MUSSELpdb | tribe Leoparreysiini Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe LEOPARREYSIINI Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017: cladomics
1 genus

Cladomics Summary

number of species = 9 (9 total)
number of subtaxa (rank: genus) = 1
analyzed in 40 cladograms in 26 publications
maximum number of terminals in an analysis = 5 (mean = 2.9, median = 3)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 5, (mean = 3.62 median = 3)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Bolotov et al., 2017 [Publ 3764], fig. S1

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
3	3	2	1	121	35	4	0

           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysia **            
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	627	79	4	3

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	58	57	4	3

  +----------- Parreysia     
  |                          
--+        +-- Leoparreysia **  
  |  +-----+                 
  |  |     +-- Lamellidentini
  +--+                       
     |  +----- Assamnaia     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	0	1	56	52	4	4

  +----------- Parreysiini   
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	44	19	3	3

  +-------- Lamellidentini
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysia **  
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	3

  +----------- Parreysiini   
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	3

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	106	35	4	3

  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2024 [Publ 4369], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	57	57	4	3

  +-------- Lamellidentini
  |                       
--+     +-- Leoparreysia **  
  |  +--+                 
  |  |  +-- Parreysiini   
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	56	52	4	3

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	112	36	4	5

  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	3

  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S3

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	0	0	0	3

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	13	8	2	3

  +----- Leoparreysia **  
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	79	21	4	1

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	197	31	4	2

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Konopleva et al., 2020 [Publ 4072], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	9	9	1	2

  +-- Leoparreysia **  
--+                 
  +-- Lamellidentini

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	10	3	1

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	195	31	4	3

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	197	31	4	3

  +-------- Parreysiini   
  |                       
--+-------- Lamellidentini
  |                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	88	10	3	1

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	13	13	2	--

  +----- Indochinellini
--+                    
  |  +-- Parreysiini   
  +--+                 
     +-- Leoparreysia **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	136	27	4	--

        +-- Parreysiini   
  +-----+                 
  |     +-- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	136	27	4	--

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	55	8	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	55	8	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	9	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	26	25	4	--

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	38	6	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Choi et al., 2020 [Publ 4169], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	2	2	--

  +----- Indochinellini
--+                    
  |  +-- Lamellidentini
  +--+                 
     +-- Leoparreysia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	2	2	--

  +-- Coelaturini   
  |                 
--+-- Leoparreysia **  
  |                 
  +-- Indochinellini

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	35	2	--

  +----- Leoparreysia **  
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Coelaturini

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	4	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Graf et al., 2014 [Publ 3591], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	8	1	--

  +-- Leoparreysia **
--+               
  +-- Coelaturini

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	35	2	--

  +----- Leoparreysia **  
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Coelaturini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	22	4	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysia **  
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	6	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysia **  
     +--+                 
        +-- Indochinellini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	3	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	8	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	8	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

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