MUSSELpdb | subtribe Pseudodontina Frierson, 1927 cladomics

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Page last updated
Thu 3 September 2025

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subtribe PSEUDODONTINA Frierson, 1927: cladomics
10 genera

Cladomics Summary

number of species = 51 (52 total)
number of subtaxa (rank: genus) = 10
analyzed in 96 cladograms in 58 publications
maximum number of terminals in an analysis = 40 (mean = 7.66, median = 3)
maximum number of subtaxa (genus) = 10 (mean = 3.7, median = 2.5)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 86
maximum number of included terminals = 40, (mean = 8.43 median = 4)
maximum number of subtaxa (genus) = 10 (mean = 4.01, median = 3)
proportion of cladograms that support basic monophyly = 0.93

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 46
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 40 (mean = 12.7, median = 9.5)
maximum number of subtaxa (genus) = 10 (mean = 5.26, median = 5.5)
proportion of cladograms that support rigorous monophyly = 0.87

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
37	37	36	10	23	5	1	0

  +----------------- Indopseudodontina
  |                                   
  |  +-------------- Pseudodon **        
--+  |                                
  |  |     +-------- Sundadontina **     
  |  |  +--+                          
  +--+  |  |  +----- Nyeinchanconcha **  
     |  |  +--+                       
     |  |     |  +-- Thaiconcha **       
     |  |     +--+                    
     +--+        +-- Bineurus **         
        |                             
        |        +-- Isannaia **         
        |  +-----+                    
        |  |     +-- Pilsbryoconcha **   
        +--+                          
           |  +----- Namkongnaia **      
           +--+                       
              |  +-- Lannanaia **        
              +--+                    
                 +-- Songkhlanaia **

Konopleva et al., 2025 [Publ 4446], fig. 4

tT	bi	riS	oT	roT	roS	iD
40	40	38	10	5	5	1	20

  +----------------------- Indopseudodontina
  |                                         
--+  +-------------------- Lannanaia **        
  |  |                                      
  +--+  +----------------- Pilsbryoconcha **   
     |  |                                   
     +--+  +-------------- Songkhlanaia **     
        |  |                                
        |  |        +----- Nyeinchanconcha **  
        +--+  +-----+                       
           |  |     |  +-- Thaiconcha **       
           |  |     +--+                    
           +--+        +-- Bineurus **         
              |                             
              |  +-------- Sundadontina **     
              +--+                          
                 |  +----- Pseudodon **        
                 +--+                       
                    |  +-- Namkongnaia **      
                    +--+                    
                       +-- Isannaia **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

tT	bi	riS	oT	roT	roS	iD
39	39	37	10	12	5	1	23

  +-------------------- Indopseudodontina
  |                                      
  |                 +-- Lannanaia **        
--+  +--------------+                    
  |  |              +-- Pilsbryoconcha **   
  |  |                                   
  +--+  +-------------- Songkhlanaia **     
     |  |                                
     |  |        +----- Nyeinchanconcha **  
     +--+  +-----+                       
        |  |     |  +-- Thaiconcha **       
        |  |     +--+                    
        +--+        +-- Bineurus **         
           |                             
           |  +-------- Sundadontina **     
           +--+                          
              |  +----- Isannaia **         
              +--+                       
                 |  +-- Namkongnaia **      
                 +--+                    
                    +-- Pseudodon **

Bolotov et al., 2023 [Publ 4312], fig. S2

tT	bi	riS	oT	roT	roS	iD
36	36	34	8	5	5	1	30

  +----------------- Indopseudodontina
  |                                   
--+  +-------------- Songkhlanaia **     
  |  |                                
  |  |           +-- Nyeinchanconcha **  
  +--+  +--------+                    
     |  |        +-- Sundadontina **     
     |  |                             
     +--+        +-- Pseudodon **        
        |  +-----+                    
        |  |     +-- Pilsbryoconcha **   
        +--+                          
           |  +----- Namkongnaia **      
           +--+                       
              |  +-- Thaiconcha **       
              +--+                    
                 +-- Bineurus **

Bolotov et al., 2023 [Publ 4312], fig. S1

tT	bi	riS	oT	roT	roS	iD
34	34	32	8	5	5	1	20

  +----------------- Indopseudodontina
  |                                   
--+  +-------------- Pilsbryoconcha **   
  |  |                                
  |  |        +----- Nyeinchanconcha **  
  +--+  +-----+                       
     |  |     |  +-- Thaiconcha **       
     |  |     +--+                    
     +--+        +-- Bineurus **         
        |                             
        |  +-------- Songkhlanaia **     
        +--+                          
           |  +----- Sundadontina **     
           +--+                       
              |  +-- Namkongnaia **      
              +--+                    
                 +-- Pseudodon **

Bolotov et al., 2023 [Publ 4312], fig. 2

tT	bi	riS	oT	roT	roS	iD
34	34	33	8	5	5	1	20

  +-------------------- Indopseudodontina
  |                                      
--+  +----------------- Pilsbryoconcha **   
  |  |                                   
  +--+  +-------------- Songkhlanaia **     
     |  |                                
     +--+           +-- Namkongnaia **      
        |  +--------+                    
        |  |        +-- Pseudodon **        
        +--+                             
           |  +-------- Sundadontina **     
           +--+                          
              |  +----- Nyeinchanconcha **  
              +--+                       
                 |  +-- Thaiconcha **       
                 +--+                    
                    +-- Bineurus **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
26	26	25	8	57	4	1	27

  +----------------------- Indopseudodontina
--+                                         
  |  +-------------------- Pilsbryoconcha **   
  +--+                                      
     |  +----------------- Songkhlanaia **     
     +--+                                   
        |  +-------------- Namkongnaia **      
        +--+                                
           |  +----------- Pseudodon **        
           +--+                             
              |  +-------- Sundadontina **     
              +--+                          
                 |  +----- Nyeinchanconcha **  
                 +--+                       
                    |  +-- Thaiconcha **       
                    +--+                    
                       +-- Bineurus **

Konopleva et al., 2021 [Publ 4213], fig. 1

tT	bi	riS	oT	roT	roS	iD
19	19	17	7	8	4	1	27

  +-------------- Indopseudodontina       
  |                                       
  |        +----- Pilsbryoconcha_exilis__2 **
--+  +-----+                              
  |  |     |  +-- Pilsbryoconcha_exilis__1 **
  |  |     +--+                           
  |  |        +-- Namkongnaia **             
  +--+                                    
     |        +-- Pseudodon **               
     |  +-----+                           
     |  |     +-- Sundadontina **            
     +--+                                 
        |  +----- Nyeinchanconcha **         
        +--+                              
           |  +-- Thaiconcha **              
           +--+                           
              +-- Bineurus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
18	18	16	7	18	4	1	36

  +----------------- Indopseudodontina    
  |                                       
  |  +-------------- Pilsbryoconcha **       
--+  |                                    
  |  |           +-- Nyeinchanconcha **      
  |  |        +--+                        
  +--+        |  +-- Sundadontina_cumingii **
     |  +-----+                           
     |  |     |  +-- Thaiconcha **           
     |  |     +--+                        
     |  |        +-- Bineurus **             
     +--+                                 
        |        +-- Namkongnaia **          
        |  +-----+                        
        |  |     +-- Pseudodon **            
        +--+                              
           |  +----- Sundadontina_tumida **  
           +--+                           
              |  +-- Sundadontina_brandti ** 
              +--+                        
                 +-- Sundadontina_taskaevi **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
17	17	16	6	95	4	1	34

  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  +--+  +-------- Nyeinchanconcha **  
     |  |                          
     +--+     +-- Thaiconcha **       
        |  +--+                    
        |  |  +-- Pilsbryoconcha **   
        +--+                       
           |  +-- Pseudodon **        
           +--+                    
              +-- Sundadontina **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
17	17	15	7	18	4	1	29

  +----------------- Indopseudodontina           
  |                                              
  |           +----- Pilsbryoconcha_exilis **       
--+  +--------+                                  
  |  |        |  +-- Pilsbryoconcha_linguaeformis **
  |  |        +--+                               
  |  |           +-- Namkongnaia_lemeslei__2 **     
  +--+                                           
     |        +----- Nyeinchanconcha **             
     |  +-----+                                  
     |  |     |  +-- Thaiconcha **                  
     |  |     +--+                               
     +--+        +-- Bineurus **                    
        |                                        
        |  +-------- Sundadontina **                
        +--+                                     
           |  +----- Pseudodon **                   
           +--+                                  
              |  +-- Namkongnaia_lemeslei__1 **     
              +--+                               
                 +-- Namkongnaia_inkhavilayi **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
17	17	16	6	101	4	1	32

  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  +--+  +-------- Nyeinchanconcha **  
     |  |                          
     +--+     +-- Thaiconcha **       
        |  +--+                    
        |  |  +-- Pilsbryoconcha **   
        +--+                       
           |  +-- Pseudodon **        
           +--+                    
              +-- Sundadontina **

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

tT	bi	riS	oT	roT	roS	iD
16	16	14	5	0	0	0	32

  +----------- Pilsbryoconcha **
--+                          
  |  +-------- Pseudodon **     
  +--+                       
     |  +----- Namkongnaia **   
     +--+                    
        |  +-- Isannaia **      
        +--+                 
           +-- Sundadontina **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
15	15	13	7	61	2	1	31

  +----------- Indopseudodontina
  |                             
  |     +----- Pilsbryoconcha **   
--+  +--+                       
  |  |  |  +-- Namkongnaia **      
  |  |  +--+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |     +-- Nyeinchanconcha **  
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
15	15	13	7	61	2	1	31

  +----------- Indopseudodontina
  |                             
  |     +----- Pilsbryoconcha **   
--+  +--+                       
  |  |  |  +-- Namkongnaia **      
  |  |  +--+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |     +-- Nyeinchanconcha **  
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
14	14	12	6	31	3	1	32

  +----------- Indopseudodontina
  |                             
  |        +-- Nyeinchanconcha **  
--+  +-----+                    
  |  |     +-- Bineurus **         
  |  |                          
  +--+     +-- Thaiconcha **       
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Pilsbryoconcha **   
        +--+                    
           +-- Pseudodon **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
14	14	11	6	31	3	1	31

  +-------- Indopseudodontina
  |                          
  |     +-- Nyeinchanconcha **  
--+  +--+                    
  |  |  +-- Bineurus **         
  |  |                       
  |  |  +-- Thaiconcha **       
  +--+--+                    
     |  +-- Sundadontina **     
     |                       
     |  +-- Pilsbryoconcha **   
     +--+                    
        +-- Pseudodon **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
13	14	13	6	126	1	1	39

  +-------------- Pilsbryoconcha **            
  |                                         
  |  +----------- Indopseudodontina         
--+  |                                      
  |  |        +-- Thaiconcha **                
  |  +--------+                             
  |  |        +-- Bineurus **                  
  +--+                                      
     |     +----- Pseudodon_cambodjensis **    
     |  +--+                                
     |  |  |  +-- Pseudodon_mekongi **         
     |  |  +--+                             
     |  |     +-- Pseudodon_vondembuschianus **
     +--+                                   
        |     +-- Pseudodon_inoscularis **     
        |  +--+                             
        |  |  +-- Sundadontina_tumida **       
        +--+                                
           |  +-- Nyeinchanconcha **           
           +--+                             
              +-- Sundadontina_cumingii **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	6	113	3	1	37

  +-------- Indopseudodontina    
  |                              
  |  +----- Bineurus **             
--+  |                           
  |  |  +-- Nyeinchanconcha **      
  |  +--+                        
  |  |  +-- Sundadontina_cumingii **
  +--+                           
     |  +-- Sundadontina_tumida **  
     +--+                        
     |  +-- Pseudodon **            
     |                           
     |  +-- Thaiconcha **           
     +--+                        
        +-- Pilsbryoconcha **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	10	6	36	0	0	28

  +-------------- Pilsbryoconcha ** 
--+                              
  |  +----------- Pseudodon **      
  +--+                           
     |  +-------- Sundadontina **   
     +--+                        
        |  +----- Nyeinchanconcha **
        +--+                     
           |  +-- Bineurus **       
           +--+                  
              +-- Thaiconcha **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
12	12	10	6	35	0	0	28

  +-------------- Pilsbryoconcha ** 
--+                              
  |  +----------- Pseudodon **      
  +--+                           
     |  +-------- Sundadontina **   
     +--+                        
        |  +----- Nyeinchanconcha **
        +--+                     
           |  +-- Bineurus **       
           +--+                  
              +-- Thaiconcha **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
12	12	13	6	187	3	1	35

  +-------------- Indopseudodontina    
  |                                    
--+  +----------- Pilsbryoconcha **       
  |  |                                 
  |  |        +-- Sundadontina_tumida **  
  +--+  +-----+                        
     |  |     +-- Pseudodon **            
     |  |                              
     +--+     +-- Nyeinchanconcha **      
        |  +--+                        
        |  |  +-- Sundadontina_cumingii **
        +--+                           
           |  +-- Thaiconcha **           
           +--+                        
              +-- Bineurus **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
12	12	12	6	189	3	1	32

  +-------------- Indopseudodontina
  |                                
--+  +----------- Pilsbryoconcha **   
  |  |                             
  +--+        +-- Pseudodon **        
     |  +-----+                    
     |  |     +-- Sundadontina **     
     +--+                          
        |  +----- Nyeinchanconcha **  
        +--+                       
           |  +-- Thaiconcha **       
           +--+                    
              +-- Bineurus **

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	10	6	37	0	0	28

  +-------------- Pilsbryoconcha ** 
--+                              
  |  +----------- Pseudodon **      
  +--+                           
     |  +-------- Sundadontina **   
     +--+                        
        |  +----- Nyeinchanconcha **
        +--+                     
           |  +-- Bineurus **       
           +--+                  
              +-- Thaiconcha **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
12	12	6	6	189	3	1	33

  +----- Indopseudodontina     
  |                            
  |  +-- Nyeinchanconcha **       
  |  |                         
--+  +-- Thaiconcha **            
  |  |                         
  |  +-- Sundadontina_cumingii ** 
  |  |                         
  |  +-- Sundadontina_tumida__1 **
  +--+                         
     +-- Sundadontina_tumida__2 **
     |                         
     +-- Pilsbryoconcha **        
     |                         
     +-- Pseudodon **             
     |                         
     +-- Bineurus **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	12	6	127	1	1	37

  +-------------- Indopseudodontina    
  |                                    
--+  +----------- Pilsbryoconcha **       
  |  |                                 
  |  |        +-- Sundadontina_tumida **  
  +--+  +-----+                        
     |  |     +-- Pseudodon **            
     |  |                              
     +--+     +-- Nyeinchanconcha **      
        |  +--+                        
        |  |  +-- Sundadontina_cumingii **
        +--+                           
           |  +-- Thaiconcha **           
           +--+                        
              +-- Bineurus **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	9	7	2	2	1	43

  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  |  |     +----- Namkongnaia **      
  +--+  +--+                       
     |  |  |  +-- Sundadontina **     
     |  |  +--+                    
     +--+     +-- Nyeinchanconcha **  
        |                          
        |  +----- Pseudodon **        
        +--+                       
           |  +-- Pilsbryoconcha **   
           +--+                    
              +-- Thaiconcha **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	0	3	75	3	1	36

  +----- Indopseudodontina         
  |                                
  |  +-- Bineurus_mouhotii **         
  |  |                             
--+  +-- Pseudodon_cambodjensis **    
  |  |                             
  |  +-- Pseudodon_mekongi **         
  |  |                             
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Pilsbryoconcha_carinifera ** 
     |                             
     +-- Bineurus_loeiensis **        
     |                             
     +-- Bineurus_exilis **           
     |                             
     +-- Pseudodon_vondembuschianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	6	5	38	2	1	34

  +----------- Indopseudodontina
  |                             
--+        +-- Pilsbryoconcha **   
  |  +-----+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |  +----- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
8	8	0	3	44	3	1	36

  +----- Indopseudodontina         
  |                                
  |  +-- Bineurus_mouhotii **         
  |  |                             
--+  +-- Pseudodon_cambodjensis **    
  |  |                             
  |  +-- Pseudodon_mekongi **         
  |  |                             
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Pilsbryoconcha_carinifera ** 
     |                             
     +-- Bineurus_loeiensis **        
     |                             
     +-- Bineurus_exilis **           
     |                             
     +-- Pseudodon_vondembuschianus **

Zieritz et al., 2024 [Publ 4381], fig. 8A

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	0	0	0	34

  +-- Namkongnaia **
--+              
  +-- Pseudodon **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	5	25	2	1	33

  +-------------- Indopseudodontina
--+                                
  |  +----------- Pilsbryoconcha **   
  +--+                             
     |  +-------- Pseudodon **        
     +--+                          
        |  +----- Sundadontina **     
        +--+                       
           |  +-- Thaiconcha **       
           +--+                    
              +-- Bineurus **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	31	2	1	34

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	31	2	1	34

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	31	2	1	36

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Bineurus **         
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Pseudodon **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
6	6	3	4	31	2	1	37

  +----------- Indopseudodontina   
  |                                
--+  +-------- Pilsbryoconcha **      
  |  |                             
  +--+  +----- Bineurus_anodontinum **
     |  |                          
     +--+----- Bineurus_loeiensis **  
        |                          
        |  +-- Thaiconcha **          
        +--+                       
           +-- Pseudodon **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
6	6	4	5	25	0	0	36

  +-------- Sundadontina **   
  |                        
--+     +-- Nyeinchanconcha **
  |  +--+                  
  |  |  +-- Bineurus **       
  +--+                     
     |  +-- Pilsbryoconcha ** 
     +--+                  
        +-- Pseudodon **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	31	2	1	34

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	31	2	1	34

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	5	34	0	0	34

  +----------- Pilsbryoconcha ** 
--+                           
  |  +-------- Pseudodon **      
  +--+                        
     |  +----- Sundadontina **   
     +--+                     
        |  +-- Nyeinchanconcha **
        +--+                  
           +-- Bineurus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
6	6	3	5	116	2	1	35

  +----------- Indopseudodontina
  |                             
--+  +-------- Pilsbryoconcha **   
  |  |                          
  +--+  +----- Pseudodon **        
     |  |                       
     +--+  +-- Nyeinchanconcha **  
        |  |                    
        +--+-- Bineurus **         
           |                    
           +-- Sundadontina **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
5	5	3	4	75	0	0	35

  +-------- Pilsbryoconcha ** 
--+                        
  |  +----- Pseudodon **      
  +--+                     
     |  +-- Bineurus **       
     +--+                  
        +-- Nyeinchanconcha **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	1	1	38

  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Bineurus **         
     +--+                       
        |  +-- Pseudodon **        
        +--+                    
           +-- Sundadontina **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	1	1	38

  +----------- Indopseudodontina
--+                             
  |  +-------- Bineurus **         
  +--+                          
     |  +----- Pilsbryoconcha **   
     +--+                       
        |  +-- Pseudodon **        
        +--+                    
           +-- Sundadontina **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	88	3	1	37

  +-------- Indopseudodontina        
--+                                  
  |  +----- Pilsbryoconcha_exilis **    
  +--+                               
     |  +-- Pseudodon **                
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	88	3	1	37

  +-------- Indopseudodontina        
--+                                  
  |  +----- Pilsbryoconcha_exilis **    
  +--+                               
     |  +-- Pseudodon **                
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	4	0	3	55	1	1	37

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	4	0	3	58	1	1	37

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	4	0	3	55	1	1	37

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	4	0	3	58	1	1	37

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	4	0	3	55	1	1	37

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Kongim et al., 2023 [Publ 4290], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	1	1	--

  +----- Indopseudodontina
--+                       
  |  +-- Namkongnaia **      
  +--+                    
     +-- Pilsbryoconcha **

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	86	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Froufe et al., 2019 [Publ 3936], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	17	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	22	2	1	--

  +----- Indopseudodontina
--+                       
  |  +-- Pseudodon **        
  +--+                    
     +-- Pilsbryoconcha **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	84	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	67	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	16	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	37	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	52	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	25	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	47	4	1	--

  +----- Indopseudodontina
--+                       
  |  +-- Pseudodon **        
  +--+                    
     +-- Sundadontina **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Chen et al., 2020 [Publ 4170], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Froufe et al., 2019 [Publ 3936], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	17	0	0	--

  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Pseudodon **   
--+               
  +-- Sundadontina **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	0	0	--

  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Pilsbryoconcha **   
--+                    
  +-- Indopseudodontina

Walker et al., 2006 [Publ 2666], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Pseudodon_vondembuschianus **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Pseudodon_vondembuschianus **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	0	0	--

-- Pilsbryoconcha_exilis **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Pilsbryoconcha_exilis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Sundadontina_tumida **

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Sundadontina_tumida **

Walker et al., 2006 [Publ 2666], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Pseudodon_vondembuschianus **

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