MUSSELpdb | tribe Lanceolariini Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe LANCEOLARIINI Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017: cladomics
1 genus

Cladomics Summary

number of species = 14 (16 total)
number of subtaxa (rank: genus) = 1
analyzed in 109 cladograms in 65 publications
maximum number of terminals in an analysis = 10 (mean = 2.55, median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 80
maximum number of included terminals = 10, (mean = 3.11 median = 3)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Sano et al., 2020 [Publ 4138], fig. 1

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
3	22	5	1	31	19	4	0

  +----- Lanceolaria_oxyrhyncha__1 **
  |                               
  +----- Lanceolaria_oxyrhyncha__2 **
  |                               
  +----- Cristaria_plicata__1     
  |                               
  +----- Cristaria_plicata__2     
  |                               
  +----- Sinanodonta_lauta        
  |                               
  +----- Beringiana_japonica      
  |                               
  +----- Beringiana_fukuharai__1  
  |                               
  +----- Beringiana_beringiana    
  |                               
  +----- Beringiana_fukuharai__2  
  |                               
  +----- Beringiana_fukuharai__3  
  |                               
  +----- Buldowskia_iwakawai__1   
  |                               
  +----- Buldowskia_iwakawai__2   
  |                               
  +----- Sinanodonta_tumens__1    
--+                               
  +----- Sinanodonta_tumens__2    
  |                               
  +----- Pletholophus             
  |                               
  +----- Anemina                  
  |                               
  +----- Lanceolaria_lanceolata **   
  |                               
  +----- Cristaria_plicata__3     
  |                               
  +----- Lanceolaria_gladiola **     
  |                               
  +----- Sinanodonta_angula       
  |                               
  +----- Lepidodesmini            
  |                               
  |  +-- Aculamprotulina          
  +--+                            
  |  +-- Acuticostini             
  |                               
  |  +-- Cuneopsis                
  +--+                            
  |  +-- Nodularia                
  |                               
  |  +-- Schistodesmus            
  +--+                            
     +-- Inversiunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	7	5	1	625	149	4	35

                                +----- Amuranodonta                
  +-----------------------------+                                  
  |                             |  +-- Anemina                     
  |                             +--+                               
  |                                +-- Buldowskia                  
  |                                                                
  |                                +-- Beringiana                  
  |                          +-----+                               
--+                          |     +-- Pletholophus                
  |  +-----------------------+                                     
  |  |                       |  +----- Mimo                        
  |  |                       +--+                                  
  |  |                          |  +-- Acudonta                    
  |  |                          +--+                               
  |  |                             +-- Sinanodonta                 
  |  |                                                             
  +--+  +----------------------------- Cristaria                   
     |  |                                                          
     |  |           +----------------- Lepidodesmini               
     |  |           |                                              
     |  |           |     +----------- Lanceolaria **                 
     |  |           |  +--+                                        
     |  |  +--------+  |  |  +-------- Pseudoacuticosta_retiaria__2
     |  |  |        |  |  +--+                                     
     +--+  |        |  |     |  +----- Acuticosta_chinensis__2     
        |  |        |  |     +--+                                  
        |  |        +--+        |  +-- Acuticosta_chinensis__1     
        |  |           |        +--+                               
        |  |           |           +-- Pseudoacuticosta_retiaria__1
        |  |           |                                           
        |  |           |  +----------- Globunio                    
        |  |           +--+                                        
        |  |              |  +-------- Aculamprotulina             
        +--+              +--+                                     
           |                 |  +----- Middendorffinaiina          
           |                 +--+                                  
           |                    |  +-- Unionina                    
           |                    +--+                               
           |                       +-- Oxynaiina                   
           |                                                       
           |                 +-------- Pseudodontoideus            
           |  +--------------+                                     
           |  |              |  +----- Lasmigona                   
           |  |              +--+                                  
           |  |                 |  +-- Pterosyna                   
           +--+                 +--+                               
              |                    +-- Arcidens                    
              |                                                    
              |  +-------------------- Anodontina                  
              |  |                                                 
              +--+                 +-- Utterbackia                 
                 |  +--------------+                               
                 |  |              +-- Pyganodon                   
                 +--+                                              
                    |  +-------------- Utterbackiana               
                    |  |                                           
                    +--+  +----------- Simpsonaias                 
                       |  |                                        
                       |  |        +-- Alasminota                  
                       +--+  +-----+                               
                          |  |     +-- Alasmidonta                 
                          |  |                                     
                          +--+     +-- Prolasmidonta               
                             |  +--+                               
                             |  |  +-- Platynaias                  
                             +--+                                  
                                |  +-- Anodontoides                
                                +--+                               
                                   +-- Strophitus

Wu et al., 2026 [Publ 4825], fig. 3

tT	bi	riS	oT	roT	roS	iD
10	10	8	1	2	2	1	37

  +-- Unionini   
--+              
  +-- Lanceolaria **

Wu et al., 2026 [Publ 4825], fig. 5

tT	bi	riS	oT	roT	roS	iD
10	10	8	1	9	7	1	37

  +----- Oxynaiina      
--+                     
  |  +-- Aculamprotulina
  +--+                  
     +-- Lanceolaria **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
8	8	6	1	114	63	4	36

  +----- Acuticostini 
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolaria **  
  +--+                
     +-- Anodontini

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

tT	bi	riS	oT	roT	roS	iD
8	8	6	1	4	2	1	36

  +-- Unionini   
--+              
  +-- Lanceolaria **

Wu et al., 2026 [Publ 4825], fig. 6B

tT	bi	riS	oT	roT	roS	iD
7	7	5	1	58	56	3	35

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	41	27	4	34

  +----------------- Anemina        
  |                                 
--+  +-------------- Acuticostini   
  |  |                              
  +--+  +----------- Sinanodonta    
     |  |                           
     +--+  +-------- Aculamprotulina
        |  |                        
        +--+     +-- Cristaria      
           |  +--+                  
           |  |  +-- Lanceolaria **    
           +--+                     
              |  +-- Lepidodesmini  
              +--+                  
                 +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	194	42	4	34

  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	2	1	196	43	4	34

  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	2	1	42	28	4	34

  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticostini 
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolaria **  
           +--+                
              +-- Unionini

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	196	43	4	34

  +-------- Acuticostini         
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesmini        
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  +-------- Lanceolaria **          
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	1	49	29	4	33

  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Lanceolaria **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
4	4	2	1	77	25	2	33

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	50	48	3	33

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	70	30	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	92	62	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	82	40	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	70	30	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	43	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	70	32	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	66	33	3	33

                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Pletholophus 
  |  +--------+                   
  |  |        |  +-- Beringiana   
  |  |        +--+                
  +--+           +-- Sinanodonta  
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	43	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	53	23	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	92	43	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	66	33	3	33

                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	136	25	2	34

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	21	19	1	21	19	3	46

  +----------- Acuticostini          
  |                                  
--+        +-- Lanceolaria_lanceolata **
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii **    
  +--+                               
     |  +----- Lanceolaria_gladiola **  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	72	68	3	33

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	66	33	3	33

                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lepidodesmini
           +--+                   
              |  +-- Lanceolaria **  
              +--+                
                 +-- Unionini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	53	30	4	34

  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolaria **  
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Anodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	92	43	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	21	14	1	21	19	3	44

  +-------- Acuticostini          
  |                               
--+  +----- Lanceolaria_grayii **    
  |  |                            
  |  +----- Lanceolaria_gladiola **  
  +--+                            
     +----- Lanceolaria_lanceolata **
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	65	29	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	18	0	1	122	25	2	39

  +----- Unionini                
  |                              
  |  +-- Alasmidonta             
  |  |                           
  |  +-- Anodonta_anatina        
  |  |                           
  |  +-- Anodonta_cygnea         
  |  |                           
  |  +-- Brachyanodon            
--+  |                           
  |  +-- Lasmigona               
  |  |                           
  |  +-- Pseudanodonta           
  |  |                           
  |  +-- Pyganodon               
  |  |                           
  |  +-- Simpsonaias             
  |  |                           
  |  +-- Strophitus              
  +--+                           
     +-- Sinanodonta_lauta       
     |                           
     +-- Cristaria_plicata       
     |                           
     +-- Cristaria_radiata       
     |                           
     +-- Pletholophus            
     |                           
     +-- Sinanodonta_lucida      
     |                           
     +-- Sinanodonta_jourdyi     
     |                           
     +-- Lanceolaria_gladiola **    
     |                           
     +-- Lanceolaria_grayii **      
     |                           
     +-- Lanceolaria_danzhouensis **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	23	4	33

  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	23	4	33

  +-------- Acuticostini 
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	20	18	3	33

  +-------- Acuticostini
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Anodontini  
     +--+               
        +-- Unionini

Kim et al., 2021 [Publ 4197], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	15	14	3	33

  +-------- Aculamprotulina
  |                        
--+     +-- Acuticostini   
  |  +--+                  
  |  |  +-- Lanceolaria **    
  +--+                     
     |  +-- Unionina       
     +--+                  
        +-- Oxynaiina

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	35	35	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	84	39	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Xiang et al., 2025 [Publ 4433], fig. 7

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	10	9	1	33

  +-- Lanceolaria **
--+              
  +-- Anodontini

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	136	25	2	34

  +----------- Unionini        
  |                            
--+  +-------- Lanceolaria **     
  |  |                         
  |  +-------- Cristariina     
  +--+                         
     |  +----- Anodonta_anatina
     |  |                      
     |  +----- Pseudanodonta   
     +--+                      
        +----- Alasmidontina   
        |                      
        |  +-- Anodonta_cygnea 
        +--+                   
           +-- Brachyanodon

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	14	14	2	33

        +-- Unionina       
  +-----+                  
  |     +-- Oxynaiina      
--+                        
  |  +----- Aculamprotulina
  +--+                     
     |  +-- Acuticostini   
     +--+                  
        +-- Lanceolaria **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	51	25	4	33

  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolaria **  
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	14	3	--

  +-------- Unionini    
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Anodontini

Ouyang et al., 2011 [Publ 3408], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	4	0	1	11	7	3	34

  +----------- Anodontini          
  |                                
--+  +-------- Oxynaiina           
  |  |                             
  +--+  +----- Lanceolaria_grayii **  
     |  |                          
     +--+----- Acuticostini        
        |                          
        |  +-- Lanceolaria_gladiola **
        +--+                       
           +-- Aculamprotulina

Wang et al., 2018 [Publ 3846], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	15	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	26	0	1	42	33	4	41

              +-- Anemina               
  +-----------+                         
  |           +-- Amuranodonta          
  |                                     
  |        +----- Pletholophus          
--+  +-----+                            
  |  |     |  +-- Sinanodonta           
  |  |     +--+                         
  |  |        +-- Beringiana            
  +--+                                  
     |        +-- Alasmidontina         
     |  +-----+                         
     |  |     +-- Anodontina            
     |  |                               
     +--+  +----- Lanceolaria_gladiola **  
        |  |                            
        |  +----- Lanceolaria_oxyrhyncha **
        |  |                            
        +--+----- Cristaria             
           |                            
           |  +-- Acuticostini          
           |  |                         
           +--+-- Lepidodesmini         
              |                         
              +-- Unionini

Riccardi et al., 2020 [Publ 4089], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	17	15	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	27	0	1	39	30	4	41

           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola **  
     |  |                            
     |  +----- Acuticostini          
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha **
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	20	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	74	24	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	39	30	4	--

  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	74	24	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	14	3	--

  +-------- Lanceolaria **    
  |                        
--+     +-- Aculamprotulina
  |  +--+                  
  |  |  +-- Acuticostini   
  +--+                     
     |  +-- Anodontini     
     +--+                  
        +-- Oxynaiina

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	39	30	4	--

  +-------------------- Unionini     
  |                                  
--+  +----------------- Acuticostini 
  |  |                               
  |  |              +-- Alasmidontina
  +--+  +-----------+                
     |  |           +-- Anodontina   
     |  |                            
     |  |        +----- Lepidodesmini
     +--+  +-----+                   
        |  |     |  +-- Cristaria    
        |  |     +--+                
        |  |        +-- Lanceolaria **  
        +--+                         
           |        +-- Anemina      
           |  +-----+                
           |  |     +-- Amuranodonta 
           +--+                      
              |  +----- Pletholophus 
              +--+                   
                 |  +-- Sinanodonta  
                 +--+                
                    +-- Beringiana

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	46	20	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	44	24	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	35	14	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	138	66	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	19	11	3	--

     +----- Aculamprotulina
  +--+                     
  |  |  +-- Tchangsinaia   
  |  +--+                  
--+     +-- Nodularia      
  |                        
  |  +----- Schistodesmus  
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Acuticostini

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	46	24	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	37	15	4	--

  +----------- Anodontini     
  |                           
  |        +-- Acuticostini   
--+  +-----+                  
  |  |     +-- Lanceolaria **    
  |  |                        
  +--+     +-- Lepidodesmini  
     |  +--+                  
     |  |  +-- Aculamprotulina
     +--+                     
        |  +-- Unionina       
        +--+                  
           +-- Oxynaiina

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	12	10	3	--

  +-------- Anodontini  
--+                     
  |  +----- Unionini    
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	39	30	4	--

  +-------------- Globunionina   
  |                              
  |        +----- Aculamprotulina
--+  +-----+                     
  |  |     |  +-- Unionina       
  |  |     +--+                  
  +--+        +-- Oxynaiina      
     |                           
     |  +-------- Acuticostini   
     +--+                        
        |  +----- Lepidodesmini  
        +--+                     
           |  +-- Lanceolaria **    
           +--+                  
              +-- Anodontini

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	20	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	80	15	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Fonseca et al., 2016 [Publ 3775], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	16	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	30	16	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Xue et al., 2016 [Publ 3659], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	13	9	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	33	4	--

           +----- Pletholophus 
  +--------+                   
  |        |  +-- Sinanodonta  
  |        +--+                
  |           +-- Beringiana   
--+                            
  |  +----------- Cristaria    
  |  |                         
  |  |        +-- Alasmidontina
  +--+--------+                
     |        +-- Anodontina   
     |                         
     |  +-------- Lanceolaria **  
     +--+                      
        |  +----- Lepidodesmini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Unionini

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	21	21	3	--

  +-------- Anodontini   
--+                      
  |  +----- Lanceolaria **  
  +--+                   
     |  +-- Lepidodesmini
     +--+                
        +-- Unionini

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	32	12	4	--

  +----------- Anodontini   
  |                         
  |     +----- Acuticostini 
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Wu et al., 2026 [Publ 4811], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	15	8	3	--

  +----------- Unionina    
--+                        
  |  +-------- Lanceolaria ** 
  +--+                     
     |  +----- Globunionina
     +--+                  
        |  +-- Acuticostini
        +--+               
           +-- Anodontini

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	38	24	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	32	12	4	--

  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticostini 
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolaria **  
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona

Sano et al., 2017 [Publ 3799], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	31	19	4	--

  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticostini 
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolaria **

Wang et al., 2018 [Publ 3865], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	15	3	--

  +-------- Unionini    
--+                     
  |  +----- Anodontini  
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Bolotov et al., 2020 [Publ 4084], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	34	17	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	16	4	--

  +----------- Acuticostini      
  |                              
  +----------- Cristariina       
  |                              
--+----------- Anodontina        
  |                              
  |  +-------- Lanceolaria **       
  |  |                           
  |  |     +-- Aculamprotulina   
  +--+-----+                     
     |     +-- Lepidodesmini     
     |                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	7	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	12	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	21	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Wu et al., 2022 [Publ 4238], fig. 2e

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	17	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Huang et al., 2019 [Publ 3843], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	9	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	8	2	--

-- Lanceolaria_grayii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	0	0	--

-- Lanceolaria_grayii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	18	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	12	2	--

  +-------- Aculamprotulina
  |                        
--+     +-- Oxynaiina      
  |  +--+                  
  |  |  +-- Unionina       
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Anodontini

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	14	2	--

  +----------- Anodontini     
--+                           
  |  +-------- Oxynaiina      
  +--+                        
     |  +----- Unionina       
     +--+                     
        |  +-- Aculamprotulina
        +--+                  
           +-- Lanceolaria **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	9	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	7	2	--

  +-------- Anodontini 
--+                    
  |  +----- Nodularia  
  +--+                 
     |  +-- Lanceolaria **
     +--+              
        +-- Inversiunio

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	9	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	21	2	--

  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Liu et al., 2017 [Publ 3939], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	13	3	--

  +-- Lanceolaria **
--+              
  +-- Unionini

Kano et al., 2019 [Publ 4194], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	5	2	--

  +-- Lanceolaria **
--+              
  +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	16	4	--

  +----------- Anodontina   
  |                         
--+        +-- Acuticostini 
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Unionini

Cha et al., 2018 [Publ 3916], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	20	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	27	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	7	2	--

  +----- Lanceolaria **
--+                 
  |  +-- Unionini   
  +--+              
     +-- Anodontini

Sano et al., 2017 [Publ 3799], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	12	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	9	2	--

  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

Zhou et al., 2017 [Publ 3788], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	5	2	--

  +-- Lanceolaria **
--+              
  +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	26	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	16	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	9	2	--

  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	17	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	14	2	--

  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

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