The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe LANCEOLARIINI Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017: cladomics
1 genus Cladomics Summarynumber of species = 13 (15 total)
number of subtaxa (rank: genus) = 1
analyzed in 94 cladograms in 58 publications
maximum number of terminals in an analysis =
8 (mean = 2.26, median = 2)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 68
maximum number of included terminals = 8, (mean =
2.74 median = 2)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly
= 0.9
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Sano et al., 2020 [Publ 4138], fig. 1 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 22 | 5 | 1 | 31 | 19 | 4 | 0 | +----- Lanceolaria_grayii__1 **
|
+----- Lanceolaria_grayii__2 **
|
+----- Cristaria_plicata__1
|
+----- Cristaria_plicata__2
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata **
|
+----- Cristaria_plicata__3
|
+----- Lanceolaria_gladiola **
|
+----- Sinanodonta_angula
|
+----- Lepidodesmini
|
| +-- Aculamprotula
+--+
| +-- Acuticostini
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 113 | 62 | 4 | 36 | +----- Acuticostini
|
+----- Lepidodesmini
--+
+----- Unionini
|
| +-- Lanceolaria **
+--+
+-- Anodontini
Lopes-Lima et al., 2020 [Publ 4080], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 4 | 2 | 1 | 36 | +-- Unionini
--+
+-- Lanceolaria **
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 40 | 26 | 4 | 30 | +-------------- Anemina
|
--+ +----------- Sinanodonta
| |
+--+ +-------- Acuticostini
| |
+--+ +-- Lepidodesmini
| +--+
| | +-- Cristaria
+--+
| +-- Lanceolaria **
+--+
+-- Unionini
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 49 | 29 | 4 | 33 | +-------- Lepidodesmini
|
--+-------- Unionini
|
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 195 | 41 | 4 | 33 | +-------- Acuticostini
|
+-------- Alasmidonta
|
+-------- Anodonta_anatina
|
+-------- Anemina
|
+-------- Anodonta_cygnea
|
+-------- Cristaria
|
+-------- Anodonta_nuttalliana
|
+-------- Lasmigona
|
+-------- Lepidodesmini
|
+-------- Pseudanodonta
|
+-------- Pyganodon
|
--+-------- Simpsonaias
|
+-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Utterbackia
|
+-------- Lanceolaria **
|
+-------- Unio
|
+-------- Aculamprotula
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 195 | 41 | 4 | 33 | +-------- Acuticostini
|
--+ +----- Lepidodesmini
| |
+--+----- Unionini
|
| +-- Lanceolaria **
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 193 | 40 | 4 | 33 | +----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 39 | 25 | 4 | 33 | +----------------------------------- Anemina
|
--+ +-------------------------------- Acuticostini
| |
+--+ +----------------------------- Sinanodonta
| |
+--+ +-------------------------- Aculamprotula
| |
+--+ +-- Cristaria
| +--------------------+
| | +-- Lanceolaria **
+--+
| +-------------------- Lepidodesmini
+--+
| +----------------- Schistodesmus
+--+
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 23 | 4 | 32 | +----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Lepidodesmini
+--+
+-- Anodontini
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 14 | 13 | 3 | 32 | +-------------- Aculamprotula
|
--+ +-- Acuticostini
| +--------+
| | +-- Lanceolaria **
+--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 50 | 48 | 3 | 32 | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 70 | 32 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 13 | 13 | 2 | 32 | +----- Aculamprotula
+--------+
| | +-- Acuticostini
| +--+
--+ +-- Lanceolaria **
|
| +----------- Unio
| |
+--+ +-------- Pseudocuneopsis
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 21 | 19 | 1 | 21 | 19 | 3 | 46 | +----------- Acuticostini
|
--+ +-- Lanceolaria_lanceolata **
| +-----+
| | +-- Lanceolaria_grayii **
+--+
| +----- Lanceolaria_gladiola **
+--+
| +-- Anodontini
+--+
+-- Unionini
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 73 | 69 | 3 | 32 | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 53 | 23 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 92 | 43 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 65 | 29 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 70 | 30 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 43 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Xiang et al., 2025 [Publ 4433], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 10 | 9 | 1 | 32 | +-- Lanceolaria **
--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 70 | 30 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 35 | 35 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 53 | 30 | 4 | 33 | +-------- Unionini
|
--+ +-- Lepidodesmini
| +--+
| | +-- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 1 | 77 | 25 | 2 | 32 | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 20 | 18 | 3 | 32 | +-------- Acuticostini
--+
| +----- Lanceolaria **
+--+
| +-- Anodontini
+--+
+-- Unionini
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 23 | 4 | 32 | +-------- Acuticostini
|
--+ +-- Lepidodesmini
| +--+
| | +-- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 85 | 41 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 21 | 14 | 1 | 21 | 19 | 3 | 43 | +-------- Acuticostini
|
--+ +----- Lanceolaria_grayii **
| |
| +----- Lanceolaria_gladiola **
+--+
+----- Lanceolaria_lanceolata **
|
| +-- Anodontini
+--+
+-- Unionini
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 43 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 92 | 43 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 50 | 24 | 4 | 32 | +----------- Lepidodesmini
--+
| +-------- Lanceolaria **
+--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 15 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 47 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 27 | 0 | 1 | 43 | 34 | 4 | 42 | +-- Anemina
+-----------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +-----+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
+--+
| +-- Alasmidontina
| +-----+
| | +-- Anodontina
| |
+--+ +----- Lanceolaria_gladiola **
| |
| +----- Lanceolaria_oxyrhyncha **
| |
+--+----- Cristaria
|
| +-- Acuticostini
| |
+--+-- Lepidodesmini
|
+-- Unionini
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 137 | 25 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 42 | 20 | 3 | -- | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 43 | 34 | 4 | -- | +----- Pletholophus
+--------+
| | +-- Sinanodonta
| +--+
| +-- Beringiana
--+
| +----------- Cristaria
| |
| | +-- Alasmidontina
+--+--------+
| +-- Anodontina
|
| +-------- Lanceolaria **
+--+
| +----- Lepidodesmini
+--+
| +-- Acuticostini
+--+
+-- Unionini
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 13 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 33 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolaria **
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 38 | 24 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 17 | 15 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 38 | 29 | 4 | -- | +-------- Acuticostini
|
--+ +----- Lepidodesmini
| |
+--+----- Unionini
|
| +-- Lanceolaria **
+--+
+-- Anodontini
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 137 | 25 | 2 | -- | +----------- Unionini
|
--+ +-------- Lanceolaria **
| |
| +-------- Cristariina
+--+
| +----- Anodonta_anatina
| |
| +----- Pseudanodonta
+--+
+----- Alasmidontina
|
| +-- Anodonta_cygnea
+--+
+-- Anodonta_nuttalliana
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 44 | 24 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 26 | 0 | 1 | 38 | 29 | 4 | 42 | +-- Anemina
+--------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +--+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
| |
+--+ +----- Lanceolaria_gladiola **
| |
| +----- Acuticostini
| |
| +----- Lepidodesmini
| |
+--+----- Lanceolaria_oxyrhyncha **
|
+----- Cristaria
|
+----- Unionini
|
| +-- Alasmidontina
+--+
+-- Anodontina
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 74 | 24 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 33 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolaria **
+--+
+-- Unionini
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 1 | 11 | 7 | 3 | 32 | +----------- Anodontini
|
--+ +-- Nodularia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Lanceolaria_grayii **
| |
+--+----- Acuticostini
|
| +-- Lanceolaria_gladiola **
+--+
+-- Aculamprotula
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 33 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Pletholophus
| +--------+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolaria **
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 17 | 0 | 1 | 123 | 25 | 2 | 37 | +----- Unionini
|
| +-- Alasmidonta
| |
| +-- Anodonta_anatina
| |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
--+ |
| +-- Lasmigona
| |
| +-- Pseudanodonta
| |
| +-- Pyganodon
| |
| +-- Simpsonaias
| |
| +-- Strophitus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola **
|
+-- Lanceolaria_grayii__1 **
|
+-- Lanceolaria_grayii__2 **
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 19 | 11 | 3 | -- | +----- Aculamprotula
+--+
| | +-- Tchangsinaia
| +--+
--+ +-- Nodularia
|
| +----- Schistodesmus
+--+
| +-- Lanceolaria **
+--+
+-- Acuticostini
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 74 | 24 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 81 | 15 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 30 | 16 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 12 | 10 | 3 | -- | +-------- Anodontini
--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 32 | 12 | 4 | -- | +-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Lepidodesmini
|
+-------- Acuticostini
|
+-------- Cristariina
--+
+-------- Lanceolaria **
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia
| |
+--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 21 | 21 | 3 | -- | +-------- Anodontini
--+
| +----- Lanceolaria **
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 32 | 12 | 4 | -- | +----------- Anodontini
|
| +----- Acuticostini
--+ +--+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +----- Lanceolaria **
+--+
| +-- Schistodesmus
+--+
+-- Lepidodesmini
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 46 | 20 | 3 | -- | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 31 | 19 | 4 | -- | +-------------- Anodontini
|
| +-- Tchangsinaia
--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticostini
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesmini
+--+
+-- Lanceolaria **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 15 | 3 | -- | +-------- Unionini
--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 16 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 42 | 20 | 3 | -- | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 36 | 14 | 4 | -- | +----------------- Anodontini
|
| +-- Acuticostini
--+ +-----------+
| | +-- Lanceolaria **
| |
+--+ +-- Lepidodesmini
| +--------+
| | +-- Aculamprotula
+--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 17 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 13 | 9 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 9 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 20 | 3 | -- | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 5 | 2 | -- | +-- Lanceolaria **
--+
+-- Unionini
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 9 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 16 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolaria **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 9 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 7 | 2 | -- | +----- Lanceolaria **
--+
| +-- Unionini
+--+
+-- Anodontini
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 17 | 3 | -- | +-------- Lepidodesmini
--+
| +----- Unionini
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 21 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 13 | 3 | -- | +-- Lanceolaria **
--+
+-- Unionini
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 12 | 2 | -- | +-------- Aculamprotula
|
--+ +-- Schistodesmus
| +--+
| | +-- Unio
+--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 12 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 16 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 8 | 2 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 7 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 15 | 4 | -- | +----------- Anodontina
|
--+ +-- Acuticostini
| +-----+
| | +-- Cristariina
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolaria **
+--+
+-- Unionini
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 26 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolaria **
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 5 | 2 | -- | +-- Lanceolaria **
--+
+-- Unionini
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 15 | 4 | -- | +----------------- Acuticostini
|
+----------------- Cristariina
|
+----------------- Anodontina
--+
| +-------------- Lanceolaria **
| |
| | +-- Aculamprotula
| +-----------+
+--+ +-- Lepidodesmini
|
| +----------- Unio
| |
+--+ +-------- Middendorffinaia
| |
+--+ +----- Cuneopsis
| |
+--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 7 | 2 | -- | +-------- Anodontini
--+
| +----- Nodularia
+--+
| +-- Lanceolaria **
+--+
+-- Inversiunio
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 9 | 2 | -- | +----- Lanceolaria **
--+
| +-- Anodontini
+--+
+-- Unionini
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 21 | 2 | -- | +----- Unionini
--+
| +-- Lanceolaria **
+--+
+-- Anodontini
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 9 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolaria **
+--+
+-- Unionini
|
