MUSSELpdb | tribe Parreysiini Henderson, 1935 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe PARREYSIINI Henderson, 1935: cladomics
2 genera

Cladomics Summary

number of species = 18 (18 total)
number of subtaxa (rank: genus) = 2
analyzed in 32 cladograms in 19 publications
maximum number of terminals in an analysis = 7 (mean = 3, median = 3)
maximum number of subtaxa (genus) = 2 (mean = 1.09, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included terminals = 7, (mean = 4.2 median = 4.5)
maximum number of subtaxa (genus) = 2 (mean = 1.15, median = 1)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 3
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 7 (mean = 7, median = 7)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.67

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Bolotov et al., 2024 [Publ 4369], fig. S1

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	62	60	2	56	55	4	0

  +----------- Parreysia **     
  |                          
--+        +-- Leoparreysiini
  |  +-----+                 
  |  |     +-- Lamellidentini
  +--+                       
     |  +----- Assamnaia **     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	7	5	2	625	77	4	59

  +----- Coelaturini
--+                 
  |  +-- Assamnaia **  
  +--+              
     +-- Parreysia **

Bolotov et al., 2024 [Publ 4369], fig. 2

tT	bi	riS	oT	roT	roS	iD
7	7	5	2	55	55	4	59

  +----------- Lamellidentini
  |                          
--+        +-- Coelaturini   
  |  +-----+                 
  |  |     +-- Indochinellini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Assamnaia **     
        +--+                 
           +-- Parreysia **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	57

  +----------- Parreysia **     
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	0	1	56	52	4	60

  +----------- Parreysia **     
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	59

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	106	35	4	57

  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	112	36	4	61

  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	56	52	4	59

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	53	49	4	61

  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	1	120	34	4	60

           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysia **               
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	135	26	4	60

        +-- Parreysia **     
  +-----+                 
  |     +-- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	197	31	4	60

  +-------- Parreysia **     
  |                       
--+-------- Lamellidentini
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	135	26	4	60

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	195	31	4	59

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	197	31	4	59

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	25	24	4	59

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	80	22	4	--

  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Lyubas et al., 2022 [Publ 4255], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	0	0	--

Lyubas et al., 2022 [Publ 4255], fig. 7

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	0	0	--

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	2	2	--

  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Imsong & Murali, 2023 [Publ 4370], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	2	2	--

  +----- Lamellidentini
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Parreysia **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	6	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Parreysia **     
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	2	2	--

  +-- Coelaturini   
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysia **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	22	4	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysia **     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	23	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Parreysia **     
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +-- Coelaturini   
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysia **

Bolotov et al., 2022 [Publ 4250], fig. S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	14	2	--

  +----- Indochinellini
--+                    
  |  +-- Parreysia **     
  +--+                 
     +-- Leoparreysiini

Graf, 2000 [Publ 2647], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	1	1	--

-- Parreysia **

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