MUSSELpdb | subtribe Rectidentina Modell, 1942 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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subtribe RECTIDENTINA Modell, 1942: cladomics
4 genera

Cladomics Summary

number of species = 20 (20 total)
number of subtaxa (rank: genus) = 4
analyzed in 68 cladograms in 44 publications
maximum number of terminals in an analysis = 17 (mean = 5.16, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.43, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 65
maximum number of included terminals = 17, (mean = 5.35 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.49, median = 2)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 60
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 17 (mean = 5.32, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.53, median = 3)
proportion of cladograms that support rigorous monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Wu et al., 2024 [Publ 4350], fig. 5b

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	13	11	3	32	9	2	0

  +----------- Hyriopsis_bialata **  
  |                               
--+        +-- Rectidens **          
  |  +-----+                      
  |  |     +-- Ctenodesmina       
  +--+                            
     |  +----- Ensidens **           
     +--+                         
        |  +-- Hyriopsis_desowitzi **
        +--+                      
           +-- Contradentina

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
17	17	15	3	615	36	2	26

  +-------------------------- Khairuloconcha         
  |                                                  
--+  +----------------------- Ctenodesma             
  |  |                                               
  +--+  +-------------------- Contradentina          
     |  |                                            
     +--+  +----------------- Ensidens **               
        |  |                                         
        |  |           +----- Rectidens **              
        +--+  +--------+                             
           |  |        |  +-- Hyriopsis_delaportei **   
           |  |        +--+                          
           |  |           +-- Hyriopsis_desowitzi **    
           +--+                                      
              |  +----------- Hyriopsis_panhai **       
              |  |                                   
              |  |        +-- Hyriopsis_bialata **      
              +--+  +-----+                          
                 |  |     +-- Hyriopsis_khoratensis **  
                 |  |                                
                 +--+     +-- Hyriopsis_kratiensis **   
                    |  +--+                          
                    |  |  +-- Hyriopsis_phuphaniensis **
                    +--+                             
                       |  +-- Hyriopsis_myersiana **    
                       +--+                          
                          +-- Hyriopsis_sakhonensis **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
16	16	14	3	66	19	1	25

  +-------------------------- Contradentina          
  |                                                  
--+  +----------------------- Ensidens **               
  |  |                                               
  +--+                    +-- Hyriopsis_desowitzi **    
     |  +-----------------+                          
     |  |                 +-- Hyriopsis_delaportei **   
     +--+                                            
        |  +----------------- Rectidens **              
        |  |                                         
        +--+  +-------------- Hyriopsis_panhai **       
           |  |                                      
           +--+  +----------- Hyriopsis_myersiana **    
              |  |                                   
              +--+        +-- Hyriopsis_khoratensis **  
                 |  +-----+                          
                 |  |     +-- Hyriopsis_bialata **      
                 +--+                                
                    |  +----- Hyriopsis_sakhonensis **  
                    +--+                             
                       |  +-- Hyriopsis_phuphaniensis **
                       +--+                          
                          +-- Hyriopsis_kratiensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

tT	bi	riS	oT	roT	roS	iD
13	13	11	3	16	15	1	22

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Rectidens **    
     +--+                
        +-- Hyriopsis **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
12	12	0	3	22	19	1	19

  +----- Contradentina        
  |                           
  |  +-- Ensidens_ingallsianus **
  |  |                        
  |  +-- Ensidens_telus **       
  |  |                        
  |  +-- Ensidens_dugasti **     
--+  |                        
  |  +-- Hyriopsis_khoratensis **
  |  |                        
  |  +-- Hyriopsis_bialata **    
  |  |                        
  |  +-- Hyriopsis_myersiana **  
  |  |                        
  +--+-- Hyriopsis_desowitzi **  
     |                        
     +-- Rectidens **            
     |                        
     +-- Hyriopsis_panhai **     
     |                        
     +-- Ensidens_jaculus__1 **  
     |                        
     +-- Ensidens_jaculus__2 **  
     |                        
     +-- Ensidens_spiculus **    
     |                        
     +-- Ensidens_sagittarius **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	105	23	1	21

  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+  +-------- Rectidens **            
        |  |                              
        +--+     +-- Hyriopsis_khoratensis **
           |  +--+                        
           |  |  +-- Hyriopsis_bialata **    
           +--+                           
              |  +-- Hyriopsis_myersiana **  
              +--+                        
                 +-- Hyriopsis_panhai **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	99	23	1	21

  +-------------------- Contradentina        
--+                                          
  |  +----------------- Ensidens **             
  +--+                                       
     |  +-------------- Hyriopsis_desowitzi **  
     +--+                                    
        |  +----------- Rectidens **            
        +--+                                 
           |  +-------- Hyriopsis_panhai **     
           +--+                              
              |  +----- Hyriopsis_myersiana **  
              +--+                           
                 |  +-- Hyriopsis_khoratensis **
                 +--+                        
                    +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	128	12	1	20

  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+           +-- Hyriopsis_bialata **    
     |  +--------+                        
     |  |        +-- Hyriopsis_khoratensis **
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Rectidens **            
              +--+                        
                 +-- Hyriopsis_desowitzi **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	188	12	1	20

  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+  +-------- Rectidens **            
        |  |                              
        +--+     +-- Hyriopsis_panhai **     
           |  +--+                        
           |  |  +-- Hyriopsis_khoratensis **
           +--+                           
              |  +-- Hyriopsis_bialata **    
              +--+                        
                 +-- Hyriopsis_myersiana **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	5	3	190	12	1	18

  +-------- Contradentina        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Rectidens **            
  |  |                           
  +--+----- Ensidens **             
     |                           
     |  +-- Hyriopsis_bialata **    
     |  |                        
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_panhai **     
        |                        
        +-- Hyriopsis_khoratensis **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	20	15	2	19

  +----------------- Contradentina        
  |                                       
--+  +-------------- Ctenodesmina         
  |  |                                    
  +--+  +----------- Ensidens **             
     |  |                                 
     +--+  +-------- Hyriopsis_desowitzi **  
        |  |                              
        |  +-------- Rectidens **            
        +--+                              
           |  +----- Hyriopsis_panhai **     
           |  |                           
           +--+  +-- Hyriopsis_khoratensis **
              |  |                        
              +--+-- Hyriopsis_bialata **    
                 |                        
                 +-- Hyriopsis_myersiana **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	10	10	1	21

  +-------------------- Contradentina          
  |                                            
--+  +----------------- Ensidens **               
  |  |                                         
  |  |              +-- Hyriopsis_desowitzi **    
  +--+  +-----------+                          
     |  |           +-- Hyriopsis_delaportei **   
     |  |                                      
     +--+  +----------- Rectidens **              
        |  |                                   
        |  |     +----- Hyriopsis_myersiana **    
        +--+  +--+                             
           |  |  |  +-- Hyriopsis_bialata **      
           |  |  +--+                          
           +--+     +-- Hyriopsis_panhai **       
              |                                
              |  +----- Hyriopsis_khoratensis **  
              +--+                             
                 |  +-- Hyriopsis_phuphaniensis **
                 +--+                          
                    +-- Hyriopsis_sakhonensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	8	3	40	16	1	19

  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+----------- Rectidens **            
        |                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	114	12	1	19

  +-------------- Contradentina        
  |                                    
--+  +----------- Ensidens **             
  |  |                                 
  +--+  +-------- Hyriopsis_desowitzi **  
     |  |                              
     |  +-------- Rectidens **            
     +--+                              
        |  +----- Hyriopsis_panhai **     
        |  |                           
        +--+  +-- Hyriopsis_myersiana **  
           |  |                        
           +--+-- Hyriopsis_bialata **    
              |                        
              +-- Hyriopsis_khoratensis **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
11	11	6	3	190	12	1	19

  +-------- Contradentina        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Hyriopsis_myersiana **  
  |  |                           
  |  +----- Hyriopsis_panhai **     
  +--+                           
     |  +-- Hyriopsis_bialata **    
     +--+                        
     |  +-- Hyriopsis_khoratensis **
     |                           
     |  +-- Rectidens **            
     +--+                        
        +-- Ensidens **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	128	11	1	17

  +----------- Contradentina        
  |                                 
  |        +-- Hyriopsis_desowitzi **  
--+        |                        
  |  +-----+-- Rectidens **            
  |  |     |                        
  |  |     +-- Ensidens **             
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+----- Hyriopsis_khoratensis **
        |                           
        |  +-- Hyriopsis_myersiana **  
        +--+                        
           +-- Hyriopsis_bialata **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

tT	bi	riS	oT	roT	roS	iD
10	10	6	2	1	1	1	19

  +----- Ctenodesmina
--+                  
  |  +-- Rectidens **   
  +--+               
     +-- Hyriopsis **

Muanta et al., 2019 [Publ 3921], fig. 2

tT	bi	riS	oT	roT	roS	iD
9	9	8	2	0	0	0	19

  +-- Hyriopsis **
--+            
  +-- Ensidens **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	0	2	74	10	1	17

  +----- Contradentina        
  |                           
  |  +-- Ensidens_telus **       
  |  |                        
--+  +-- Ensidens_ingallsianus **
  |  |                        
  |  +-- Ensidens_jaculus **     
  |  |                        
  |  +-- Hyriopsis_desowitzi **  
  +--+                        
     +-- Hyriopsis_myersiana **  
     |                        
     +-- Hyriopsis_khoratensis **
     |                        
     +-- Ensidens_spiculus **    
     |                        
     +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

tT	bi	riS	oT	roT	roS	iD
7	7	5	2	0	0	0	16

  +-- Rectidens **
--+            
  +-- Ensidens **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
7	7	2	3	74	4	1	16

  +-------- Contradentina        
  |                              
--+  +----- Hyriopsis_desowitzi **  
  |  |                           
  |  +----- Rectidens **            
  +--+                           
     +----- Ensidens **             
     |                           
     |  +-- Hyriopsis_khoratensis **
     +--+                        
        +-- Hyriopsis_myersiana **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

tT	bi	riS	oT	roT	roS	iD
6	6	3	2	0	0	0	15

  +-- Rectidens **
--+            
  +-- Hyriopsis **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	40	13	2	14

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	32	9	2	13

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	27	10	2	13

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Zieritz et al., 2026 [Publ 4812], fig. 7a

tT	bi	riS	oT	roT	roS	iD
4	4	1	2	0	0	0	13

  +-- Hyriopsis **
--+            
  +-- Rectidens **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	32	9	2	13

  +-------------- Ctenodesmina       
--+                                  
  |  +----------- Contradentina      
  +--+                               
     |  +-------- Ensidens **           
     +--+                            
        |  +----- Hyriopsis_bialata **  
        +--+                         
           |  +-- Hyriopsis_desowitzi **
           +--+                      
              +-- Rectidens **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	4	1	13

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	32	9	2	13

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	4	1	13

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
4	13	3	3	32	9	2	8

  +-- Ensidens **           
  |                      
  +-- Hyriopsis_desowitzi **
  |                      
  +-- Rectidens **          
--+                      
  +-- Hyriopsis_bialata **  
  |                      
  +-- Ctenodesmina       
  |                      
  +-- Contradentina

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	32	9	2	13

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	56	6	2	13

  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Hyriopsis **    
        +--+                
           +-- Rectidens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	54	5	1	13

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	31	3	1	13

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Zieritz et al., 2024 [Publ 4381], fig. 6A

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	0	0	0	13

  +-- Ensidens ** 
--+            
  +-- Rectidens **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	4	4	1	13.5

  +----- Contradentina
  |                   
--+  +-- Ensidens **     
  |  |                
  |  +-- Hyriopsis__1 ** 
  +--+                
     +-- Hyriopsis__2 ** 
     |                
     +-- Rectidens **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	32	3	1	13

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	54	5	1	13

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	73	3	1	12

  +----- Contradentina
  |                   
--+  +-- Ensidens **     
  |  |                
  +--+-- Hyriopsis **    
     |                
     +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	3	1	13

  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Rectidens **    
     +--+                
        +-- Hyriopsis **

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
2	5	0	2	120	3	1	13

  +-- Lens     
  |            
  +-- Ensidens ** 
  |            
--+-- Physunio 
  |            
  +-- Rectidens **
  |            
  +-- Yaukthwa

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	3	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	83	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	5	2	--

  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	138	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	3	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	85	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	25	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	5	2	--

  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	2	1	--

  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	5	2	--

  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	6	1	--

  +-- Ensidens **     
--+                
  +-- Contradentina

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	3	1	--

  +-- Ensidens **     
--+                
  +-- Contradentina

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	3	1	--

  +-- Ensidens **     
--+                
  +-- Contradentina

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