The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Search the database pages:
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae tribe LEPIDODESMINI Huang & Wu in Huang et al. 2019: cladomics
1 genus Cladomics Summarynumber of species = 2 (3 total)
number of subtaxa (rank: genus) = 1
analyzed in 67 cladograms in 39 publications
maximum number of terminals in an analysis =
2 (mean = 1.18, median = 1)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 12
maximum number of included terminals = 2, (mean =
2 median = 2)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly
= 1
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 629 | 155 | 4 | -- | +----------------- Lepidodesma **
|
| +----------- Lanceolariini
| +--+
--+ | | +-------- Pseudoacuticosta_retiaria__2
| | +--+
| | | +----- Acuticosta_chinensis__2
| | +--+
+--+ | +-- Acuticosta_chinensis__1
| +--+
| +-- Pseudoacuticosta_retiaria__1
|
| +----------- Globunio
+--+
| +-------- Aculamprotulina
+--+
| +----- Middendorffinaiina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 26 | 26 | 1 | -- | +-- Lepidodesma **
--+
+-- Unionini
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 17 | 9 | 1 | -- | +-- Lepidodesma **
--+
+-- Anodontini
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 139 | 67 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 94 | 45 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 94 | 45 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 74 | 70 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 90 | 45 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wang et al., 2025 [Publ 4488], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 63 | 61 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 90 | 45 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 93 | 63 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 26 | 26 | 1 | -- | +-- Lepidodesma **
--+
+-- Unionini
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 16 | 4 | -- | +----------- Anodontina
|
--+ +-- Acuticostini
| +-----+
| | +-- Cristariina
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 32 | 4 | -- | +-------------- Anemina
|
--+ +----------- Sinanodonta
| |
+--+ +-------- Acuticostini
| |
+--+ +-- Lepidodesma **
| +--+
| | +-- Cristaria
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 16 | 4 | -- | +----------- Acuticostini
|
+----------- Cristariina
|
--+----------- Anodontina
|
| +-------- Lanceolariini
| |
| | +-- Aculamprotulina
+--+-----+
| +-- Lepidodesma **
|
| +----- Unionina
+--+
| +-- Middendorffinaiina
+--+
+-- Oxynaiina
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 26 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 16 | 4 | -- | +----------- Anodontini
|
| +-- Acuticostini
--+ +-----+
| | +-- Lanceolariini
| |
+--+ +-- Lepidodesma **
| +--+
| | +-- Aculamprotulina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 42 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 16 | 3 | -- | -- Lepidodesma_languilati **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 21 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 46 | 4 | -- | +-------- Acuticostini
|
+-------- Alasmidonta
|
+-------- Anodonta_anatina
|
+-------- Anemina
|
+-------- Anodonta_cygnea
|
+-------- Cristaria
|
+-------- Brachyanodon
|
+-------- Lasmigona
|
+-------- Lepidodesma **
|
+-------- Pseudanodonta
|
+-------- Pyganodon
|
--+-------- Simpsonaias
|
+-------- Strophitus
|
+-------- Utterbackia
|
+-------- Lanceolariini
|
+-------- Schistodesmus
|
+-------- Unionina
|
+-------- Aculamprotulina
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 6 | 2 | -- | +-------- Anodontini
--+
| +----- Aculamprotulina
+--+
| +-- Lepidodesma **
+--+
+-- Unionina
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 21 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 33 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 32 | 4 | -- | +-------- Lepidodesma **
|
--+-------- Unionini
|
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 8 | 2 | -- | +----- Anodontini
--+
| +-- Lepidodesma **
+--+
+-- Unionini
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 31 | 4 | -- | +-- Anemina
+--------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +--+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Acuticostini
| |
| +----- Lepidodesma **
| |
+--+----- Lanceolaria_oxyrhyncha
|
+----- Cristaria
|
+----- Unionini
|
| +-- Alasmidontina
+--+
+-- Anodontina
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 31 | 4 | -- | +-------------- Globunionina
|
| +----- Aculamprotulina
--+ +-----+
| | | +-- Unionina
| | +--+
+--+ +-- Oxynaiina
|
| +-------- Acuticostini
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 25 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 31 | 4 | -- | +-------- Acuticostini
|
--+ +----- Lepidodesma **
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 45 | 4 | -- | +----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 34 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 14 | 2 | -- | +-------- Anodontini
--+
| +----- Oxynaiina
+--+
| +-- Lepidodesma **
+--+
+-- Aculamprotulina
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 70 | 4 | -- | +----- Acuticostini
|
+----- Lepidodesma **
--+
+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 27 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Lanceolariini
+--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 32 | 4 | -- | +-------- Unionini
|
--+ +-- Lepidodesma **
| +--+
| | +-- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 85 | 43 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 34 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 27 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 16 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 27 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 2 | -- | +----------- Unionina
|
--+ +-- Middendorffinaiina
| +-----+
| | +-- Oxynaiina
+--+
| +----- Aculamprotulina
+--+
| +-- Anodontini
+--+
+-- Lepidodesma **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 33 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 35 | 4 | -- | +----- Pletholophus
+--------+
| | +-- Sinanodonta
| +--+
| +-- Beringiana
--+
| +----------- Cristaria
| |
| | +-- Alasmidontina
+--+--------+
| +-- Anodontina
|
| +-------- Lanceolariini
+--+
| +----- Lepidodesma **
+--+
| +-- Acuticostini
+--+
+-- Unionini
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 22 | 3 | -- | +-------- Anodontini
--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 25 | 4 | -- | +----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Anodontini
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 35 | 4 | -- | +-- Anemina
+-----------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +-----+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
+--+
| +-- Alasmidontina
| +-----+
| | +-- Anodontina
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Lanceolaria_oxyrhyncha
| |
+--+----- Cristaria
|
| +-- Acuticostini
| |
+--+-- Lepidodesma **
|
+-- Unionini
Hou et al., 2025 [Publ 4432], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 10 | 1 | -- | +-- Lepidodesma **
--+
+-- Unionini
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 31 | 4 | -- | +-------------------- Unionini
|
--+ +----------------- Acuticostini
| |
| | +-- Alasmidontina
+--+ +-----------+
| | +-- Anodontina
| |
| | +----- Lepidodesma **
+--+ +-----+
| | | +-- Cristaria
| | +--+
| | +-- Lanceolariini
+--+
| +-- Anemina
| +-----+
| | +-- Amuranodonta
+--+
| +----- Pletholophus
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 20 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 13 | 4 | -- | +----------- Anodontini
|
| +----- Acuticostini
--+ +--+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +----- Lanceolariini
+--+
| +-- Schistodesmus
+--+
+-- Lepidodesma **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 25 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 21 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 30 | 4 | -- | +----------------- Anemina
|
--+ +-------------- Acuticostini
| |
+--+ +----------- Sinanodonta
| |
+--+ +-------- Aculamprotulina
| |
+--+ +-- Cristaria
| +--+
| | +-- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 46 | 4 | -- | +-------- Acuticostini
|
--+ +----- Lepidodesma **
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 13 | 4 | -- | +-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Lepidodesma **
|
+-------- Acuticostini
|
+-------- Cristariina
--+
+-------- Lanceolariini
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia
| |
+--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 17 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 15 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 25 | 4 | -- | +-------- Acuticostini
|
--+ +-- Lepidodesma **
| +--+
| | +-- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 25 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 32 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 21 | 4 | -- | +----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1
|
+----- Cristaria_plicata__2
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula
|
+----- Lepidodesma **
|
| +-- Aculamprotulina
+--+
| +-- Acuticostini
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 51 | 3 | -- | +-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 35 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 38 | 4 | -- | +----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 34 | 3 | -- | +-- Anemina
+--------------+
| +-- Buldowskia
|
--+ +----- Pletholophus
| +--------+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 20 | 4 | -- | +-------------- Anodontini
|
| +-- Tchangsinaia
--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticostini
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesma **
+--+
+-- Lanceolariini
|
