    |  |     |  +-- Microcondylaea            
                    |  |     +--+                             
                    +--+        +-- Leguminaia                
                       |                                      
                       |  +-------- Inversidens               
                       +--+                                   
                          |  +----- Sinosolenaia_recognita    
                          +--+                                
                             |  +-- Sinosolenaia_iridinea     
                             +--+                             
                                +-- Sinosolenaia_oleivora     

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
66521352046
  +----------- Chamberlainiini
  |                           
--+  +-------- Rectidentini   
  |  |                        
  +--+     +-- Potomida **       
     |  +--+                  
     |  |  +-- Lamprotula **     
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini      

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
6653892246
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5533911546
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
5533222046
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5533871646
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
55331955044
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
5533151346
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55431172846
           +-- Pseudodontini  
  +--------+                  
  |        +-- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
55331934944
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
55031066648
  +-------------- Rectidentini      
  |                                 
--+  +----------- Pseudodontini     
  |  |                              
  +--+  +-------- Gonideini         
     |  |                           
     +--+  +----- Chamberlainiini   
        |  |                        
        |  |  +-- Lamprotula_leaii **  
        +--+  |                     
           |  +-- Lamprotula **        
           |  |                     
           +--+-- Lamprotula_caveata **
              |                     
              +-- Potomida **          
              |                     
              +-- Pronodularia **      

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5533911546
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
57332218
  +----- Potomida **      
  |                    
--+  +-- Pronodularia **  
  |  |                 
  |  +-- Lamprotula **    
  +--+                 
     +-- Leguminaia    
     |                 
     +-- Microcondylaea

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
55331955044
  +----- Lamprotula **  
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5533871646
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
5533151346
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
554326647
  +-------------- Chamberlainiini
  |                              
--+           +-- Gonideini      
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Pronodularia **   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
5533481746
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5160311266420
  +-------- Rectidentini             
  |                                  
  |  +----- Pseudodontini            
  |  |                               
  |  |  +-- Sinosolenaia_iridinea    
--+  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  |  |  +-- Leguminaia               
  +--+  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii **         
        |                            
        +-- Lamprotula **               
        |                            
        +-- Lamprotula_caveata **       
        |                            
        +-- Potomida **                 
        |                            
        +-- Pronodularia **             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora    

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
442340937
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
4423433857
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
4423423755
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Lamprotula **     
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Potomida **       
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
4423201535
  +-------------------- Chamberlainiini
  |                                    
  |  +----------------- Pseudodontini  
--+  |                                 
  |  |        +-------- Sinosolenaia   
  |  |  +-----+                        
  +--+  |     |  +----- Lamprotula **     
     |  |     +--+                     
     |  |        |  +-- Potomida **       
     |  |        +--+                  
     +--+           +-- Pronodularia **   
        |                              
        |           +-- Ptychorhynchus 
        |  +--------+                  
        |  |        +-- Parvasolenaia  
        +--+                           
           |  +-------- Inversidens    
           +--+                        
              |  +----- Gonidea        
              +--+                     
                 |  +-- Microcondylaea 
                 +--+                  
                    +-- Leguminaia     

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442352927
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4423701037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
44231344445
  +-------------- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Pronodularia **   

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4423701037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4130312046416
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida **              
  |                         
  +-- Pronodularia **          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
442340937
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442352627
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
443226426
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
4423443957
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
442350835
  +-------------- Chamberlainiini
--+                              
  |  +----------- Pseudodontini  
  +--+                           
     |  +-------- Gonideini      
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
4423413947
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |  +----------------- Inversidens    
  |  |  |                                 
  +--+  |        +-------- Chamberlainiini
     |  +--------+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     +--+           |  +-- Pronodularia **   
        |           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4423872547
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **     

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
442334527
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
4423413947
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |        +----------- Inversidens    
  |  |  +-----+                           
  +--+  |     |  +-------- Chamberlainiini
     |  |     +--+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     |  |           |  +-- Pronodularia **   
     +--+           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis       

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
442338427
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4423701037
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
4423491547
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
442338637
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4423762445
        +-------- Chamberlainiini
  +-----+                        
  |     |  +----- Lamprotula **     
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Pronodularia **   
  |                              
  |           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia     

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
44324215
  +-------- Chamberlainia
--+                      
  |  +----- Sinohyriopsis
  +--+                   
     |  +-- Pronodularia ** 
     +--+                
        +-- Lamprotula **   

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4423841647
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
44231344445
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442210115
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
4423821447
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4423441347
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **     

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4423872547
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3313323248
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Lamprotula **     
              +--+                  
                 +-- Pronodularia **   

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
331346528
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3313732848
  +-------- Chamberlainiini
--+                        
  |  +----- Potomida **       
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
331233336
  +-- Pronodularia **
--+               
  +-- Potomida **    

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
331250626
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
331365526
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
331319118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
313037931317
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana   
  |                 
  +-- Lampsilis     
  |                 
  +-- Cambarunio    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini    

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
331231116
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
331343528
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
331233336
  +-- Pronodularia **
--+               
  +-- Potomida **    

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
331231116
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
331365526
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Ptychorhynchus 
     |  +-----+                  
     |  |     +-- Obovalis       
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
331336938
  +-------------- Chamberlainiini
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Pronodularia **   

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331115006
1

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
331317228
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3603333110
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
--+                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
331338118
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
3312333156
              +-- Schepmaniini   
  +-----------+                  
  |           +-- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Chamberlainiini
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
331310226
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **   

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
331228526
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3313313148
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Pronodularia **   
              +--+                  
                 +-- Lamprotula **     

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
331310228
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
331311218
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
332144927
  +----------------- Chamberlainiini           
  |                                            
  |           +----- Parvasolenaia_rivularis   
--+  +--------+                                
  |  |        |  +-- Parvasolenaia_triangularis
  |  |        +--+                             
  |  |           +-- Ptychorhynchus            
  +--+                                         
     |           +-- Sinosolenaia_carinata     
     |  +--------+                             
     |  |        +-- Lamprotula **                
     +--+                                      
        |  +-------- Sinosolenaia_recognita    
        |  |                                   
        +--+  +----- Sinosolenaia_oleivora__2  
           |  |                                
           +--+  +-- Sinosolenaia_oleivora__1  
              |  |                             
              +--+-- Sinosolenaia_iridinea     
                 |                             
                 +-- Sinosolenaia_oleivora__3  

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
331319118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
33139428
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
331329428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3313371138
  +-------------- Rectidentini 
--+                            
  |  +----------- Pseudodontini
  +--+                         
     |  +-------- Gonideini    
     +--+                      
        |  +----- Potomida **     
        +--+                   
           |  +-- Pronodularia ** 
           +--+                
              +-- Lamprotula **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3313732846
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotula **     
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Pronodularia **   

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
3312441246
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
331365526
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Obovalis       
     |  +-----+                  
     |  |     +-- Ptychorhynchus 
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
3312333156
  +-------------------------- Chamberlainiini
  |                                          
--+  +----------------------- Rectidentini   
  |  |                                       
  |  |                    +-- Schepmaniini   
  +--+  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+              +----- Inversidens    
        |  +-----------+                     
        |  |           |  +-- Potomida **       
        |  |           +--+                  
        +--+              +-- Lamprotula **     
           |                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus 

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3313565658
           +-- Chamberlainiini
  +--------+                  
  |        +-- Rectidentini   
  |                           
--+     +----- Pronodularia **   
  |  +--+                     
  |  |  |  +-- Lamprotula **     
  |  |  +--+                  
  +--+     +-- Potomida **       
     |                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmaniini   
        +--+                  
           +-- Pseudodontini  

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
331122316
  +-- Potomida ** 
--+            
  +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3312661036
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Lamprotula **     
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
3422333157
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
--+                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
  +--+  +----------------------- Rectidentini              
     |  |                                                  
     +--+  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula **                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida **                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus            

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
342118217
  +----------- Chamberlainiini        
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata **     
     +--+                             
        |  +-- Gibbosula              
        +--+                          
           +-- Lamprotula_cornuumlunae **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
331338118
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **    

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
331220006
  +-- Potomida **  
--+             
  +-- Lamprotula **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
3313474738
           +-- Schepmaniini 
  +--------+                
  |        +-- Pseudodontini
--+                         
  |  +-------- Gonideini    
  +--+                      
     |  +----- Potomida **     
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Pronodularia ** 

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
331350838
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
3312333156
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **     

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
3422333157
  +----------------------------- Schepmaniini   
  |                                             
--+  +-------------------------- Pseudodontini  
  |  |                                          
  +--+  +----------------------- Rectidentini   
     |  |                                       
     +--+  +-------------------- Inversidens    
        |  |                                    
        |  |              +----- Lamprotula **     
        +--+  +-----------+                     
           |  |           |  +-- Chamberlainiini
           |  |           +--+                  
           +--+              +-- Potomida **       
              |                                 
              |  +-------------- Sinosolenaia   
              |  |                              
              +--+           +-- Gonidea        
                 |  +--------+                  
                 |  |        +-- Microcondylaea 
                 +--+                           
                    |  +-------- Koreosolenaia  
                    +--+                        
                       |  +----- Parvasolenaia  
                       +--+                     
                          |  +-- Postolata      
                          +--+                  
                             +-- Ptychorhynchus 

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
331320428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3313121236
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Potomida **       

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
331228526
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **     

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
331142927
  +-- Lamprotula **
--+             
  +-- Gonideini 

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
331314118
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
331328428
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **   

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
3312333156
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Potomida **       
     |  +-----+                  
     |  |     +-- Lamprotula **     
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
220221104--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **       

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
220280663--
  +----------- Pseudodontini
--+                         
  |  +-------- Rectidentini 
  +--+                      
     |  +----- Gonideini    
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Potomida **     

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2201661--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
2202611--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
22021300--
  +-- Lamprotula **
--+             
  +-- Potomida **  

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2902130014
  +-- Aculamprotulina
  |                  
  +-- Lamprotula **     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida **       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini 
  |                  
  +-- Pleurobemini   

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
22021842--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2201421--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
27024721312
  +-- Gibbosula    
  |                
  +-- Margaritifera
  |                
  +-- Sinosolenaia 
  |                
--+-- Leguminaia   
  |                
  +-- Gonidea      
  |                
  +-- Lamprotula **   
  |                
  +-- Potomida **     

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
220110101--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
241024332647
     +----------- Rectidentini   
     |                           
  +--+        +-- Schepmaniini   
  |  |  +-----+                  
  |  |  |     +-- Pseudodontini  
  |  +--+                        
  |     |  +----- Caudiculatus   
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Gonideini      
  |                              
  |     +-------- Lamellidentini 
  |  +--+                        
  |  |  |  +----- Leoparreysiini 
  |  |  +--+                     
  |  |     |  +-- Indochinellini 
  |  |     +--+                  
  +--+        +-- Coelaturini    
     |                           
     |        +-- Anodontini     
     |     +--+                  
     |     |  +-- Unionini       
     +-----+                     
           |  +-- Chamberlainiini
           +--+                  
              +-- Discomya **       

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
22024300--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22023593--
  +-------- Chamberlainiini
  |                        
--+     +-- Lamprotula **     
  |  +--+                  
  |  |  +-- Potomida **       
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini      

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
22023852--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Potomida **       

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
2202711--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
22023900--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
23401329339
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontini           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Oxynaiina               
  |              +-----+                              
  |              |     |  +-- Lepidodesmini           
--+              |     +--+                           
  |              |        +-- Aculamprotulina         
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |              +-----+                           
     |              |     +-- Potamilus_alatus        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis           
     +--+              +--+                           
        |                 +-- Gonideini               
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulini              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022700--
  +-- Potomida **  
--+             
  +-- Lamprotula **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
22021763--
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini  

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
22012942--
  +----- Lamprotula **     
--+                     
  |  +-- Chamberlainiini
  +--+                  
     +-- Gonideini      

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
22024300--
  +-- Pronodularia **
--+               
  +-- Lamprotula **  

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
22022042--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **     

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
22011300--
1

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
22021942--
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
22023262--
  +-------- Gonideini      
--+                        
  |  +----- Chamberlainiini
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **   

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
2202811--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2902140014
  +----------------------- Lamprotula **     
--+                                       
  |  +-------------------- Potomida **       
  +--+                                    
     |  +----------------- Lampsilini     
     +--+                                 
        |  +-------------- Pleurobemini   
        +--+                              
           |  +----------- Aculamprotulina
           +--+                           
              |  +-------- Unio_pictorum  
              +--+                        
                 |  +----- Anodontini     
                 +--+                     
                    |  +-- Unio_ravoisieri
                    +--+                  
                       +-- Lamellidentini 

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22011452--
  +----- Chamberlainiini
--+                     
  |  +-- Lamprotula **     
  +--+                  
     +-- Gonideini      

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
22023162--
           +-- Obovalis       
  +--------+                  
  |        +-- Ptychorhynchus 
  |                           
--+        +-- Inversidens    
  |  +-----+                  
  |  |     +-- Sinosolenaia   
  +--+                        
     |  +----- Chamberlainiini
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **     

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
220210022--
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
22012942--
1

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
226022424232
  +-- Lamprotula **   
  |                
  +-- Leguminaia   
  |                
--+-- Potomida **     
  |                
  +-- Gonidea      
  |                
  +-- Pseudodontini

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
110130265--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Potomida **       
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini  

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101611--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101721--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1101511--
-- Potomida **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011721--
-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101911--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
11011311--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
11011111--
  +-- Potomida ** 
--+            
  +-- Gonideini

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101500--
-- Potomida_littoralis **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Lamprotula_leaii **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012032--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Chamberlainiini

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1101811--
-- Potomida_littoralis **

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Pronodularia_japanensis **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Potomida_littoralis **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011132--
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Chamberlainiini
     +--+                  
        +-- Pronodularia **   

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1101100--
-- Pronodularia_japanensis **

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1101300--
-- Potomida_littoralis **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011922--
  +----- Chamberlainiini
--+                     
  |  +-- Rectidentini   
  +--+                  
     +-- Lamprotula **     

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101311--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
11011111--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101311--
  +-- Chamberlainiini
--+                  
  +-- Pronodularia **   

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1101911--
  +-- Gonideini
--+            
  +-- Potomida ** 

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012411--
-- Pronodularia_japanensis **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
110130304--
  +----------- Chamberlainiini
--+                           
  |  +-------- Lamprotula **     
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Rectidentini   

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1101921--
  +----- Inversidens 
--+                  
  |  +-- Obovalis    
  +--+               
     +-- Pronodularia **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011200--
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1101511--
  +-- Gonideini   
--+               
  +-- Pronodularia **

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1101221--
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101900--
-- Potomida_littoralis **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Potomida_littoralis **

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Potomida_littoralis **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012311--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
110156164--
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Lamprotula **     

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011231--
  +-- Lamprotula **
--+             
  +-- Gonideini 

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013332--
-- Lamprotula_leaii **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012700--
-- Potomida_littoralis **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012500--
-- Potomida_littoralis **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011500--
-- Lamprotula_leaii **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
11011611--
  +-- Potomida ** 
--+            
  +-- Gonideini

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013711--
-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012442--
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini      

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
110128281--
  +-- Lamprotula **  
--+               
  +-- Rectidentini

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
11011532--
  +----- Chamberlainiini
--+                     
  |  +-- Gonideini      
  +--+                  
     +-- Pronodularia **   

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
11011511--
  +-- Pronodularia **
--+               
  +-- Gonideini   

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1101661--
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Potomida_littoralis **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Potomida_littoralis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011422--
-- Pronodularia_japanensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
110156164--
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Pseudodontini  

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012211--
  +-- Gonideini 
--+             
  +-- Lamprotula **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011811--
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **     
 
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