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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae

tribe SCHEPMANIINI Lopes-Lima, Pfeiffer & Zieritz in Zieritz et al., 2021: cladomics
     1 genus

Cladomics Summary

number of species = 2 (2 total)
number of subtaxa (rank: genus) = 1
analyzed in 14 cladograms in 11 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = , (mean = median = )
maximum number of subtaxa (genus) = (mean = , median = )
proportion of cladograms that support basic monophyly = 0

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11016301346--
     +----------------------------- Indopseudodontina         
     |                                                        
     |                          +-- Lannanaia                 
  +--+  +-----------------------+                             
  |  |  |                       +-- Pilsbryoconcha            
  |  |  |                                                     
  |  +--+  +----------------------- Songkhlanaia              
  |     |  |                                                  
  |     |  |                    +-- Nyeinchanconcha           
  |     +--+  +-----------------+                             
  |        |  |                 +-- Sundadontina              
  |        |  |                                               
  |        |  |                 +-- Isannaia                  
  |        +--+              +--+                             
  |           |              |  +-- Namkongnaia               
  |           |  +-----------+                                
  |           |  |           |  +-- Thaiconcha                
  |           |  |           +--+                             
  |           +--+              +-- Bineurus                  
  |              |                                            
  |              |  +-------------- Pseudodon_walpolei        
  |              |  |                                         
  |              +--+  +----------- Pseudodon_vagulus         
  |                 |  |                                      
  |                 |  |     +----- Pseudodon_vondembuschianus
--+                 +--+  +--+                                
  |                    |  |  |  +-- Pseudodon_mekongi         
  |                    |  |  +--+                             
  |                    +--+     +-- Schepmania **                
  |                       |                                   
  |                       |  +----- Pseudodon_lenyanensis__2  
  |                       +--+                                
  |                          |  +-- Pseudodon_inoscularis     
  |                          +--+                             
  |                             +-- Pseudodon_lenyanensis__1  
  |                                                           
  |           +-------------------- Chamberlainiini           
  |           |                                               
  |        +--+  +----------------- Parvasolenaia             
  |        |  |  |                                            
  |        |  +--+  +-------------- Pseudopostolata           
  |        |     |  |                                         
  |        |     +--+  +----------- Koreosolenaia             
  |        |        |  |                                      
  |        |        +--+        +-- Cosmopseudodon            
  |        |           |  +-----+                             
  |        |           |  |     +-- Guiunio                   
  |        |           +--+                                   
  +--------+              |  +----- Ptychorhynchus            
           |              +--+                                
           |                 |  +-- Obovalis                  
           |                 +--+                             
           |                    +-- Postolata                 
           |                                                  
           |                    +-- Caudiculatus              
           |     +--------------+                             
           |     |              +-- Lamprotulini              
           |     |                                            
           +-----+  +-------------- Sinosolenaia_carinata     
                 |  |                                         
                 |  |        +----- Gonidea                   
                 +--+  +-----+                                
                    |  |     |  +-- Microcondylaea            
                    |  |     +--+                             
                    +--+        +-- Leguminaia                
                       |                                      
                       |  +-------- Inversidens               
                       +--+                                   
                          |  +----- Sinosolenaia_recognita    
                          +--+                                
                             |  +-- Sinosolenaia_iridinea     
                             +--+                             
                                +-- Sinosolenaia_oleivora

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
110135335--
  +----------------------------- Schepmania **     
  |                                             
--+  +-------------------------- Pseudodontini  
  |  |                                          
  +--+  +----------------------- Rectidentini   
     |  |                                       
     +--+  +-------------------- Inversidens    
        |  |                                    
        |  |              +----- Lamprotula     
        +--+  +-----------+                     
           |  |           |  +-- Chamberlainiini
           |  |           +--+                  
           +--+              +-- Potomida       
              |                                 
              |  +-------------- Sinosolenaia   
              |  |                              
              +--+           +-- Gonidea        
                 |  +--------+                  
                 |  |        +-- Microcondylaea 
                 +--+                           
                    |  +-------- Koreosolenaia  
                    +--+                        
                       |  +----- Parvasolenaia  
                       +--+                     
                          |  +-- Postolata      
                          +--+                  
                             +-- Ptychorhynchus

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
110135335--
           +-- Schepmania **     
  +--------+                  
  |        +-- Pseudodontini  
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Rectidentini   
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
110130265--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Lamprotulini   
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmania **     
           +--+                  
              +-- Pseudodontini

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
110145405--
  +-------------- Rectidentini   
--+                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmania **     
           +--+                  
              +-- Pseudodontini

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
110135335--
  +----------- Chamberlainiini
  |                           
--+  +-------- Rectidentini   
  |  |                        
  +--+     +-- Schepmania **     
     |  +--+                  
     |  |  +-- Pseudodontini  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
110135335--
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmania **     
           +--+                  
              +-- Pseudodontini

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
110147425--
  +-------------- Rectidentini   
--+                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmania **     
           +--+                  
              +-- Pseudodontini

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
110144336--
  +----------- Rectidentini 
  |                         
--+        +-- Schepmania **   
  |  +-----+                
  |  |     +-- Pseudodontini
  +--+                      
     |  +----- Caudiculatus 
     +--+                   
        |  +-- Lamprotulini 
        +--+                
           +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
110135335--
  +-------------------------- Chamberlainiini
  |                                          
--+  +----------------------- Rectidentini   
  |  |                                       
  |  |                    +-- Schepmania **     
  +--+  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+                 +-- Inversidens    
        |  +--------------+                  
        |  |              +-- Lamprotulini   
        +--+                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
110149493--
     +-- Lamprotulini 
  +--+                
  |  +-- Gonideini    
--+                   
  |  +-- Schepmania **   
  +--+                
     +-- Pseudodontini

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
110146415--
  +-------------- Rectidentini   
--+                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmania **     
           +--+                  
              +-- Pseudodontini

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
110158585--
           +-- Chamberlainiini
  +--------+                  
  |        +-- Rectidentini   
--+                           
  |  +-------- Lamprotulini   
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmania **     
        +--+                  
           +-- Pseudodontini

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
1340135335--
  +----------------------------- Schepmania_nieuwenhuisi__1 **
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2 **
--+                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
  +--+  +----------------------- Rectidentini              
     |  |                                                  
     +--+  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus
 
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