The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini subtribe MIDDENDORFFINAIINA Lopes-Lima, Bolotov & Bogan in Lopes-Lima et al., 2020: cladomics
1 genus Cladomics Summarynumber of species = 1 (1 total)
number of subtaxa (rank: genus) = 1
analyzed in 23 cladograms in 15 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1)
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 65 | 4 | -- | +----------- Globunionina
--+
| +-------- Aculamprotulina
+--+
| +----- Middendorffinaia **
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 20 | 4 | -- | +----------------------- Globunionina
|
--+ +-------------------- Aculamprotulina
| |
+--+ +----------------- Middendorffinaia **
| |
+--+ +-------------- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Unionina
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Wu et al., 2026 [Publ 4825], fig. 6B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 64 | 28 | 3 | -- | +-------- Aculamprotulina
--+
| +----- Unionina
+--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 27 | 3 | -- | +-------- Aculamprotulina
--+
| +----- Middendorffinaia **
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 11 | 3 | -- | +----- Unionina
--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 7 | 2 | -- | +----- Unionina
--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 74 | 47 | 3 | -- | +-------- Aculamprotulina
--+
| +----- Unionina
+--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Wang et al., 2025 [Publ 4488], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 63 | 31 | 3 | -- | +-------- Aculamprotulina
--+
| +----- Unionina
+--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Hou et al., 2025 [Publ 4432], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 9 | 1 | -- | +-------- Tchangsinaia
--+
| +----- Pseudocuneopsis
+--+
| +-- Middendorffinaia **
+--+
+-- Cuneopsis
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 12 | 2 | -- | +-------- Unionina
--+
| +----- Inversiunio
+--+
| +-- Middendorffinaia **
+--+
+-- Nodularia
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 20 | 4 | -- | +-------------------- Globunionina
|
--+ +----------------- Aculamprotulina
| |
| | +-- Middendorffinaia **
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 10 | 3 | -- | +-- Middendorffinaia **
--+
+-- Oxynaiina
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 11 | 3 | -- | +-------- Middendorffinaia **
--+
| +----- Aculamprotulina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Hou et al., 2024 [Publ 4400], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 12 | 1 | -- | +----- Cuneopsis_rufescens
+--------+
| | +-- Middendorffinaia **
| +--+
--+ +-- Pseudocuneopsis
|
| +----------- Tchangsinaia
| |
+--+ +-- Cuneopsis_kiangsiensis
| +-----+
| | +-- Cuneopsis_celtiformis
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 31 | 4 | -- | +----------- Globunionina
--+
| +-------- Aculamprotulina
+--+
| +----- Unionina
+--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 25 | 1 | -- | +-- Middendorffinaia **
--+
+-- Oxynaiina
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 139 | 45 | 4 | -- | +----------- Aculamprotulina
--+
| +-------- Globunionina
+--+
| +----- Oxynaiina
+--+
| +-- Middendorffinaia **
+--+
+-- Unionina
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 25 | 1 | -- | +-- Middendorffinaia **
--+
+-- Oxynaiina
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- | +----------------- Aculamprotulina
|
| +-- Middendorffinaia **
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Lopes-Lima et al., 2020 [Publ 4080], fig. 13 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 7 | 1 | -- | +-- Middendorffinaia **
--+
+-- Oxynaiina
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 2 | -- | +----- Middendorffinaia **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- | +----------------- Aculamprotulina
|
| +-- Middendorffinaia **
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 11 | 3 | -- | +-------- Aculamprotulina
--+
| +----- Unionina
+--+
| +-- Middendorffinaia **
+--+
+-- Oxynaiina
|
