The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae tribe LAMPSILINI Ihering, 1901: cladomics
32 genera Cladomics Summarynumber of species = 158 (183 total)
number of subtaxa (rank: genus) = 32
analyzed in 282 cladograms in 166 publications
maximum number of terminals in an analysis =
93 (mean = 5.93, median = 3)
maximum number of subtaxa (genus) =
27 (mean = 3.88, median = 3) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 205
maximum number of included terminals = 93, (mean =
7.78 median = 5)
maximum number of subtaxa (genus) =
27 (mean = 4.96, median = 4)
proportion of cladograms that support basic monophyly
= 0.78 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 176
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 93 (mean = 8.1, median = 5)
maximum number of subtaxa (genus) =
27 (mean = 5.34, median = 4)
proportion of cladograms that support rigorous monophyly = 0.74
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 93 | 97 | 113 | 27 | 539 | 98 | 4 | 0 | +-- Quadrulini
+--------------------------------------------------------------------------+
| +-- Pleurobemini
|
| +----- Amblema
--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias
| | +--+
| | +-- Psoronaias
| |
+--+ +----------------------------------------------------------------------- Reginaia
| |
| | +----- Popenaias
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias **
+--+ | +--+
| | +-- Glebula **
| |
| | +----------------------------------------------------------------- Paetulunio **
| | |
+--+ | +----------------- Toxolasma_parvum **
| | |
| | | +----- Toxolasma_lividum__2 **
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus **
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1 **
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2 **
| | | +--+
| | | | | +----- Toxolasma_glans **
| | | | +--+
| | | | | +-- Toxolasma_paulum **
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1 **
| | |
| | | +-------- Toxolasma__4 **
| | +--+
+--+ | +----- Toxolasma_granulatum **
| +--+
| | +-- Toxolasma__2 **
| +--+
| +-- Toxolasma__3 **
|
| +----- Toxolasma_pullus **
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum **
| | +--+
| | +-- Toxolasma__1 **
| |
| | +-- Obliquaria **
| | +-----------------------------------------------------+
| | | +-- Plectomerus
+--+ |
| | +-------- Potamilus_inflatus **
| | +--------+
| | | | +----- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni **
| | | |
+--+ | | +----- Potamilus_fragilis__2 **
| | | +--------+
| | | | | +-- Potamilus_leptodon **
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1 **
| | |
| | | +----------- Potamilus_metnecktayi **
| | | |
| | +--+ +-------- Potamilus_alatus__3 **
| | | |
| | +--+ +----- Potamilus_alatus__2 **
+--+ | |
| +--+----- Potamilus_purpuratus__2 **
| |
| | +-- Potamilus_alatus__1 **
| +--+
| +-- Potamilus_purpuratus__1 **
|
| +-- Villosa_villosa **
| +-----------------------------------------------+
| | +-- Atlanticoncha **
| |
| | +-- Cambarunio_nebulosus **
| | +--------------------------------------------+
+--+ | +-- Ligumia **
| |
| | +-------- Leaunio_umbrans **
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus **
+--+ | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Cambarunio_simus **
| |
| | +----------------------------------------- Hamiota **
| | |
+--+ | +----------------------- Pachynaias **
| | |
| | +--------------+ +-------------------- Sagittunio **
| | | | |
| | | | | +----- Ellipsaria **
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax **
| | | | | +--+
| | | | | +-- Truncilla **
+--+ | +--+
| | | +----- Medionidus_acutissimus **
| | | +--------+
| | | | | +-- Lemiox **
| | | | +--+
| | +--+ +-- Medionidus_penicillatus **
| | |
| | | +----------- Ptychobranchus **
| | | |
| | +--+ +-- Cyprogenia_aberti__2 **
| | | +-----+
+--+ | | +-- Dromus **
| +--+
| | +----- Cyprogenia_stegaria__2 **
| +--+
| | +-- Cyprogenia_aberti__1 **
| +--+
| +-- Cyprogenia_stegaria__1 **
|
| +-- Villosa_delumbis **
| +--------------------------------+
| | +-- Villosa_vibex **
| |
| | +-------------- Obovaria_olivaria **
| | |
| | +-----------------+ +-- Venustaconcha **
| | | | +--------+
| | | | | +-- Epioblasma **
| | | +--+
| | | | +-------- Obovaria_retusa **
+--+ | +--+
| | | +----- Obovaria_choctawensis **
| | +--+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_subrotunda **
| |
| | +-- Mesonaias **
| | +-----------------------+
| | | +-- Obovaria_arkansasensis **
| | |
+--+ +--+ +-- Lampsilis_splendida **
| | | +--------------------+
| | | | +-- Lampsilis_bracteata **
| | | |
| | +--+ +-- Lampsilis_fasciola **
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----------------- Lampsilis_ornata **
| | | |
| | +--+ +-------------- Lampsilis_ovata__2 **
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis **
| | | |
| | | | +----- Lampsilis_cardium__2 **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1 **
+--+ | | +--+
| +--+ +-- Lampsilis_satura **
| |
| | +----- Lampsilis_cariosa__2 **
| +--+
| | +-- Lampsilis_ovata__1 **
| +--+
| +-- Lampsilis_cariosa__1 **
|
| +----------- Ortmanniana_ligamentina__2 **
| +--------------+
| | | +-------- Lampsilis_abrupta **
| | +--+
| | | +----- Lampsilis_higginsii__2 **
| | +--+
| | | +-- Ortmanniana_ligamentina__1 **
| | +--+
+--+ +-- Lampsilis_higginsii__1 **
|
| +----- Medionidus_conradicus **
| +-----------------+
| | | +-- Leaunio_lienosus **
| | +--+
| | +-- Leaunio_vanuxemensis **
| |
+--+ +-- Lampsilis_floridensis **
| +-----------------+
| | +-- Lampsilis_teres **
| |
| | +-- Lampsilis_siliquoidea__1 **
+--+ |
| +-- Lampsilis_siliquoidea__2 **
| +--------------+
| | +-- Lampsilis_powellii__1 **
| | |
+--+ +-- Lampsilis_powellii__2 **
|
| +-------------- Lampsilis_sietmani **
| |
+--+ +-- Lampsilis_straminea__2 **
| +--------+
| | +-- Lampsilis_virescens **
+--+
| +-------- Lampsilis_hydiana__2 **
+--+
| +----- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
+--+
+-- Lampsilis_straminea__1 **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 48 | 48 | 45 | 23 | 8 | 6 | 3 | 120 | +----- Pleurobemini
+--------------------------------+
| | +-- Amblemini
| +--+
| +-- Popenaiadini
--+
| +-- Cyrtonaias **
| +--------------------------------+
| | +-- Glebula **
| |
+--+ +-------------------------------- Paetulunio **
| |
| | +-- Obliquaria **
+--+ +--------------------------+
| | +-- Toxolasma **
| |
| | +-------------------------- Sagittunio **
+--+ |
| | +----------- Potamilus **
| | |
| | +-----------+ +-- Medionidus_conradicus **
| | | | +-----+
+--+ | | | +-- Ptychobranchus **
| | +--+
| | | +----- Medionidus_acutissimus **
| | | |
| | +--+----- Lemiox **
| | |
| | | +-- Dromus **
+--+ +--+
| +-- Cyprogenia **
|
| +----- Lampsilis_teres **
| +--------------+
| | | +-- Truncilla **
| | +--+
| | +-- Ellipsaria **
| |
+--+ +----------------- Leaunio **
| |
| | +----------- Hamiota **
| | |
| | +--+ +-- Lampsilis_fasciola **
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
+--+ +--+
| +-- Lampsilis_cardium **
|
| +----- Venustaconcha **
| +-----+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +-------- Lampsilis_siliquoidea **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Cambarunio **
+--+
+-- Ligumia **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 48 | 48 | 42 | 23 | 9 | 7 | 3 | 117 | +-------------------------------------------- Pleurobemini
|
| +----------------------------------------- Amblemini
--+ |
| +----------------------------------------- Popenaiadini
| |
| | +-- Paetulunio **
+--+ +-----------------------------------+
| | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +-----------------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
| | +----- Ptychobranchus **
| | +--+
+--+ | | +-- Dromus **
| | +--+
| +-----------------------+ +-- Cyprogenia **
| | |
| | | +----- Medionidus_conradicus **
| | +--+
| | | +-- Medionidus_acutissimus **
| | +--+
+--+ +-- Lemiox **
|
| +----------------------------- Sagittunio **
| |
| +----------------------------- Potamilus **
| |
| | +-- Truncilla **
+--+ +-----------------------+
| | +-- Ellipsaria **
| |
| | +-- Ortmanniana_pectorosa **
| | +-----+
| | | +-- Lampsilis_fasciola **
+--+ +--------------+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lampsilis_cardium **
+--+
| +-- Ligumia **
| +--+
| | +-- Cambarunio **
| +--------------+
| | | +-- Lampsilis_teres **
| | +--+
+--+ +-- Hamiota **
|
| +----------------- Leaunio **
| |
+--+ +-------------- Ortmanniana_ligamentina **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
+--+ +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha **
+--+
+-------- Epioblasma **
|
| +----- Obovaria_unicolor **
+--+
| +-- Obovaria_subrotunda **
+--+
+-- Obovaria_arkansasensis **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 43 | 43 | 46 | 23 | 59 | 47 | 4 | 130 | +----------------------------------------- Quadrulini
|
| +-------------------------------------- Pleurobemini
--+ |
| | +-- Plectomerus
| | +--------------------------------+
+--+ | +-- Reginaia
| |
| | +-- Popenaiadini
| | +-----------------------------+
+--+ | +-- Amblema
| |
| | +-------- Toxolasma **
| | +--------------------+
| | | | +----- Obliquaria **
| | | +--+
+--+ | | +-- Cyrtonaias **
| | +--+
| | +-- Glebula **
| |
| | +-------------- Potamilus **
| | |
| | | +-------- Medionidus_acutissimus **
+--+ | +--+
| +-----------+ | | +----- Lemiox **
| | | | +--+
| | | | | +-- Dromus **
| | | | +--+
| | +--+ +-- Cyprogenia **
| | |
| | | +-- Sagittunio **
| | | +--+
| | | | +-- Medionidus_conradicus **
+--+ +-----+
| | +-- Ptychobranchus **
| +--+
| +-- Venustaconcha_ellipsiformis__2 **
|
| +-- Ellipsaria **
| +--------------------+
| | +-- Truncilla **
| |
| | +-------------------- Villosa **
+--+ |
| | +-- Obovaria_olivaria **
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1 **
| | +-----+
+--+ | | +-------- Epioblasma **
| | +--+
| | | +----- Obovaria_unicolor **
| | +--+
| | | +-- Obovaria_arkansasensis **
| | +--+
| | +-- Obovaria_subrotunda **
+--+
| +-------- Hamiota **
| |
| +-----+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ovata **
| | +--+
| | | +-- Ortmanniana_pectorosa **
+--+ +--+
| +-- Lampsilis_ornata **
|
| +-- Ligumia **
| +--------+
| | +-- Cambarunio **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Leaunio **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 43 | 44 | 37 | 23 | 59 | 47 | 4 | 115 | +-------------------------- Quadrulini
|
| +----------------------- Pleurobemini
--+ |
| | +----- Popenaiadini
| | +--------------+
| | | | +-- Amblema
+--+ | +--+
| | +-- Reginaia
| |
| | +-- Plectomerus
| | +--------------+
+--+ | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +--------+
| | | | +-- Cyrtonaias **
| | | +--+
| | | +-- Glebula **
| | |
+--+ | +----------- Sagittunio **
| | |
| | +----------- Leaunio **
| | |
| | +----------- Potamilus **
| | |
| | +----------- Hamiota **
| | |
| | | +-- Ellipsaria **
+--+ +--------+
| | +-- Truncilla **
| |
| | +----- Lampsilis_teres **
| +-----+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
| | +----- Villosa **
| +-----+
| | | +-- Ligumia **
| | +--+
| | +-- Cambarunio **
| |
+--+ +-------- Ortmanniana_pectorosa **
+--+
| | +----- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha_ellipsiformis__2 **
+--+
| +-------- Epioblasma **
| |
| | +----- Obovaria_unicolor **
| +--+
| | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Ptychobranchus **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1 **
| |
+--+ +----- Medionidus_conradicus **
| |
| | +-- Lemiox **
+--+--+
| +-- Medionidus_acutissimus **
|
| +-- Dromus **
+--+
+-- Cyprogenia **
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 36 | 37 | 36 | 20 | 5 | 5 | 3 | 122 | +-------------------------------------- Quadrulini
|
| +-- Amblema
--+ +--------------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------------- Plectomerus
| |
| | +----------------------------- Toxolasma **
+--+ |
| | +----- Popenaiadini
| | +--------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Medionidus **
| | +--+
| | | +-- Lemiox **
+--+ +--------------+
| | | +-- Cyprogenia **
| | +--+
| | +-- Ptychobranchus **
| |
+--+ +-------- Potamilus **
| +--------+
| | | +----- Atlanticoncha **
| | +--+
| | | +-- Ellipsaria **
| | +--+
+--+ +-- Truncilla **
|
| +-- Hamiota **
| +-----------+
| | +-- Villosa **
| |
+--+ +----------- Ligumia **
| |
| | +----- Venustaconcha **
+--+ +--+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +----- Ortmanniana_pectorosa **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis **
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 27 | 7 | 15 | 14 | 3 | 104 | +-------------------------------------------- Quadrulini
|
--+ +----------------------------------------- Pleurobemini
| |
+--+ +-------------------------------------- Amblemini
| |
+--+ +----------------------------------- Toxolasma **
| |
+--+ +-------------------------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +--------------------------+
| | +-- Truncilla **
| |
| | +-------------------------- Leaunio **
+--+ |
| | +-------------------- Lampsilis_bracteata **
| | |
| | +--+ +----------------- Lampsilis_fasciola **
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata **
| | | |
| | +--+ +-- Lampsilis_ovata **
| | | +--------+
| | | | +-- Lampsilis_cariosa **
| | +--+
| | | +-------- Lampsilis_cardium__3 **
+--+ +--+
| | +----- Lampsilis_cardium__2 **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium__1 **
|
| +-- Lampsilis_higginsii **
| +--------------+
| | +-- Ortmanniana **
+-----+
| +-------------- Lampsilis_teres **
| |
+--+ +-- Lampsilis_radiata **
| +--------+
| | +-- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_bergmanni **
+--+
| +----- Lampsilis_hydiana **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 30 | 18 | 18 | 7 | 7 | 4 | 115 | +-------------------------- Quadrulini
|
| +----------------------- Plectomerus
--+ |
| | +-------------------- Popenaiadini
| | |
| | +-------------------- Pleurobemini
+--+ |
| | +-- Cyrtonaias **
| | +--+
| | | +-- Glebula **
| +--------------+
| | | +-- Reginaia
+--+ +--+
| +-- Obliquaria **
|
| +----------------- Toxolasma **
| |
| | +-------------- Potamilus **
| | |
| | | +-- Ellipsaria **
| | +-----------+
+--+ | +-- Truncilla **
| |
| | +----- Medionidus **
| +--------+
| | | +-- Lemiox **
| | +--+
+--+ +-- Dromus **
|
| +----------- Cambarunio **
| |
| +----------- Leaunio **
| |
| +----------- Villosa **
| |
| | +-- Sagittunio **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha **
+-----+
| | +-- Epioblasma **
| +--+
| +-- Obovaria **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 26 | 7 | 15 | 14 | 3 | 114 | +----------------------------------------------- Quadrulini
|
--+ +-------------------------------------------- Reginaia
| |
+--+ +----------------------------------------- Pleurobemini
| |
+--+ +-------------------------------------- Amblema
| |
+--+ +----------------------------------- Toxolasma **
| |
+--+ +-------------------------------- Ellipsaria **
| |
+--+ +----------------------------- Truncilla **
| |
| | +-------------------------- Potamilus **
+--+ |
| | +----------------------- Leaunio **
| | |
+--+ | +----------------- Lampsilis_bracteata **
| | |
| | +--+ +-- Lampsilis_ornata **
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola **
| | +--+
| | | +----------- Lampsilis_ovata **
| | +--+
| | | +-------- Lampsilis_cardium__3 **
| | +--+
+--+ | +----- Lampsilis_cardium__2 **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium__1 **
|
| +-------------- Lampsilis_teres **
| |
| | +-- Lampsilis_higginsii **
+-----+ +--------+
| | +-- Ortmanniana **
| |
+--+ +-- Lampsilis_radiata **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Smith et al., 2020 [Publ 4129], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 18 | 8 | 8 | 8 | 4 | 119 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Obliquaria **
| |
+--+ +----------- Cyrtonaias **
| |
| | +-- Pachynaias **
+--+ +-----+
| | +-- Lampsilis **
| |
+--+ +-- Ellipsaria **
| +--+
| | +-- Truncilla **
+--+
| +-- Atlanticoncha **
+--+
+-- Potamilus **
Smith et al., 2020 [Publ 4129], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 18 | 8 | 8 | 8 | 4 | 119 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Obliquaria **
| |
+--+ +----------- Cyrtonaias **
| |
| | +-- Pachynaias **
+--+ +-----+
| | +-- Lampsilis **
| |
+--+ +-- Ellipsaria **
| +--+
| | +-- Truncilla **
+--+
| +-- Atlanticoncha **
+--+
+-- Potamilus **
Porto-Hannes et al., 2019 [Publ 3969], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 18 | 13 | 6 | 1 | 1 | 1 | 114 | +-------------------- Obliquaria **
|
+-------------------- Amblemini
|
| +----------------- Lampsilis_teres **
| |
| +----------------- Obovaria **
--+ |
| | +-- Leaunio **
| +--------------+
| | +-- Cambarunio **
| |
| | +-------- Ortmanniana **
| | |
+--+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
Porto-Hannes et al., 2019 [Publ 3969], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 6 | 1 | 1 | 1 | 115 | +-------------------------- Amblemini
|
| +----------------------- Obliquaria **
--+ |
| | +-- Cambarunio **
| | +-----------------+
| | | +-- Leaunio **
+--+ |
| | +----------- Lampsilis_teres **
| | |
| +--------+ +-------- Ortmanniana **
| | | |
| | +--+ +-- Lampsilis_hydiana **
+--+ | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +----------------- Obovaria **
| |
+--+ +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
Grobler et al., 2011 [Publ 3359], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 12 | 10 | 1 | 1 | 1 | 117 | +----------------- Amblemini
|
| +-------- Lemiox **
| +-----+
--+ | | +----- Medionidus **
| | +--+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------- Potamilus **
| |
| | +----- Lampsilis_siliquoidea **
| | +--+
+--+ | | +-- Lampsilis_higginsii **
| | +--+
| | +-- Ortmanniana **
| |
| | +----- Lampsilis_fasciola **
+--+--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-- Cambarunio **
| |
+-----+-- Ligumia **
|
+-- Epioblasma **
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 9 | 10 | 19 | 13 | 3 | 114 | +-------------------- Reginaia
|
--+ +-- Pleurobemini
| +--------------+
| | +-- Quadrulini
+--+
| +-------------- Amblema
| |
+--+ +----------- Toxolasma **
| |
| | +-------- Obliquaria **
+--+ |
| | +----- Obovaria **
| | |
| | +----- Lampsilis_cardium **
+--+ |
| +----- Potamilus **
| |
| | +-- Epioblasma **
| +--+
| | +-- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
| +-- Lampsilis_higginsii **
|
| +-- Ligumia **
+--+
| +-- Lampsilis_teres **
|
| +-- Ellipsaria **
+--+
+-- Truncilla **
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 22 | 18 | 10 | 22 | 13 | 3 | 117 | +-------------------------------- Quadrulina
|
--+ +----------------------------- Megalonaias
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Amblema
| |
| | +-------------------- Pleurobemini
+--+ |
| | +-- Reginaia
| +-----------------+
+--+ +-- Obliquaria **
|
| +----------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +-----------+
| | +-- Truncilla **
| |
+--+ +----------- Lampsilis_cardium **
| |
| | +----- Ligumia **
| +-----+
+--+ | +-- Obovaria **
| +--+
| +-- Epioblasma **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis_higginsii **
Inoue et al., 2020 [Publ 3941], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 2 | 0 | 0 | 0 | 113 | +-- Truncilla **
--+
+-- Lampsilis **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 9 | 11 | 58 | 58 | 4 | 122 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
| | +-- Plectomerus
+--+ +-----------------+
| | +-- Reginaia
| |
| | +-- Popenaiadini
+--+ +--------------+
| | +-- Amblema
| |
| | +-------------- Potamilus_alatus **
| | |
+--+ +-------------- Potamilus_capax **
| |
| +-------------- Glebula **
| |
| | +-- Toxolasma **
| +-----------+
+--+ +-- Paetulunio **
|
| +-- Dromus **
+-----------+
| +-- Lemiox **
|
| +-- Lampsilis_ovata **
| +--------+
| | +-- Obliquaria **
+--+
| +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 16 | 14 | 11 | 58 | 58 | 4 | 121 | +----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
| | +-- Amblema
+--+ +--------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------- Popenaiadini
| |
| | +-- Glebula **
+--+ +--------------------+
| | +-- Paetulunio **
| |
+--+ +-- Toxolasma **
| +-----------------+
| | +-- Plectomerus
| |
+--+ +----------------- Obliquaria **
| |
| | +----- Potamilus **
+--+ +--------+
| | | +-- Dromus **
| | +--+
+--+ +-- Lemiox **
|
| +----------- Lampsilis_ovata **
| |
+--+ +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
Szumowski et al., 2012 [Publ 3569], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 11 | 10 | 1 | 1 | 1 | 120 | +-------------------------- Amblemini
|
--+ +----------------------- Ellipsaria **
| |
+--+ +-------------------- Truncilla **
| |
+--+ +----------------- Obliquaria **
| |
| | +-------------- Potamilus **
+--+ |
| | +----- Lampsilis_ornata **
| | +-----+
| | | | +-- Lampsilis_cardium **
+--+ | +--+
| | +-- Lampsilis_ovata **
| |
| | +-------- Lampsilis_teres **
| +--+
+--+ | +----- Lampsilis_siliquoidea **
| +--+
| | +-- Ortmanniana **
| +--+
| +-- Lampsilis_higginsii **
|
| +----- Hamiota **
| |
+-----+ +-- Epioblasma **
| |
+--+-- Cambarunio **
|
+-- Ligumia **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 15 | 13 | 12 | 185 | 11 | 3 | 119 | +----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
+--+ +----------------------------- Amblema
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +-- Obliquaria **
| +--------------------+
| | +-- Plectomerus
+--+
| +-------------------- Potamilus_leptodon **
| |
+--+ +----------------- Potamilus_alatus **
| |
| | +-- Medionidus **
+--+ +-----------+
| | +-- Ptychobranchus **
| |
+--+ +-- Ellipsaria **
| +--------+
| | +-- Truncilla **
+--+
| +-------- Leaunio **
| |
+--+ +-- Venustaconcha **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 15 | 10 | 12 | 187 | 11 | 3 | 118 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblema
| |
| | +----- Toxolasma **
+--+ +-----------+
| | | +-- Obliquaria **
| | +--+
| | +-- Plectomerus
+--+
| +-------------- Potamilus_leptodon **
| |
| | +----------- Potamilus_alatus **
+--+ |
| | +-------- Venustaconcha **
| | |
+--+ +-------- Leaunio **
| |
| | +-- Ellipsaria **
| +-----+
+--+ +-- Truncilla **
|
| +-- Medionidus **
+-----+
| +-- Ptychobranchus **
|
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 21 | 8 | 12 | 187 | 11 | 3 | 122 | +-------- Amblema
|
+-------- Toxolasma **
|
+-------- Pleurobemini
|
| +-- Obliquaria **
+-----+
| +-- Plectomerus
--+
| +----- Ellipsaria **
| |
| +----- Potamilus_leptodon **
| |
| +----- Medionidus **
| |
| +----- Potamilus_alatus **
| |
| +----- Ptychobranchus **
+--+
+----- Truncilla **
|
+----- Venustaconcha **
|
+----- Cambarunio **
|
+----- Leaunio **
|
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 8 | 25 | 6 | 3 | 122 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Ellipsaria **
| |
+--+ +----------- Truncilla **
| |
+--+ +-- Lemiox **
| +-----+
| | +-- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Cambarunio **
+--+
+-- Venustaconcha **
Chapman et al., 2008 [Publ 2744], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 11 | 8 | 7 | 4 | 114 | +----------------------------------------- Quadrulini
|
--+ +-------------------------------------- Pleurobemini
| |
+--+ +----------------------------------- Amblema
| |
+--+ +-------------------------------- Plectomerus
| |
+--+ +----------------------------- Popenaiadini
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Cyrtonaias **
+--+ +-----------------+
| | +-- Glebula **
| |
+--+ +-- Ptychobranchus **
| +--------------+
| | +-- Lemiox **
| |
+--+ +-- Venustaconcha **
| +-----------+
| | +-- Obovaria **
+--+
| +----------- Hamiota **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 8 | 26 | 5 | 3 | 116 | +----------------- Toxolasma **
|
--+ +-------------- Venustaconcha **
| |
+--+ +-- Lemiox **
| +--------+
| | +-- Lampsilis **
+--+
| +-------- Potamilus **
+--+
| +----- Cambarunio **
+--+
| +-- Ellipsaria **
+--+
+-- Truncilla **
Chapman et al., 2008 [Publ 2667], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 11 | 8 | 7 | 4 | 114 | +----------------------------------------- Quadrulini
|
--+ +-------------------------------------- Pleurobemini
| |
+--+ +----------------------------------- Amblema
| |
+--+ +-------------------------------- Plectomerus
| |
+--+ +----------------------------- Popenaiadini
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Cyrtonaias **
+--+ +-----------------+
| | +-- Glebula **
| |
+--+ +-- Ptychobranchus **
| +--------------+
| | +-- Lemiox **
| |
+--+ +-- Venustaconcha **
| +-----------+
| | +-- Obovaria **
+--+
| +----------- Hamiota **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 12 | 19 | 18 | 4 | 119 | +----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
+--+ +----------------------------- Amblemini
| |
+--+ +-------------------------- Popenaiadini
| |
+--+ +-- Cyrtonaias **
| +--------------------+
| | +-- Glebula **
+--+
| +-------------------- Potamilus **
| |
+--+ +-- Truncilla **
| +--------------+
| | +-- Ptychobranchus **
+--+
| +-------------- Pachynaias **
| |
+--+ +----------- Mesonaias **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Ortmanniana **
+--+
| +----- Cambarunio **
+--+
| +-- Epioblasma **
+--+
+-- Ligumia **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 6 | 128 | 6 | 4 | 115 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +-- Amblemini
| +--------------------+
| | +-- Popenaiadini
+--+
| +-------------------- Toxolasma **
| |
+--+ +----------------- Potamilus **
| |
+--+ +-------------- Leaunio **
| |
| | +----- Ortmanniana **
+--+ +-----+
| | | +-- Lampsilis_hydiana **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-------- Venustaconcha **
+--+
| +----- Lampsilis_ornata **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_cariosa **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 9 | 18 | 5 | 3 | 123 | +-- Quadrulini
+-----------------------+
| +-- Pleurobemini
--+
| +----------------------- Amblemini
| |
+--+ +-------------------- Truncilla **
| |
+--+ +----------------- Ligumia **
| |
| | +-- Ptychobranchus **
+--+ +-----------+
| | +-- Sagittunio **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
| | +----- Ortmanniana **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_fasciola **
|
| +----- Epioblasma **
+--+
| +-- Cambarunio **
+--+
+-- Leaunio **
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 5 | 12 | 11 | 11 | 3 | 114 | +----------------------- Quadrulini
|
--+ +-------------------- Reginaia
| |
+--+ +----------------- Amblema
| |
+--+ +-------------- Popenaiadini
| |
| | +-- Glebula **
+--+ +--------+
| | +-- Cyrtonaias **
| |
+--+ +-------- Potamilus **
| |
| | +-- Ptychobranchus **
+--+ +--+
| | +-- Truncilla **
| |
| | +-- Ortmanniana **
+--+ |
| +-- Epioblasma **
| |
| +-- Lampsilis **
| |
+--+-- Ligumia **
|
+-- Cambarunio **
|
+-- Pachynaias **
|
+-- Mesonaias **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 6 | 9 | 18 | 5 | 3 | 119 | +-- Quadrulini
+--------------+
| +-- Pleurobemini
|
--+ +-------------- Amblemini
| |
| | +----------- Ligumia **
| | |
+--+ +----------- Truncilla **
| |
| | +-- Ptychobranchus **
| +--------+
+--+ +-- Sagittunio **
|
| +-- Lampsilis_siliquoidea **
| +-----+
| | +-- Epioblasma **
| |
+--+ +----- Ortmanniana **
| |
| +----- Cambarunio **
+--+
+----- Leaunio **
|
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_fasciola **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 12 | 0 | 8 | 94 | 72 | 4 | 117 | +-------- Reginaia
|
+-------- Amblema
|
| +-- Quadrulini
+-----+
| +-- Pleurobemini
--+
| +----- Popenaiadini
| |
| | +-- Dromus **
| | |
| | +-- Epioblasma **
| | |
+--+ +-- Glebula **
| |
| +-- Lampsilis_ovata **
| |
| +-- Lampsilis_virescens **
| |
| +-- Lemiox **
+--+
+-- Obliquaria **
|
+-- Obovaria_olivaria **
|
+-- Obovaria_unicolor **
|
+-- Plectomerus
|
+-- Potamilus_alatus **
|
+-- Potamilus_capax **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 8 | 5 | 80 | 6 | 4 | 113 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-------- Venustaconcha **
| |
| | +-- Lampsilis_powellii **
+--+-----+
| +-- Lampsilis_siliquoidea **
|
| +----- Lampsilis_ornata **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_cariosa **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 5 | 80 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 5 | 84 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 5 | 84 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 11 | 11 | 3 | 124 | +----------------------------------- Quadrulini
|
--+ +-------------------------------- Reginaia
| |
+--+ +----------------------------- Amblema
| |
+--+ +-------------------------- Popenaiadini
| |
+--+ +----------------------- Cyrtonaias **
| |
+--+ +-------------------- Ptychobranchus **
| |
+--+ +----------------- Ligumia **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Pachynaias **
| |
| | +-- Epioblasma **
+--+ +-----+
| | +-- Truncilla **
| |
+--+ +-- Mesonaias **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 12 | 10 | 8 | 22 | 10 | 3 | 122 | +-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----------- Elliptoideus
| +-----------+
--+ | | +-------- Pleurobema
| | +--+
| | | +----- Uniomerus
| | +--+
| | | +-- Fusconaia
+--+ +--+
| +-- Eurynia
|
| +-------------------- Amblema
| |
| | +----- Toxolasma **
| | +-----------+
+--+ | | +-- Plectomerus
| | +--+
| | +-- Obliquaria **
| |
+--+ +----- Ptychobranchus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus **
+--+
| +-- Villosa **
| +--------+
| | +-- Ellipsaria **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
Roe, 2013 [Publ 3571], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 6 | 0 | 0 | 0 | 119 | +----------- Obliquaria **
|
--+----------- Lampsilis **
|
| +-------- Medionidus **
+--+
| +----- Ptychobranchus **
+--+
| +-- Cyprogenia **
+--+
+-- Dromus **
Roe et al., 2002 [Publ 2689], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 0 | 0 | 0 | 115 | +----- Ligumia **
|
+----- Obliquaria **
--+
+----- Lampsilis **
|
| +-- Villosa **
+--+
+-- Hamiota **
Stodola et al., 2021 [Publ 4266], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 7 | 2 | 0 | 0 | 0 | 118 | +-- Ortmanniana **
|
+-----------+-- Lampsilis_abrupta **
| |
| +-- Lampsilis_higginsii **
--+
| +----------- Lampsilis_sietmani **
| |
| | +-- Lampsilis_siliquoidea **
+--+ +-----+
| | +-- Lampsilis_radiata **
| |
+--+ +----- Lampsilis_hydiana__2 **
| |
| +----- Lampsilis_straminea__2 **
| |
+--+ +-- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
|
| +-- Lampsilis_straminea__1 **
+--+
+-- Lampsilis_virescens **
Inoue et al., 2013 [Publ 3528], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 13 | 10 | 6 | 7 | 5 | 3 | 122 | +-- Pleurobemini
+-----------+
| +-- Quadrulini
|
--+ +-- Paetulunio **
| +--------+
| | +-- Amblemini
+--+
| +-------- Lampsilis **
| |
+--+ +----- Leaunio **
| |
+--+----- Obovaria **
|
| +-- Villosa **
+--+
+-- Cambarunio **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 77 | 6 | 4 | 115 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 5 | 64 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Roe & Lydeard, 1998 [Publ 2693], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 10 | 5 | 3 | 1 | 1 | 1 | 115 | +-------------- Pleurobemini
|
+-------------- Obliquaria **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus__1 **
| |
+--+-- Potamilus_purpuratus__2 **
|
+-- Potamilus_alatus **
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 5 | 76 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 5 | 64 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 15 | 13 | 4 | 6 | 6 | 4 | 121 | +----------- Toxolasma **
+--+
| | +-------- Popenaiadini
| +--+
| | +----- Amblemini
| +--+
--+ | +-- Quadrulini
| +--+
| +-- Pleurobemini
|
| +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 5 | 64 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Turner et al., 2000 [Publ 2722], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 5 | 3 | 3 | 117 | +-- Amblemini
+--------------------+
| +-- Pleurobemini
--+
| +-------------------- Medionidus **
| |
+--+ +----------------- Lampsilis_teres **
| |
| +----------------- Villosa **
+--+
| +-- Lampsilis_satura **
| +-----------+
| | +-- Lampsilis_cardium **
+--+
| +----------- Lampsilis_reeveiana **
| |
+--+ +-------- Lampsilis_hydiana__1 **
| |
+--+-------- Lampsilis_hydiana__2 **
|
| +----- Lampsilis_siliquoidea__2 **
+--+
| +-- Lampsilis_siliquoidea__1 **
+--+
+-- Lampsilis_powellii **
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 10 | 7 | 7 | 18 | 9 | 3 | 119 | +-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----- Elliptoideus
--+ |
| +-----------------+ +-- Eurynia
| | | |
| | +--+-- Pleurobema
| | |
+--+ +-- Uniomerus
|
| +-------------------- Amblema
| |
+--+ +-- Plectomerus
| +--------------+
| | +-- Obliquaria **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus **
| |
+--+ +-------- Medionidus **
| |
+--+ +----- Potamilus_purpuratus **
| |
+--+----- Lampsilis **
|
| +-- Ellipsaria **
+--+
+-- Villosa **
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 7 | 19 | 9 | 3 | 121 | +-- Megalonaias
+-----------------------------+
| +-- Quadrulina
|
| +-------- Elliptoideus
--+ +--------------------+
| | | +----- Uniomerus
| | +--+
| | | +-- Eurynia
+--+ +--+
| +-- Pleurobema
|
| +-------------------------- Amblema
+--+
| +----------------------- Plectomerus
+--+
| +-------------------- Obliquaria **
+--+
| +----------------- Medionidus **
+--+
| +-------------- Ptychobranchus **
+--+
| +----------- Potamilus_fragilis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ellipsaria **
+--+
| +-- Villosa **
+--+
+-- Lampsilis **
Porto-Hannes et al., 2021 [Publ 4227], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 4 | 6 | 6 | 3 | 117 | +-------------------- Quadrula_quadrula
|
+-------------------- Quadrula_verrucosa
--+
| +----------------- Pleurobemini
| |
| | +-------------- Amblemini
+--+ |
| | +-- Obovaria **
| | +--------+
+--+ | +-- Ortmanniana **
| |
| | +-- Obliquaria **
| +--------+
+--+ +-- Lampsilis_ornata **
|
| +----- Lampsilis_radiata__2 **
| +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_radiata__1 **
|
| +----- Lampsilis_virescens **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 5 | 44 | 6 | 4 | 114 | +----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 8 | 39 | 6 | 3 | 117 | +----------------------- Quadrulini
|
--+ +-------------------- Amblemini
| |
+--+ +----------------- Pleurobemini
| |
+--+ +-------------- Obliquaria **
| |
| +-------------- Truncilla **
+--+
+-------------- Ptychobranchus **
|
| +----------- Ligumia **
+--+
| +-------- Cambarunio **
+--+
| +----- Epioblasma **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
Keogh & Simons, 2019 [Publ 3935], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 3 | 2 | 2 | 2 | 113 | +-- Amblemini
+-----+
| +-- Pleurobemini
--+
| +----- Potamilus **
+--+
| +-- Leaunio **
+--+
+-- Lampsilis **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 4 | 11 | 6 | 4 | 113 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Toxolasma **
| |
+--+ +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 11 | 9 | 8 | 17 | 5 | 3 | 127 | +----- Eurynia
+-----------+
| | +-- Cambarunio **
| +--+
| +-- Ptychobranchus **
--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema
+--+
| +----------- Truncilla **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Epioblasma **
+--+
| +----- Ligumia **
+--+
| +-- Obliquaria **
+--+
+-- Ortmanniana **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 4 | 60 | 4 | 4 | 113 | +-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Toxolasma **
| |
+--+ +----------- Potamilus **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 14 | 12 | 8 | 39 | 6 | 3 | 126 | +-------- Amblemini
+-----------+
| | +----- Pleurobemini
| +--+
| | +-- Obliquaria **
--+ +--+
| +-- Quadrulini
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 9 | 7 | 7 | 26 | 6 | 3 | 121 | +----------------------------- Megalonaias
|
| +----- Elliptoideus
--+ +--------------------+
| | | +-- Uniomerus
| | +--+
+--+ +-- Pleurobema
|
| +----------------------- Amblema
| |
+--+ +-- Obliquaria **
| +-----------------+
| | +-- Plectomerus
+--+
| +----------------- Medionidus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis **
+--+
| +-- Villosa **
+--+
+-- Ellipsaria **
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 13 | 10 | 8 | 38 | 5 | 2 | 126 | +----- Pleurobemini
+--------------+
| | +-- Obliquaria **
| +--+
--+ +-- Quadrulini
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 10 | 2 | 7 | 26 | 6 | 3 | 119 | +----------- Megalonaias
|
| +----- Elliptoideus
| +--+
--+ | | +-- Uniomerus
| | +--+
| | +-- Pleurobema
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus **
| |
| +----- Amblema
| |
| | +-- Obliquaria **
+--+--+
| +-- Plectomerus
|
| +-- Villosa **
| |
| +-- Ellipsaria **
| |
+--+-- Lampsilis **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 4 | 46 | 3 | 2 | 115 | +-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 29 | 27 | 6 | 22 | 22 | 3 | 135 | +-------------------------- Lampsilis **
|
--+ +----------------------- Potamilus **
| |
| | +----- Medionidus **
+--+ +--------------+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------------- Obliquaria **
| |
+--+ +-------------- Amblema
| |
| | +----- Reginaia
+--+ +-----+
| | | +-- Elliptoideus
| | +--+
+--+ +-- Plectomerus
|
| +-------- Quadrulini
+--+
| +----- Hemistena
+--+
| +-- Elliptio
+--+
+-- Fusconaia
Serb & Barnhart, 2008 [Publ 2674], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 10 | 5 | 0 | 0 | 0 | 114 | +-------------------------- Lampsilis **
|
--+ +----------------------- Obliquaria **
| |
+--+ +-------------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus **
+--+
| +-------------- Cyprogenia_aberti__4 **
| |
+--+ +----------- Cyprogenia_aberti__3 **
| |
+--+ +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Serb & Barnhart, 2008 [Publ 2674], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 9 | 5 | 0 | 0 | 0 | 113 | +----------------------- Obliquaria **
|
+----------------------- Lampsilis **
--+
| +-------------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus **
+--+
| +-------------- Cyprogenia_aberti__5 **
| |
+--+ +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 13 | 7 | 7 | 39 | 6 | 3 | 124 | +-- Obliquaria **
+--------+
| +-- Quadrulini
|
--+ +-------- Amblemini
| |
| +-------- Pleurobemini
+--+
| +----- Truncilla **
| |
| +----- Ptychobranchus **
+--+
| +-- Ortmanniana **
| |
| +-- Cambarunio **
+--+
+-- Ligumia **
|
+-- Epioblasma **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 3 | 3 | 4 | 4 | 3 | 117 | +-------------- Quadrula
|
--+-------------- Pustulosa
|
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Obovaria **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Buhay et al., 2002 [Publ 2661], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 3 | 4 | 0 | 0 | 0 | 117 | +----- Lampsilis **
|
--+----- Epioblasma **
|
| +-- Cambarunio **
+--+
+-- Villosa **
Jones & Neves, 2010 [Publ 2646], fig. 17 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 1 | 1 | 0 | 0 | 0 | 118 | Buhay et al., 2002 [Publ 2661], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 4 | 0 | 0 | 0 | 118 | +----- Lampsilis **
|
--+----- Epioblasma **
|
| +-- Cambarunio **
+--+
+-- Villosa **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 49 | 3 | 2 | 114 | +----------------- Quadrulini
--+
| +-------------- Pleurobemini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Jones & Neves, 2010 [Publ 2646], fig. 16 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 0 | 0 | 0 | 116 | Hewitt et al., 2019 [Publ 3962], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 8 | 2 | 3 | 1 | 1 | 116 | +----------------------- Pleurobemini
|
--+ +-------------------- Lampsilis_siliquoidea **
| |
| | +----------------- Lampsilis_fasciola **
+--+ |
| | +-- Epioblasma **
| +--------------+
+--+ +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_ornata **
| |
+--+ +-- Lampsilis_cardium__1 **
| +--------+
| | +-- Lampsilis_ovata__1 **
+--+
| +-------- Lampsilis_cardium__6 **
| |
+--+ +----- Lampsilis_cardium__5 **
| |
+--+ +-- Lampsilis_cardium__2 **
| |
| +-- Lampsilis_ovata__2 **
+--+
+-- Lampsilis_cardium__3 **
|
+-- Lampsilis_cardium__4 **
Serb, 2006 [Publ 2678], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 7 | 5 | 0 | 0 | 0 | 115 | +-------------- Obliquaria **
|
| +-- Ptychobranchus **
+-----------+
--+ +-- Lampsilis **
|
| +-- Cyprogenia_aberti__6 **
| +--------+
| | +-- Dromus **
+--+
| +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_aberti__4 **
| |
| | +-- Cyprogenia_stegaria__1 **
+--+ |
| +-- Cyprogenia_stegaria__2 **
| |
+--+-- Cyprogenia_aberti__1 **
|
+-- Cyprogenia_aberti__2 **
|
+-- Cyprogenia_aberti__3 **
Serb, 2006 [Publ 2678], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 8 | 5 | 0 | 0 | 0 | 113 | +-------------------- Obliquaria **
|
+-------------------- Lampsilis **
--+
| +----------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus **
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Serb, 2006 [Publ 2678], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 8 | 5 | 0 | 0 | 0 | 113 | +-------------------- Obliquaria **
|
+-------------------- Lampsilis **
--+
| +----------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus **
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 29 | 20 | 6 | 22 | 22 | 3 | 135 | +----------- Lampsilis **
|
| +-------- Obliquaria **
| |
| +-------- Amblema
| |
--+ +-------- Reginaia
| |
| +-------- Potamilus **
| |
| +-------- Quadrulini
| |
+--+ +-- Elliptoideus
+-----+
| +-- Plectomerus
|
| +----- Medionidus **
+--+
| | +-- Dromus **
| +--+
| +-- Cyprogenia **
|
| +----- Hemistena
+--+
| +-- Elliptio
+--+
+-- Fusconaia
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 11 | 8 | 6 | 25 | 5 | 3 | 118 | +-- Glebula **
+--------+
| +-- Cyrtonaias **
|
--+ +-------- Toxolasma **
| |
| | +----- Potamilus **
| +--+
+--+ | +-- Ligumia **
| +--+
| +-- Ortmanniana **
|
| +----- Amblemini
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
Keogh & Simons, 2019 [Publ 3935], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 6 | 1 | 0 | 0 | 0 | 117 | Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 25 | 5 | 3 | 115 | +----------------- Amblemini
|
| +-- Quadrulini
--+ +-----------+
| | +-- Pleurobemini
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Glebula **
+--+
| +-------- Toxolasma **
+--+
| +----- Ortmanniana **
+--+
| +-- Ligumia **
+--+
+-- Potamilus **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 4 | 31 | 1 | 1 | 115 | +-------------- Quadrulini
|
--+ +----------- Toxolasma **
| |
+--+ +-------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Venustaconcha **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 4 | 33 | 1 | 1 | 115 | +-------------- Quadrulini
|
--+ +----------- Toxolasma **
| |
+--+ +-------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Venustaconcha **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 4 | 41 | 1 | 1 | 113 | +----------- Quadrulini
--+
| +-------- Potamilus **
+--+
| +----- Venustaconcha **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 11 | 5 | 6 | 28 | 5 | 3 | 119 | +-------- Toxolasma **
|
+-------- Amblemini
|
| +-- Glebula **
+-----+
--+ +-- Cyrtonaias **
|
| +-- Quadrulini
+-----+
| +-- Pleurobemini
|
| +----- Potamilus **
+--+
| +-- Ortmanniana **
+--+
+-- Ligumia **
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 25 | 5 | 3 | 113 | +----------------- Amblemini
|
| +-- Quadrulini
--+ +-----------+
| | +-- Pleurobemini
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Glebula **
+--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 32 | 31 | 3 | 115 | +----------------------------- Amblemini
|
| +-- Cyrtonaias **
| +-----+
| | +-- Glebula **
--+ |
| +-----------------+ +-- Lampsilis **
| | | +--+
| | | | +-- Leaunio **
| | +--+
| | | +-- Truncilla **
+--+ +--+
| +-- Potamilus **
|
| +----------------------- Elliptoideus
| |
+--+ +-------------------- Quadrulini
| |
+--+ +----------------- Elliptio
| |
| | +-------------- Parvaspina
+--+ |
| +-------------- Plethobasus
| |
| | +-- Hemistena
+--+-----------+
| +-- Eurynia
|
| +----------- Fusconaia
| |
| | +----- Pleurobema_riddellii
+--+ +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +----- Pleuronaia
| |
+--+ +-- Pleurobema_pyriforme
| |
+--+-- Pleurobema_strodeanum
|
+-- Pleurobema_clava
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 12 | 9 | 6 | 25 | 5 | 3 | 127 | +----------------------- Fusconaia
|
+----------------------- Amblemini
--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema
| |
| | +-------------- Eurynia
+--+ |
| | +----- Cyrtonaias **
| +--------+
+--+ | +-- Unionini
| +--+
| +-- Anodontini
|
| +----------- Ortmanniana **
+--+
| +-------- Glebula **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Toxolasma **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 28 | 16 | 6 | 25 | 5 | 3 | 138 | +-------------------- Amblemini
|
+-------------------- Coelaturini
|
+-------------------- Eurynia
|
+-------------------- Fusconaia
|
+-------------------- Gonideini
|
+-------------------- Pleurobema
|
+-------------------- Quadrulini
--+
| +----------- Ortmanniana **
+--------+
| | +-------- Glebula **
| +--+
| | +----- Potamilus **
| +--+
| | +-- Ligumia **
| +--+
| +-- Toxolasma **
|
| +----------------- Cyrtonaias **
| |
+--+ +-------------- Unionini
| |
| | +----------- Anodontini
+--+ |
| | +----- Mutela
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaea
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 47 | 6 | 4 | 115 | +----------------------- Quadrulini
--+
| +-------------------- Pleurobemini
+--+
| +----------------- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 16 | 14 | 3 | 28 | 1 | 1 | 126 | +-- Anodontini
+-----+
| +-- Potamilus_alatus **
+-----------+
| | +----- Lamprotulini
| +--+
| | +-- Chamberlainiini
--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 26 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 10 | 1 | 1 | 119 | +----------- Mesonaias **
--+
| +-------- Potamilus **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Walker et al., 2006 [Publ 2666], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 9 | 1 | 1 | 116 | +-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Ligumia **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 8 | 6 | 4 | 37 | 37 | 4 | 115 | +-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
| | +-- Popenaiadini
+--+ +-----------+
| | +-- Amblema
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Reginaia
+--+
| +-------- Plectomerus
+--+
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 52 | 10 | 4 | 118 | +-------------------- Quadrulini
|
--+ +-- Pleurobemini
| +--------------+
| | +-- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Obliquaria **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Pachynaias **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 39 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 6 | 3 | 4 | 37 | 37 | 4 | 114 | +-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
+--+ +-- Reginaia
| +-----------+
| | +-- Amblema
+--+
| +----------- Popenaiadini
| |
+--+ +-------- Cyrtonaias **
| |
+--+-------- Plectomerus
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 11 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 10 | 1 | 1 | 117 | +-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Mesonaias **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 10 | 1 | 1 | 119 | +-------------- Pleurobemini
--+
| +----------- Mesonaias **
+--+
| +-------- Potamilus **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Walker et al., 2006 [Publ 2666], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 6 | 3 | 5 | 31 | 1 | 1 | 117 | +----------- Cyrtonaias **
|
--+----------- Pleurobemini
|
| +-------- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 36 | 35 | 4 | 38 | 38 | 4 | 136 | +-------------------------------- Quadrulini
|
| +-------------- Elliptoideus
| +--------------+
| | | +----------- Plectomerus
--+ | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis **
+--+ +--+
| +-- Lemiox **
|
| +-------- Cyrtonaias **
| +-----------------+
| | | +----- Reginaia
| | +--+
| | | +-- Popenaiadini
+--+ +--+
| +-- Amblema
|
| +----------------------- Elliptio
| |
+--+ +-------------------- Eurynia
| |
| | +----------------- Parvaspina
+--+ |
| | +-- Hemistena
| +--------------+
+--+ +-- Plethobasus
|
| +-------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
+--+ +--------+
| | +-- Fusconaia
| |
+--+ +-- Pleurobema_stabile
| +-----+
| | +-- Pleurobema_sintoxia
+--+
| +----- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
+--+
+-- Pleurobema_pyriforme
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 43 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 41 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 8 | 5 | 4 | 37 | 37 | 4 | 116 | +----------------- Quadrulini
|
--+ +-------------- Pleurobemini
| |
| | +-- Popenaiadini
+--+ +--------+
| | +-- Amblema
| |
+--+ +-------- Plectomerus
| |
| | +-- Cyrtonaias **
+--+-----+
| +-- Reginaia
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Jones et al., 2006 [Publ 2681], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 0 | 0 | 0 | 117 | Curole & Kocher, 2005 [Publ 2682], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 9 | 7 | 4 | 6 | 4 | 2 | 120 | +----------------- Glebula **
|
--+ +-- Amblema
| +-----------+
| | +-- Quadrulini
+--+
| +----------- Plectomerus
+--+
| +-------- Leaunio **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 43 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 1 | 1 | 1 | 114 | +-------- Quadrulini
--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Klymus et al., 2025 [Publ 4814], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 31 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Potamilus **
+--+
| +----- Venustaconcha **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 39 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Saxon et al., 2023 [Publ 4338], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 116 | +-- Ellipsaria **
--+
+-- Truncilla **
Buhay et al., 2002 [Publ 2661], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 117 | +-- Lampsilis_siliquoidea__1 **
|
--+-- Lampsilis_siliquoidea__2 **
|
+-- Epioblasma **
Hoeh et al., 2002 [Publ 2673], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 4 | 1 | 1 | 118 | +-------------- Pleurobemini
--+
| +----------- Cyrtonaias **
+--+
| +-------- Ortmanniana **
+--+
| +----- Lampsilis **
+--+
| +-- Potamilus **
+--+
+-- Ligumia **
Lane et al., 2016 [Publ 3735], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 1 | 3 | 0 | 0 | 0 | 117 | +-- Paetulunio **
|
--+-- Epioblasma **
|
+-- Venustaconcha **
Krebs et al., 2013 [Publ 3515], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 118 | +-------- Lampsilis_radiata **
|
--+ +-- Lampsilis_ovata **
| +--+
| | +-- Ortmanniana **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 37 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 52 | 10 | 4 | 118 | +----------------------- Quadrulini
--+
| +-------------------- Pleurobemini
+--+
| +----------------- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Obliquaria **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Pachynaias **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 7 | 4 | 2 | 116 | +-------------- Quadrulini
|
--+ +----------- Plectomerus
| |
+--+ +-------- Amblema
| |
+--+ +-- Potamilus **
| +--+
| | +-- Glebula **
+--+
| +-- Leaunio **
+--+
+-- Lampsilis **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 26 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Hoeh et al., 2002 [Publ 2673], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 4 | 1 | 1 | 116 | +-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Ligumia **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
Klymus et al., 2025 [Publ 4814], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 31 | 1 | 1 | 114 | +----------- Quadrulini
--+
| +-------- Potamilus **
+--+
| +----- Venustaconcha **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 48 | 5 | 3 | 116 | +-------------------- Quadrulini
--+
| +----------------- Pleurobemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Gangloff et al., 2013 [Publ 3551], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 0 | 0 | 0 | 117 | +----- Venustaconcha **
--+
| +-- Ligumia **
+--+
+-- Sagittunio **
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 15 | 2 | 2 | 117 | +-- Pleurobemini
+--------+
| +-- Quadrulini
--+
| +-------- Obliquaria **
+--+
| +----- Truncilla **
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
Zanatta & Harris, 2013 [Publ 3530], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 1 | 0 | 0 | 115 | +----- Lampsilis **
--+
| +-- Epioblasma **
+--+
+-- Venustaconcha **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 87 | 2 | 2 | 115 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Obliquaria **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 87 | 2 | 2 | 115 | +-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Obliquaria **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 34 | 1 | 1 | 115 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
Freeman et al., 2025 [Publ 4443], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 8 | 8 | 4 | 117 | +----- Amblemini
+--------+
| | +-- Pleurobemini
| +--+
--+ +-- Quadrulini
|
| +----------- Popenaiadini
+--+
| +-------- Toxolasma **
+--+
| +----- Obliquaria **
+--+
| +-- Glebula **
+--+
+-- Cyrtonaias **
Freeman et al., 2025 [Publ 4443], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 6 | 2 | 4 | 8 | 8 | 4 | 117 | +----- Amblemini
+--+
| | +-- Pleurobemini
| +--+
| +-- Quadrulini
--+
| +----- Obliquaria **
| |
| +----- Toxolasma **
+--+
+----- Popenaiadini
|
| +-- Glebula **
+--+
+-- Cyrtonaias **
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 16 | 5 | 3 | 115 | +-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Ouyang et al., 2015 [Publ 4493], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 9 | 7 | 3 | 28 | 6 | 1 | 122 | +-- Epioblasma **
+--+
| +-- Lampsilis **
--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 22 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 17 | 0 | 2 | 19 | 9 | 3 | 127 | +-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea **
|
+-- Lampsilis_teres **
|
+-- Megalonaias
|
+-- Obliquaria **
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unionini
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 13 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Lydeard et al., 2000 [Publ 2654], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 11 | 9 | 3 | 116 | +-------- Amblemini
|
--+ +-- Lampsilis **
| +--+
| | +-- Obovaria **
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
Lydeard et al., 2000 [Publ 2654], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 11 | 9 | 3 | 116 | +-- Lampsilis **
+-----+
| +-- Obovaria **
--+
| +----- Amblemini
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 6 | 3 | 118 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Cambarunio **
+--+
+-- Lampsilis **
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 17 | 0 | 2 | 19 | 8 | 3 | 127 | +-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea **
|
+-- Lampsilis_teres **
|
+-- Megalonaias
|
+-- Obliquaria **
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotulina
|
+-- Unionina
Freeman et al., 2025 [Publ 4443], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 8 | 8 | 4 | 116 | +----- Amblemini
+-----+
| | +-- Pleurobemini
| +--+
--+ +-- Quadrulini
|
| +-------- Popenaiadini
+--+
| +----- Obliquaria **
+--+
| +-- Glebula **
+--+
+-- Cyrtonaias **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 6 | 3 | 118 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis **
+--+
+-- Cambarunio **
Stewart et al., 2021 [Publ 4451], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 13 | 0 | 0 | 118 | Smith & Johnson, 2020 [Publ 4229], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 116 | Ouyang et al., 2015 [Publ 4493], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 28 | 6 | 1 | 116 | +-------- Pleurobemini
--+
| +----- Cyrtonaias **
+--+
| +-- Lampsilis **
+--+
+-- Epioblasma **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 15 | 2 | 2 | 116 | +-------- Quadrulini
|
--+-------- Pleurobemini
|
| +----- Lampsilis **
+--+
| +-- Epioblasma **
+--+
+-- Venustaconcha **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 78 | 6 | 3 | 116 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Cambarunio **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 1 | 3 | 0 | 0 | 118 | Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 10 | 0 | 2 | 16 | 8 | 3 | 122 | +-- Pyganodon_cataracta
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Lampsilis_teres **
|
+-- Lampsilis_straminea **
--+
+-- Megalonaias
|
+-- Obliquaria **
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 136 | 6 | 3 | 118 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis **
+--+
+-- Cambarunio **
Amaro et al., 2017 [Publ 3791], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 13 | 3 | 2 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 13 | 0 | 3 | 79 | 4 | 3 | 125 | +-- Lamprotula
|
+-- Potomida
|
+-- Pronodularia
|
+-- Gonidea
|
+-- Leguminaia
|
+-- Microcondylaea
|
--+-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 28 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 9 | 0 | 3 | 122 | 6 | 3 | 123 | +-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
--+-- Elliptio
|
+-- Eurynia
|
+-- Pleurobema
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
Lydeard et al., 2000 [Publ 2654], fig. 2c iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 11 | 9 | 3 | 118 | +----- Obovaria_unicolor **
--+
| +-- Obovaria_olivaria **
+--+
+-- Lampsilis **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 29 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 22 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Lydeard et al., 2000 [Publ 2654], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 11 | 9 | 3 | 116 | +-- Lampsilis **
--+
+-- Obovaria **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 37 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Martínez et al., 2026 [Publ 4820], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 13 | 6 | 1 | 118 | +-------- Nephronaias_hjalmarsoni
+--+
| | +----- Psoronaias
| +--+
| | +-- Nephronaias_goascoranensis
--+ +--+
| +-- Nephronaias_tempisquensis
|
| +-------- Popenaias
+--+
| +----- Mesonaias_tampicoensis **
+--+
| +-- Mesonaias_explicata **
+--+
+-- Pachynaias **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 7 | 0 | 0 | 116 | +-- Potamilus **
--+
+-- Lampsilis **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 38 | 1 | 1 | 116 | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 17 | 1 | 1 | 116 | +-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 136 | 6 | 3 | 116 | +-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 37 | 2 | 2 | 116 | +----------- Quadrulini
--+
| +-------- Amblemini
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 7 | 0 | 3 | 33 | 4 | 3 | 121 | +-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
--+-- Cambarunio **
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 22 | 3 | -- | +-------- Amblemini
|
--+ +-- Lampsilis **
| +--+
| | +-- Potamilus **
+--+
| +-- Pleurobemini
+--+
+-- Quadrulini
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Lampsilis **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 24 | 20 | 3 | -- | +----------- Quadrulini
--+
| +-------- Amblemini
+--+
| +----- Pleurobemini
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 2 | 2 | -- | +-- Lampsilis **
+--+
| +-- Obliquaria **
--+
| +-- Pleurobemini
+--+
+-- Amblemini
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 3 | 3 | -- | +----------- Pleurobemini
--+
| +-------- Quadrulini
+--+
| +----- Amblemini
+--+
| +-- Truncilla **
+--+
+-- Lampsilis **
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 19 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Amaro et al., 2017 [Publ 3791], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 2 | 2 | -- | +-- Pleurobemini
+--+
| +-- Quadrulini
--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 6 | 0 | 2 | 6 | 1 | 1 | 121 | +-------- Lampsilis **
|
--+ +-- Venustaconcha **
| +--+
| | +-- Quadrulini
+--+
| +-- Lamprotulini
+--+
+-- Chamberlainiini
Hauck et al., 2023 [Publ 4333], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 11 | 3 | -- | +----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Obovaria **
+--+
+-- Lampsilis **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
Amaro et al., 2017 [Publ 3791], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 1 | 1 | -- | +-- Toxolasma **
--+
+-- Venustaconcha **
Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 12 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Lampsilis **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Burlakova et al., 2012 [Publ 3486], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 39 | 0 | 2 | 39 | 39 | 3 | 141 | +----------- Amblema
|
--+ +-- Lampsilis **
| +-----+
| | +-- Plectomerus
+--+
| +----- Popenaiadini
+--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini
Schilling et al., 2025 [Publ 4737], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 9 | 1 | -- | +-- Lampsilis **
--+
+-- Pleurobemini
Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 3 | 2 | -- | +-- Lampsilis **
+--+
| +-- Sagittunio **
--+
| +-- Amblemini
+--+
+-- Pleurobemini
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 13 | 0 | 0 | -- | +-- Lampsilis **
--+
+-- Venustaconcha **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Toxolasma **
+--+
+-- Venustaconcha **
Freeman et al., 2025 [Publ 4443], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 4 | 1 | -- | +----- Popenaiadini
--+
| +-- Mesonaias **
+--+
+-- Pachynaias **
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Lampsilis **
Wen et al., 2017 [Publ 3795], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 3 | 3 | -- | +----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 3 | 3 | -- | +-------- Quadrulini
|
--+-------- Pleurobemini
|
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 14 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Lampsilis **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 27 | 23 | 3 | -- | +-------------- Megalonaias
|
--+ +-- Uniomerus
| +--------+
| | +-- Quadrulina
+--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
Amaro et al., 2017 [Publ 3791], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Toxolasma **
+--+
+-- Venustaconcha **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 1 | 1 | -- | +----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 3 | 3 | -- | +-- Amblemini
|
+-- Pleurobemini
--+
+-- Lampsilis **
|
+-- Quadrulini
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Combosch et al., 2017 [Publ 3806], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Campbell et al., 2008 [Publ 2675], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 24 | 3 | -- | +----------- Quadrulini
|
+----------- Amblemini
|
| +-- Elliptio_arctata
| +-----+
| | +-- Elliptio_crassidens
| |
| | +----- Pleurobema_decisum
--+ | |
| | +----- Pleurobema_rubellum
| | |
| | +----- Pleurobema_georgianum__2
| | |
| | +----- Pleurobema_stabile
| | |
| | +----- Pleurobema_pyriforme
| | |
+--+ +----- Pleurobema_oviforme__2
| |
| +----- Pleurobema_hanleyianum__2
| |
| +----- Pleuronaia
| |
| +----- Pleurobema_georgianum__3
| |
| +----- Pleurobema_cordatum
| |
| +----- Pleurobema_beadleianum
| |
| +----- Hemistena
+--+
+----- Fusconaia_flava
|
+----- Fusconaia_cerina
|
+----- Eurynia
|
+----- Elliptio_arca
|
+----- Cyrtonaias **
|
| +-- Plethobasus
+--+
| +-- Parvaspina
|
| +-- Pleurobema_sintoxia
| |
+--+-- Pleurobema_oviforme__1
| |
| +-- Pleurobema_clava
|
| +-- Pleurobema_athearni
| |
+--+-- Pleurobema_georgianum__1
|
+-- Pleurobema_hanleyianum__1
Metzger et al., 2018 [Publ 3952], fig. 1C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- | +-- Pleurobemini
--+
+-- Lampsilis **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Takeuchi et al., 2015 [Publ 3661], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Chen et al., 2025 [Publ 4455], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- | Combosch et al., 2017 [Publ 3806], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Baker et al., 2003 [Publ 2690], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | -- Epioblasma_triquetra **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 1 | -- | Xue et al., 2016 [Publ 3659], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Venustaconcha **
Froufe et al., 2015 [Publ 3664], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 8 | 3 | -- | +-- Amblemini
--+
+-- Obovaria **
Lemer et al., 2019 [Publ 4093], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Amaro et al., 2017 [Publ 3791], fig. 3A-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | -- Venustaconcha_ellipsiformis **
Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- | Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | Vikhrev et al., 2018 [Publ 3812], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- | Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Wu et al., 2020 [Publ 4121], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Lampsilis **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 0 | 0 | -- | Guerra et al., 2017 [Publ 3772], fig. 2b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Krebs, 2004 [Publ 2692], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 2 | 2 | -- | +-- Lampsilis **
--+
+-- Pleurobemini
Inoue et al., 2014 [Publ 3635], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 3 | 3 | -- | Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Venustaconcha **
Burdick & White, 2007 [Publ 2679], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 29 | 2 | -- | +-------------------------------- Lampsilis **
|
--+ +----------------------------- Parvaspina
| |
| | +-------------------------- Amblemini
+--+ |
| | +----- Eurynia
| | +-----------------+
+--+ | | +-- Plethobasus
| | +--+
| | +-- Elliptio
| |
| | +-------------------- Pleurobema_plenum
+--+ |
| +-------------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum__1
| | +-----+
| | | +-- Fusconaia
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1
| | | |
| | +--+ +-- Pleurobema_cordatum__2
| | | |
| | +--+-- Pleurobema_sintoxia__2
| | |
+--+ +-- Pleurobema_sintoxia__3
|
| +-------------- Pleurobema_stabile
| |
| | +-- Pleurobema_rubellum
| | +-----+
| | | +-- Pleurobema_strodeanum
+--+ +--+
| | | +----- Pleurobema_beadleianum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
+--+ +-- Pleurobema_taitianum
|
| +-- Pleurobema_clava
| +-----+
| | +-- Pleurobema_oviforme
| |
+--+ +-- Pleurobema_decisum
| +--+
| | +-- Pleurobema_hanleyianum
+--+
| +-- Pleurobema_pyriforme
+--+
+-- Pleurobema_georgianum
Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 0 | 0 | -- | Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 0 | 0 | -- | Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 74 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Amaro et al., 2017 [Publ 3791], fig. 3B-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | -- Venustaconcha_ellipsiformis **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 0 | 0 | -- | Campbell et al., 2008 [Publ 2675], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 25 | 2 | -- | +-- Cyrtonaias **
|
--+-- Amblemini
|
+-- Pleurobemini
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 1 | 1 | -- | +-- Amblemini
--+
+-- Lampsilis **
Zieritz et al., 2021 [Publ 4218], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- | +-- Amblemini
--+
+-- Lampsilis **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Amblemini
Froufe et al., 2015 [Publ 3664], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Jiang et al., 2010 [Publ 3438], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- | +-- Amblemini
--+
+-- Lampsilis **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | Wu et al., 2020 [Publ 4121], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 0 | 0 | -- | Plazzi & Passamonti, 2010 [Publ 3354], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 1 | 0 | 0 | -- | Curole & Kocher, 2002 [Publ 2712], fig. 1b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 1 | -- | Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Amblemini
--+
+-- Lampsilis **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Campbell et al., 2008 [Publ 2675], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 24 | 3 | -- | +----------------------- Quadrulini
|
+----------------------- Amblemini
|
| +-------------------- Cyrtonaias **
| |
| +-------------------- Eurynia
| |
| +-------------------- Fusconaia_cerina
--+ |
| +-------------------- Fusconaia_flava
| |
| +-------------------- Hemistena
| |
| +-------------------- Plethobasus
| |
| +-------------------- Parvaspina
| |
+--+-------------------- Pleurobema_cordatum
|
+-------------------- Pleuronaia
|
+-------------------- Pleurobema_oviforme__2
|
+-------------------- Pleurobema_stabile
|
+-------------------- Elliptio
|
| +----------------- Pleurobema_decisum
| |
| +----------------- Pleurobema_sintoxia
| |
| +----------------- Pleurobema_clava
+--+
+----------------- Pleurobema_oviforme__1
|
| +-------------- Pleurobema_beadleianum
| |
+--+ +----------- Pleurobema_pyriforme
| |
+--+ +-------- Pleurobema_rubellum
| |
| +-------- Pleurobema_hanleyianum__2
+--+
+-------- Pleurobema_georgianum__2
|
| +----- Pleurobema_hanleyianum__1
+--+
| +-- Pleurobema_georgianum__1
+--+
+-- Pleurobema_athearni
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Curole & Kocher, 2002 [Publ 2712], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Cyr et al., 2007 [Publ 2745], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Patterson et al., 2021 [Publ 4222], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- | +-- Villosa **
--+
+-- Amblemini
Krebs, 2004 [Publ 2692], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Pleurobemini
Campbell et al., 2008 [Publ 2675], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 25 | 2 | -- | +-- Cyrtonaias **
|
--+-- Amblemini
|
+-- Pleurobemini
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Venustaconcha **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Lampsilis **
--+
+-- Quadrulini
Wu et al., 2020 [Publ 4147], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Neemuchwala et al., 2023 [Publ 4292], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 25 | 4 | -- | +----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Popenaiadini
Huang et al., 2013 [Publ 3629], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | +-- Quadrulini
--+
+-- Venustaconcha **
Guerra et al., 2017 [Publ 3772], fig. 2a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- | +-- Venustaconcha **
--+
+-- Quadrulini
Baker et al., 2004 [Publ 2685], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | -- Epioblasma_triquetra **
Takeuchi et al., 2015 [Publ 3661], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Huang et al., 2018 [Publ 3828], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 0 | 0 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Schilling et al., 2025 [Publ 4737], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 9 | 1 | -- | +-- Lampsilis **
--+
+-- Pleurobemini
|
