MUSSELpdb | tribe Rectidentini Modell, 1942 cladomics

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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe RECTIDENTINI Modell, 1942: cladomics
5 genera

Cladomics Summary

number of species = 21 (21 total)
number of subtaxa (rank: genus) = 5
analyzed in 59 cladograms in 38 publications
maximum number of terminals in an analysis = 15 (mean = 5.34, median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.47, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 56
maximum number of included terminals = 15, (mean = 5.57 median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.55, median = 3)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 52
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 15 (mean = 5.52, median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.6, median = 3)
proportion of cladograms that support rigorous monophyly = 0.94

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	41	30	8	0

     +----------- Ctenodesmini 
  +--+                         
  |  |  +-------- Contradentini
  |  +--+                      
  |     |  +----- Ensidens **     
  |     +--+                   
  |        |  +-- Rectidens **    
--+        +--+                
  |           +-- Hyriopsis **    
  |                            
  |           +-- Schepmaniini 
  |     +-----+                
  |     |     +-- Pseudodontini
  +-----+                      
        |  +----- Caudiculatus 
        +--+                   
           |  +-- Lamprotulini 
           +--+                
              +-- Gonideini

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
15	15	14	3	67	53	4	16

  +-------------------------------- Pseudodontini          
  |                                                        
--+                             +-- Gonideini              
  |  +--------------------------+                          
  |  |                          +-- Lamprotulini           
  +--+                                                     
     |  +-------------------------- Contradentini          
     |  |                                                  
     +--+  +----------------------- Ensidens **               
        |  |                                               
        +--+                    +-- Hyriopsis_desowitzi **    
           |  +-----------------+                          
           |  |                 +-- Hyriopsis_delaportei **   
           +--+                                            
              |  +----------------- Rectidens **              
              |  |                                         
              +--+  +-------------- Hyriopsis_panhai **       
                 |  |                                      
                 +--+  +----------- Hyriopsis_myersiana **    
                    |  |                                   
                    +--+        +-- Hyriopsis_khoratensis **  
                       |  +-----+                          
                       |  |     +-- Hyriopsis_bialata **      
                       +--+                                
                          |  +----- Hyriopsis_sakhonensis **  
                          +--+                             
                             |  +-- Hyriopsis_phuphaniensis **
                             +--+                          
                                +-- Hyriopsis_kratiensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

tT	bi	riS	oT	roT	roS	iD
13	13	11	3	16	16	2	12

  +----------- Lamprotulini 
--+                         
  |  +-------- Contradentini
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	100	60	5	11

                 +-------- Pseudodontini        
  +--------------+                              
  |              |  +----- Gonideini            
  |              +--+                           
  |                 |  +-- Chamberlainiini      
--+                 +--+                        
  |                    +-- Lamprotulini         
  |                                             
  |  +-------------------- Contradentini        
  +--+                                          
     |  +----------------- Ensidens **             
     +--+                                       
        |  +-------------- Hyriopsis_desowitzi **  
        +--+                                    
           |  +----------- Rectidens **            
           +--+                                 
              |  +-------- Hyriopsis_panhai **     
              +--+                              
                 |  +----- Hyriopsis_myersiana **  
                 +--+                           
                    |  +-- Hyriopsis_khoratensis **
                    +--+                        
                       +-- Hyriopsis_bialata **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
12	12	0	3	22	22	2	12

  +-------- Pseudodontini        
  |                              
  |  +----- Contradentini        
  |  |                           
--+  |  +-- Ensidens_ingallsianus **
  |  |  |                        
  |  |  +-- Ensidens_telus **       
  |  |  |                        
  |  |  +-- Ensidens_dugasti **     
  +--+  |                        
     |  +-- Hyriopsis_khoratensis **
     |  |                        
     |  +-- Hyriopsis_bialata **    
     |  |                        
     |  +-- Hyriopsis_myersiana **  
     |  |                        
     +--+-- Hyriopsis_desowitzi **  
        |                        
        +-- Rectidens **            
        |                        
        +-- Hyriopsis_panhai **     
        |                        
        +-- Ensidens_jaculus__1 **  
        |                        
        +-- Ensidens_jaculus__2 **  
        |                        
        +-- Ensidens_spiculus **    
        |                        
        +-- Ensidens_sagittarius **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	106	60	5	11

     +----------------- Contradentini            
     |                                           
  +--+  +-------------- Ensidens **                 
  |  |  |                                        
  |  +--+  +----------- Hyriopsis_desowitzi **      
  |     |  |                                     
  |     +--+  +-------- Rectidens **                
  |        |  |                                  
  |        +--+     +-- Hyriopsis_khoratensis **    
  |           |  +--+                            
  |           |  |  +-- Hyriopsis_bialata **        
  |           +--+                               
  |              |  +-- Hyriopsis_myersiana **      
--+              +--+                            
  |                 +-- Hyriopsis_panhai **         
  |                                              
  |              +----- Pseudodontini            
  |              |                               
  |              |  +-- Sinosolenaia_iridinea    
  |              |  |                            
  |              |  +-- Sinosolenaia_recognita__1
  |              |  |                            
  |              |  +-- Sinosolenaia_carinata    
  |              |  |                            
  |              |  +-- Leguminaia               
  +--------------+  |                            
                 |  +-- Microcondylaea           
                 |  |                            
                 |  +-- Gonidea                  
                 |  |                            
                 |  +-- Ptychorhynchus           
                 |  |                            
                 |  +-- Parvasolenaia            
                 |  |                            
                 +--+-- Lamprotula_leaii         
                    |                            
                    +-- Lamprotula               
                    |                            
                    +-- Lamprotula_caveata       
                    |                            
                    +-- Potomida                 
                    |                            
                    +-- Pronodularia             
                    |                            
                    +-- Sinohyriopsis            
                    |                            
                    +-- Chamberlainia            
                    |                            
                    +-- Sinosolenaia_recognita__2
                    |                            
                    +-- Sinosolenaia_oleivora

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
11	11	6	3	188	45	5	11

  +-------- Contradentini        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Hyriopsis_myersiana **  
  |  |                           
  |  +----- Hyriopsis_panhai **     
  +--+                           
     |  +-- Hyriopsis_bialata **    
     +--+                        
     |  +-- Hyriopsis_khoratensis **
     |                           
     |  +-- Rectidens **            
     +--+                        
        +-- Ensidens **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	128	38	5	10

  +----------------- Contradentini        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+           +-- Hyriopsis_bialata **    
     |  +--------+                        
     |  |        +-- Hyriopsis_khoratensis **
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Rectidens **            
              +--+                        
                 +-- Hyriopsis_desowitzi **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	5	3	188	45	5	10

  +-------- Contradentini        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Rectidens **            
  |  |                           
  +--+----- Ensidens **             
     |                           
     |  +-- Hyriopsis_bialata **    
     |  |                        
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_panhai **     
        |                        
        +-- Hyriopsis_khoratensis **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	20	20	6	9

  +----------------------------- Chamberlainiini      
  |                                                   
--+  +-------------------------- Lamprotulini         
  |  |                                                
  +--+  +----------------------- Gonideini            
     |  |                                             
     +--+  +-------------------- Pseudodontini        
        |  |                                          
        +--+  +----------------- Contradentini        
           |  |                                       
           +--+  +-------------- Ctenodesmini         
              |  |                                    
              +--+  +----------- Ensidens **             
                 |  |                                 
                 +--+  +-------- Hyriopsis_desowitzi **  
                    |  |                              
                    |  +-------- Rectidens **            
                    +--+                              
                       |  +----- Hyriopsis_panhai **     
                       |  |                           
                       +--+  +-- Hyriopsis_khoratensis **
                          |  |                        
                          +--+-- Hyriopsis_bialata **    
                             |                        
                             +-- Hyriopsis_myersiana **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	10	10	1	11

  +-------------------- Contradentini          
  |                                            
--+  +----------------- Ensidens **               
  |  |                                         
  |  |              +-- Hyriopsis_desowitzi **    
  +--+  +-----------+                          
     |  |           +-- Hyriopsis_delaportei **   
     |  |                                      
     +--+  +----------- Rectidens **              
        |  |                                   
        |  |     +----- Hyriopsis_myersiana **    
        +--+  +--+                             
           |  |  |  +-- Hyriopsis_bialata **      
           |  |  +--+                          
           +--+     +-- Hyriopsis_panhai **       
              |                                
              |  +----- Hyriopsis_khoratensis **  
              +--+                             
                 |  +-- Hyriopsis_phuphaniensis **
                 +--+                          
                    +-- Hyriopsis_sakhonensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	8	3	41	27	2	9

  +----------------- Contradentini        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+----------- Rectidens **            
        |                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	128	38	5	11

  +----------- Contradentini        
  |                                 
  |        +-- Hyriopsis_desowitzi **  
--+        |                        
  |  +-----+-- Rectidens **            
  |  |     |                        
  |  |     +-- Ensidens **             
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+----- Hyriopsis_khoratensis **
        |                           
        |  +-- Hyriopsis_myersiana **  
        +--+                        
           +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	7	3	114	40	5	9

  +-------------- Contradentini        
  |                                    
--+  +----------- Ensidens **             
  |  |                                 
  +--+  +-------- Hyriopsis_desowitzi **  
     |  |                              
     |  +-------- Rectidens **            
     +--+                              
        |  +----- Hyriopsis_panhai **     
        |  |                           
        +--+  +-- Hyriopsis_myersiana **  
           |  |                        
           +--+-- Hyriopsis_bialata **    
              |                        
              +-- Hyriopsis_khoratensis **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	186	44	5	10

  +----------------- Contradentini        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+  +-------- Rectidens **            
        |  |                              
        +--+     +-- Hyriopsis_panhai **     
           |  +--+                        
           |  |  +-- Hyriopsis_khoratensis **
           +--+                           
              |  +-- Hyriopsis_bialata **    
              +--+                        
                 +-- Hyriopsis_myersiana **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

tT	bi	riS	oT	roT	roS	iD
10	10	6	2	1	1	1	9

  +----- Ctenodesmini
--+                  
  |  +-- Rectidens **   
  +--+               
     +-- Hyriopsis **

Muanta et al., 2019 [Publ 3921], fig. 2

tT	bi	riS	oT	roT	roS	iD
9	9	8	2	0	0	0	11

  +-- Hyriopsis **
--+            
  +-- Ensidens **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	0	2	75	27	4	9

  +----- Contradentini        
  |                           
  |  +-- Ensidens_telus **       
  |  |                        
--+  +-- Ensidens_ingallsianus **
  |  |                        
  |  +-- Ensidens_jaculus **     
  |  |                        
  |  +-- Hyriopsis_desowitzi **  
  +--+                        
     +-- Hyriopsis_myersiana **  
     |                        
     +-- Hyriopsis_khoratensis **
     |                        
     +-- Ensidens_spiculus **    
     |                        
     +-- Ensidens_sagittarius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
7	7	2	3	73	20	5	8

  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens **            
--+                           
  +----- Ensidens **             
  |                           
  |  +-- Hyriopsis_khoratensis **
  +--+                        
     +-- Hyriopsis_myersiana **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

tT	bi	riS	oT	roT	roS	iD
7	7	5	2	0	0	0	8

  +-- Rectidens **
--+            
  +-- Ensidens **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

tT	bi	riS	oT	roT	roS	iD
6	6	3	2	0	0	0	5

  +-- Rectidens **
--+            
  +-- Hyriopsis **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	25	5	5

  +----- Contradentini
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	28	24	7	1

  +-------------------- Chamberlainiini
  |                                    
--+  +----------------- Lamprotulini   
  |  |                                 
  |  |           +----- Gonideini      
  +--+  +--------+                     
     |  |        |  +-- Schepmaniini   
     |  |        +--+                  
     +--+           +-- Pseudodontini  
        |                              
        |  +----------- Ctenodesmini   
        +--+                           
           |  +-------- Contradentini  
           +--+                        
              |  +----- Ensidens **       
              +--+                     
                 |  +-- Rectidens **      
                 +--+                  
                    +-- Hyriopsis **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	87	25	5	5

  +----- Contradentini
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	33	31	7	3

                       +-- Schepmaniini       
  +--------------------+                      
  |                    +-- Pseudodontini      
--+                                           
  |  +-------------------- Gonideini          
  |  |                                        
  +--+                 +-- Chamberlainiini    
     |  +--------------+                      
     |  |              +-- Lamprotulini       
     +--+                                     
        |  +-------------- Ctenodesmini       
        +--+                                  
           |  +----------- Contradentini      
           +--+                               
              |  +-------- Ensidens **           
              +--+                            
                 |  +----- Hyriopsis_bialata **  
                 +--+                         
                    |  +-- Hyriopsis_desowitzi **
                    +--+                      
                       +-- Rectidens **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
4	13	3	3	33	31	7	12

  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |  +-------------------------- Pseudodontini             
--+  |                                                     
  |  |                       +-- Ensidens **                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi **       
  |  |                       |                             
  +--+                       +-- Rectidens **                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata **         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	33	31	7	1

  +----------------- Chamberlainiini
  |                                 
  |              +-- Schepmaniini   
  |           +--+                  
--+           |  +-- Pseudodontini  
  |  +--------+                     
  |  |        |  +-- Lamprotulini   
  |  |        +--+                  
  +--+           +-- Gonideini      
     |                              
     |  +----------- Ctenodesmini   
     +--+                           
        |  +-------- Contradentini  
        +--+                        
           |  +----- Ensidens **       
           +--+                     
              |  +-- Rectidens **      
              +--+                  
                 +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	33	31	7	1

  +-------------------------- Chamberlainiini
  |                                          
  |              +----------- Ctenodesmini   
  |  +-----------+                           
--+  |           |  +-------- Contradentini  
  |  |           +--+                        
  |  |              |  +----- Ensidens **       
  |  |              +--+                     
  |  |                 |  +-- Rectidens **      
  +--+                 +--+                  
     |                    +-- Hyriopsis **      
     |                                       
     |                    +-- Schepmaniini   
     |  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+                 +-- Inversidens    
        |  +--------------+                  
        |  |              +-- Lamprotulini   
        +--+                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	34	32	7	1

  +----------------- Chamberlainiini
  |                                 
  |        +-------- Lamprotulini   
--+  +-----+                        
  |  |     |  +----- Gonideini      
  |  |     +--+                     
  |  |        |  +-- Schepmaniini   
  +--+        +--+                  
     |           +-- Pseudodontini  
     |                              
     |  +----------- Ctenodesmini   
     +--+                           
        |  +-------- Contradentini  
        +--+                        
           |  +----- Ensidens **       
           +--+                     
              |  +-- Rectidens **      
              +--+                  
                 +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
4	13	11	3	33	31	7	14

  +----------------------------- Schepmaniini       
  |                                                 
  |  +-------------------------- Pseudodontini      
--+  |                                              
  |  |              +----------- Hyriopsis_bialata **  
  |  |              |                               
  |  |  +-----------+        +-- Rectidens **          
  +--+  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens **           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi **
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	54	14	5	3

  +-------------- Chamberlainiini
  |                              
  |        +----- Lamprotulini   
--+  +-----+                     
  |  |     |  +-- Gonideini      
  |  |     +--+                  
  +--+        +-- Pseudodontini  
     |                           
     |  +-------- Contradentini  
     +--+                        
        |  +----- Ensidens **       
        +--+                     
           |  +-- Hyriopsis **      
           +--+                  
              +-- Rectidens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	54	14	5	3

  +-------------- Chamberlainiini
  |                              
  |        +----- Pseudodontini  
--+  +-----+                     
  |  |     |  +-- Gonideini      
  |  |     +--+                  
  +--+        +-- Lamprotulini   
     |                           
     |  +-------- Contradentini  
     +--+                        
        |  +----- Ensidens **       
        +--+                     
           |  +-- Hyriopsis **      
           +--+                  
              +-- Rectidens **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	55	55	7	3

           +-------- Lamprotulini   
  +--------+                        
  |        |  +----- Gonideini      
  |        +--+                     
  |           |  +-- Schepmaniini   
--+           +--+                  
  |              +-- Pseudodontini  
  |                                 
  |  +-------------- Chamberlainiini
  +--+                              
     |  +----------- Ctenodesmini   
     +--+                           
        |  +-------- Contradentini  
        +--+                        
           |  +----- Ensidens **       
           +--+                     
              |  +-- Hyriopsis **      
              +--+                  
                 +-- Rectidens **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	33	5	3

     +-------- Contradentini  
  +--+                        
  |  |  +----- Ensidens **       
  |  +--+                     
  |     |  +-- Hyriopsis **      
--+     +--+                  
  |        +-- Rectidens **      
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	33	5	3

     +-------- Contradentini  
  +--+                        
  |  |  +----- Ensidens **       
  |  +--+                     
  |     |  +-- Hyriopsis **      
--+     +--+                  
  |        +-- Rectidens **      
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Zieritz et al., 2024 [Publ 4381], fig. 6A

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	0	0	0	3

  +-- Ensidens ** 
--+            
  +-- Rectidens **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	4	4	1	5.5

  +----- Contradentini
  |                   
--+  +-- Ensidens **     
  |  |                
  |  +-- Hyriopsis__1 ** 
  +--+                
     +-- Hyriopsis__2 ** 
     |                
     +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	73	28	5	4

  +----- Contradentini
  |                   
--+  +-- Ensidens **     
  |  |                
  +--+-- Hyriopsis **    
     |                
     +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	28	5	3

  +-------- Contradentini
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Rectidens **    
     +--+                
        +-- Hyriopsis **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	10	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	86	17	5	--

  +----- Contradentini
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	15	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	52	21	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	40	7	--

        +-------- Ctenodesmini   
  +-----+                        
  |     |  +----- Contradentini  
  |     +--+                     
  |        |  +-- Ensidens **       
--+        +--+                  
  |           +-- Hyriopsis **      
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	18	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	41	5	--

  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Contradentini  
  |  +--------------+                     
--+  |              |  +-- Hyriopsis **      
  |  |              +--+                  
  |  |                 +-- Ensidens **       
  |  |                                    
  +--+              +----- Inversidens    
     |  +-----------+                     
     |  |           |  +-- Chamberlainiini
     |  |           +--+                  
     |  |              +-- Lamprotulini   
     +--+                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	19	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	17	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	19	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	16	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	26	24	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	41	7	--

        +-------- Ctenodesmini   
  +-----+                        
  |     |  +----- Contradentini  
  |     +--+                     
  |        |  +-- Ensidens **       
--+        +--+                  
  |           +-- Hyriopsis **      
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
2	5	0	2	119	31	5	6

  +-- Lens     
  |            
  +-- Ensidens ** 
  |            
--+-- Physunio 
  |            
  +-- Rectidens **
  |            
  +-- Yaukthwa

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	18	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	16	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Contradentini  
--+  +--+                     
  |  |  |  +-- Hyriopsis **      
  |  |  +--+                  
  +--+     +-- Rectidens **      
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	41	5	--

  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Contradentini  
--+  +--------------+                     
  |  |              |  +-- Hyriopsis **      
  |  |              +--+                  
  |  |                 +-- Ensidens **       
  +--+                                    
     |  +----------------- Inversidens    
     |  |                                 
     |  |              +-- Chamberlainiini
     |  +--------------+                  
     +--+              +-- Lamprotulini   
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	7	2	--

  +-- Ensidens **     
--+                
  +-- Contradentini

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	9	2	--

  +----- Pseudodontini
--+                   
  |  +-- Ensidens **     
  +--+                
     +-- Contradentini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	13	4	--

        +-- Ensidens **     
  +-----+                
  |     +-- Contradentini
--+                      
  |  +----- Pseudodontini
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

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