The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Bivalvia | Unionoida | Unionoidea | Unionidae subfamily UNIONINAE Rafinesque, 1820: cladomics
5 tribes Cladomics Summarynumber of species = 216 (269 total)
number of subtaxa (rank: tribe) = 5
analyzed in 394 cladograms in 217 publications
maximum number of terminals in an analysis =
72 (mean = 10.76, median = 6)
maximum number of subtaxa (tribe) =
5 (mean = 2.02, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 347
maximum number of included terminals = 72, (mean =
12.09 median = 8)
maximum number of subtaxa (tribe) =
5 (mean = 2.16, median = 2)
proportion of cladograms that support basic monophyly
= 0.92 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 164
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 72 (mean = 16.11, median = 12)
maximum number of subtaxa (tribe) =
5 (mean = 3.12, median = 3)
proportion of cladograms that support rigorous monophyly = 0.87
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 46 | 46 | 44 | 5 | 49 | 38 | 3 | 0 | +-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. 2 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 45 | 45 | 36 | 5 | 154 | 115 | 3 | 50 | +-------------------- Parreysiinae
|
| +-------- Acuticostini **
| |
--+ +--------+ +----- Lepidodesmini **
| | | |
| | +--+----- Unionini **
| | |
| | | +-- Lanceolariini **
+--+ +--+
| +-- Anodontini **
|
| +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 72 | 72 | 70 | 4 | 4 | 2 | 1 | 40 | +----------- Ambleminae
--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 70 | 70 | 58 | 5 | 51 | 40 | 3 | 65 | +----------------- Parreysiinae
|
| +----- Acuticostini **
| |
| +----- Lepidodesmini **
| +--------+
--+ | +----- Unionini **
| | |
| | | +-- Lanceolariini **
| | +--+
| | +-- Anodontini **
| |
+--+ +-- Lens
| |
| +-- Ensidens
| |
| +--------+-- Physunio
| | |
| | +-- Rectidens
| | |
+--+ +-- Yaukthwa
|
| +-------- Ambleminae
| |
+--+ +-- Chamberlainiini
| +--+
| | +-- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 51 | 51 | 50 | 4 | 2 | 0 | 0 | 27 | +-------- Lepidodesmini **
--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 46 | 46 | 43 | 5 | 45 | 39 | 3 | 13 | +-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 46 | 46 | 43 | 5 | 45 | 39 | 3 | 13 | +-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 46 | 46 | 42 | 5 | 49 | 38 | 3 | 16 | +-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 45 | 45 | 22 | 5 | 154 | 115 | 3 | 53 | +-------------- Parreysiinae
|
| +----------- Chamberlainia
| |
| +----------- Megalonaias
| |
| +----------- Uniomerus
| |
| +----------- Quadrulina
| |
| +----------- Sinohyriopsis
| |
| +----------- Lamprotulini
| |
| +----------- Gonideini
| |
| +----------- Pseudodontini
| |
| | +-- Rectidentini
--+ +--------+
| | +-- Contradentini
| |
| | +-------- Amblema
| | |
| | +-------- Toxolasma
| | |
| | +-------- Pleurobemini
| | |
| | | +-- Obliquaria
| | +-----+
| | | +-- Plectomerus
| +--+
| | | +----- Ellipsaria
| | | |
| | | +----- Potamilus_leptodon
| | | |
| | | +----- Medionidus
| | | |
| | | +----- Potamilus_alatus
+--+ | |
| | +----- Ptychobranchus
| +--+
| +----- Truncilla
| |
| +----- Venustaconcha
| |
| +----- Cambarunio
| |
| +----- Leaunio
| |
| | +-- Ortmanniana
| +--+
| +-- Lampsilis
|
| +-------- Acuticostini **
| |
| +-------- Alasmidonta **
| |
| +-------- Anodonta_anatina **
| |
| +-------- Anemina **
| |
| +-------- Anodonta_cygnea **
| |
| +-------- Cristaria **
| |
| +-------- Anodonta_nuttalliana **
| |
| +-------- Lasmigona **
| |
| +-------- Lepidodesmini **
| |
| +-------- Pseudanodonta **
| |
| +-------- Pyganodon **
| |
+--+-------- Simpsonaias **
|
+-------- Strophitus **
|
+-------- Schistodesmus **
|
+-------- Utterbackia **
|
+-------- Lanceolariini **
|
+-------- Unio **
|
+-------- Aculamprotula **
|
| +-- Pletholophus **
+-----+
| +-- Sinanodonta **
|
| +----- Cuneopsis_celtiformis **
| |
| +----- Tchangsinaia **
| |
+--+----- Cuneopsis_rufescens **
|
+----- Nodularia **
|
| +-- Cuneopsis_szechenyii **
+--+
+-- Cuneopsis_heudei **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 44 | 44 | 41 | 5 | 153 | 114 | 3 | 50 | +----------------------- Parreysiinae
|
--+ +-------------------- Pseudodontini
| |
| | +-- Rectidentini
+--+ +--------------+
| | +-- Contradentini
| |
| | +-------- Chamberlainiini
+--+ +-----+
| | | +----- Ambleminae
| | +--+
| | | +-- Lamprotulini
+--+ +--+
| +-- Gonideini
|
| +----------- Acuticostini **
+--+
| +-------- Unionini **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 44 | 63 | 61 | 5 | 44 | 35 | 2 | 47 | +----------------------------- Ambleminae
|
| +-------------- Chamberlainiini
| |
| +-----------+ +-- Contradentini
--+ | | +--------+
| | | | +-- Rectidentini
| | +--+
| | | +-------- Lamprotulini
| | +--+
| | | +----- Pseudodontini
+--+ +--+
| | +-- Inversiunio **
| +--+
| +-- Gonideini
|
| +----------------------- Lepidodesmini **
| |
| | +----- Anodontini **
+--+ +--------------+
| | | +-- Acuticostini **
| | +--+
+--+ +-- Lanceolariini **
|
| +----------------- Globunio **
| |
+--+ +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Pseudocuneopsis **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Whelan et al., 2023 [Publ 4287], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 41 | 41 | 42 | 1 | 0 | 0 | 0 | 67 | Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 38 | 38 | 36 | 5 | 0 | 0 | 0 | 24 | +----------- Lepidodesmini **
--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 36 | 36 | 30 | 5 | 9 | 7 | 3 | 54 | +-- Gonideinae
+-----------------+
| +-- Ambleminae
|
--+ +----------------- Parreysiinae
| |
| | +-- Anemina **
+--+ +-----------+
| | +-- Amuranodonta **
| |
| | +----- Pletholophus **
+--+ +-----+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +-- Alasmidontina **
| +-----+
| | +-- Anodontina **
| |
+--+ +----- Lanceolaria_gladiola **
| |
| +----- Lanceolaria_oxyrhyncha **
| |
+--+----- Cristaria **
|
| +-- Acuticostini **
| |
+--+-- Lepidodesmini **
|
+-- Unionini **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 36 | 39 | 32 | 5 | 9 | 7 | 3 | 58 | +----------------------- Gonideinae
|
--+ +-------------------- Ambleminae
| |
| | +----------------- Parreysiinae
+--+ |
| | +-- Anemina **
| +--------------+
| | +-- Amuranodonta **
+--+
| +----- Pletholophus **
| +--------+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +----------- Cristaria **
| |
| | +-- Alasmidontina **
+--+--------+
| +-- Anodontina **
|
| +-------- Lanceolariini **
+--+
| +----- Lepidodesmini **
+--+
| +-- Acuticostini **
+--+
+-- Unionini **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 35 | 35 | 37 | 4 | 33 | 10 | 3 | 43 | +----------------------- Parreysiinae
|
| +-- Ambleminae
--+ +-----------------+
| | +-- Gonideinae
| |
+--+ +-- Anemina **
| +--------------+
| | +-- Buldowskia **
| |
+--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2025 [Publ 4442], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 35 | 35 | 33 | 2 | 0 | 0 | 0 | 67 | +-- Unionini **
--+
+-- Anodontini **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 35 | 35 | 33 | 5 | 38 | 29 | 2 | 19 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 35 | 35 | 37 | 4 | 33 | 10 | 3 | 43 | +----------------------- Parreysiinae
|
| +-------- Chamberlainiini
| +-----------+
--+ | | +----- Ambleminae
| | +--+
| | | +-- Lamprotulini
| | +--+
+--+ +-- Gonideini
|
| +-- Anemina **
| +--------------+
| | +-- Buldowskia **
| |
+--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 35 | 35 | 36 | 4 | 33 | 10 | 3 | 43 | +----------------------- Parreysiinae
|
| +-- Ambleminae
--+ +-----------------+
| | +-- Gonideinae
| |
+--+ +-- Anemina **
| +--------------+
| | +-- Buldowskia **
| |
+--+ +----- Pletholophus **
| +--------+
| | | +-- Beringiana **
| | +--+
+--+ +-- Sinanodonta **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 28 | 5 | 20 | 18 | 2 | 40 | +----------------- Quadrulini
|
+----------------- Pleurobemini
|
| +-- Amblemini
--+--------------+
| +-- Lampsilini
|
| +-- Lamprotulini
| +-----------+
| | +-- Gonideini
+--+
| +----------- Chamberlainiini
| |
+--+ +-------- Lepidodesmini **
| |
+--+-------- Unionini **
|
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 31 | 5 | 40 | 29 | 2 | 23 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 31 | 5 | 40 | 29 | 2 | 25 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 31 | 5 | 23 | 22 | 3 | 51 | +-------------- Parreysiinae
|
--+ +-- Ambleminae
| +--------+
| | +-- Gonideinae
+--+
| +-------- Unionini **
| |
+--+ +-- Lepidodesmini **
| +--+
| | +-- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 32 | 32 | 30 | 5 | 36 | 25 | 2 | 24 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 31 | 31 | 36 | 5 | 14 | 12 | 1 | 54 | +----------------- Gonideinae
|
--+ +-------------- Anemina **
| |
+--+ +----------- Sinanodonta **
| |
+--+ +-------- Acuticostini **
| |
+--+ +-- Lepidodesmini **
| +--+
| | +-- Cristaria **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 31 | 31 | 27 | 5 | 9 | 7 | 3 | 54 | +-- Gonideinae
+-----------+
| +-- Ambleminae
--+
| +----------- Parreysiinae
| |
+--+ +-------- Acuticostini **
| |
+--+ +----- Lepidodesmini **
| |
+--+----- Unionini **
|
| +-- Lanceolariini **
+--+
+-- Anodontini **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 31 | 31 | 23 | 5 | 9 | 7 | 3 | 55 | +-- Gonideinae
+--------------+
| +-- Ambleminae
|
--+ +-------------- Parreysiinae
| |
| | +-- Anemina **
+--+ +--------+
| | +-- Amuranodonta **
| |
| | +----- Pletholophus **
+--+ +--+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
| |
+--+ +----- Lanceolaria_gladiola **
| |
| +----- Acuticostini **
| |
| +----- Lepidodesmini **
| |
+--+----- Lanceolaria_oxyrhyncha **
|
+----- Cristaria **
|
+----- Unionini **
|
| +-- Alasmidontina **
+--+
+-- Anodontina **
Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 29 | 29 | 26 | 2 | 2 | 0 | 0 | 61 | +-- Unionini **
--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 29 | 29 | 27 | 5 | 14 | 12 | 1 | 54 | +-- Lamprotulini
+--------------------------------------+
| +-- Gonideini
--+
| +-------------------------------------- Chamberlainiini
| |
+--+ +----------------------------------- Anemina **
| |
+--+ +-------------------------------- Acuticostini **
| |
+--+ +----------------------------- Sinanodonta **
| |
+--+ +-------------------------- Aculamprotula **
| |
+--+ +-- Cristaria **
| +--------------------+
| | +-- Lanceolariini **
+--+
| +-------------------- Lepidodesmini **
+--+
| +----------------- Schistodesmus **
+--+
| +-------------- Nodularia **
+--+
| +----------- Cuneopsis_rufescens **
+--+
| +-------- Tchangsinaia **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_szechenyii **
+--+
+-- Cuneopsis_heudei **
Lyubas et al., 2022 [Publ 4255], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 19 | 2 | 3 | 3 | 2 | 60 | +-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Whelan et al., 2023 [Publ 4287], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 27 | 1 | 0 | 0 | 0 | 65 | Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 26 | 2 | 0 | 0 | 0 | 52 | +-- Lepidodesmini **
--+
+-- Unionini **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 25 | 3 | 52 | 40 | 3 | 47 | +----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Lyubas et al., 2022 [Publ 4255], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 25 | 2 | 3 | 3 | 2 | 60 | +-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 26 | 2 | 0 | 0 | 0 | 52 | +-- Lepidodesmini **
--+
+-- Unionini **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 3 | 112 | 87 | 3 | 50 | +-------------------- Parreysiinae
|
| +-- Rectidentini
--+ +--------------+
| | +-- Contradentini
| |
+--+ +-------------- Pseudodontini
| |
| | +----- Unionini **
+--+ +-----+
| | | +-- Lanceolariini **
| | +--+
+--+ +-- Anodontini **
|
| +-------- Ambleminae
+--+
| +----- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Lamprotulini
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 2 | 3 | 98 | 98 | 3 | 55 | +----------------- Parreysiinae
|
| +-------------- Pseudodontini
| |
--+ | +-- Rectidentini
| | +--------+
| | | +-- Contradentini
| | |
| | | +----- Ambleminae
+--+ | |
| | | +-- Chamberlainia
| | | |
| | | +-- Sinohyriopsis
| | | |
| | | +-- Lamprotula_caveata
| | +--+ |
| | | | +-- Lamprotula_leaii
+--+ | | |
| | | +-- Potomida
| | | |
| | | +-- Pronodularia
| | | |
| | +--+-- Gonidea
| | |
| | +-- Leguminaia_wheatleyi
| | |
| | +-- Leguminaia
| | |
| | +-- Microcondylaea
| | |
+--+ +-- Sinosolenaia_carinata
| |
| +-- Sinosolenaia_oleivora
| |
| +-- Sinosolenaia_recognita
|
| +----- Unionini **
| |
| | +-- Alasmidonta **
| | |
| | +-- Anodonta_anatina **
| | |
| | +-- Anodonta_cygnea **
| | |
| | +-- Anodonta_nuttalliana **
+--+ |
| +-- Lasmigona **
| |
| +-- Pseudanodonta **
| |
| +-- Pyganodon **
| |
| +-- Simpsonaias **
| |
| +-- Strophitus **
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola **
|
+-- Lanceolaria_grayii__1 **
|
+-- Lanceolaria_grayii__2 **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 18 | 3 | 112 | 87 | 3 | 51 | +-------------------- Parreysiinae
|
| +-- Contradentini
| +-----------+
| | +-- Rectidentini
--+ +--+
| | | +----------- Ambleminae
| | +--+
| | | +-------- Pseudodontini
| | +--+
| | | +----- Gonideini
+--+ +--+
| | +-- Chamberlainiini
| +--+
| +-- Lamprotulini
|
| +----------- Unionini **
| |
+-----+ +-------- Lanceolariini **
| |
| +-------- Cristariina **
+--+
| +----- Anodonta_anatina **
| |
| +----- Pseudanodonta **
+--+
+----- Alasmidontina **
|
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_nuttalliana **
Dai et al., 2024 [Publ 4385], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 1 | 4 | 2 | 1 | 63 | +-- Parreysiinae
--+
+-- Anodontini **
Dai et al., 2024 [Publ 4385], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 24 | 1 | 4 | 2 | 1 | 63 | +-- Parreysiinae
--+
+-- Anodontini **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 5 | 22 | 14 | 2 | 26 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Xiang et al., 2025 [Publ 4433], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 1 | 1 | 1 | 1 | 61 | +-- Parreysiinae
--+
+-- Anodontini **
Whelan et al., 2023 [Publ 4287], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 1 | 0 | 0 | 0 | 64 | Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 5 | 25 | 23 | 2 | 33 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 5 | 26 | 23 | 2 | 30 | +-- Ambleminae
+-----------+
| +-- Gonideinae
--+
| +----------- Lepidodesmini **
+--+
| +-------- Lanceolariini **
+--+
| +----- Unionini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 5 | 65 | 48 | 3 | 42 | +-------------------- Parreysiinae
|
| +----------- Acuticostini **
| +-----+
--+ | | +-------- Unionini **
| | +--+
| | | +----- Lanceolariini **
| | +--+
| | | +-- Lepidodesmini **
+--+ +--+
| +-- Anodontini **
|
| +-- Contradentini
| +-----------+
| | +-- Rectidentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 5 | 14 | 11 | 2 | 23 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 3 | 50 | 40 | 2 | 47 | +-------------------- Pseudodontini
|
--+ +-- Rectidentini
| +--------------+
| | +-- Contradentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 5 | 20 | 14 | 2 | 25 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 25 | 5 | 14 | 14 | 2 | 36 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 3 | 50 | 40 | 2 | 45 | +-------------------- Pseudodontini
|
--+ +-- Contradentini
| +--------------+
| | +-- Rectidentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 5 | 30 | 20 | 2 | 28 | +-- Gonideinae
+-----------+
| +-- Ambleminae
--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 5 | 65 | 48 | 3 | 40 | +-------------------- Parreysiinae
|
| +-------- Acuticostini **
| |
--+ +--------+ +-- Lepidodesmini **
| | | +--+
| | | | +-- Unionini **
| | +--+
| | | +-- Lanceolariini **
+--+ +--+
| +-- Anodontini **
|
| +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 21 | 4 | 0 | 0 | 0 | 32 | +-------- Anodontini **
--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 3 | 0 | 0 | 0 | 51 | +----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 3 | 0 | 0 | 0 | 51 | +----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 4 | 2 | 2 | 1 | 50 | +-------------- Gonideinae
|
--+ +----------- Acuticostini **
| |
+--+ +-- Lanceolaria_lanceolata **
| +-----+
| | +-- Lanceolaria_grayii **
+--+
| +----- Lanceolaria_gladiola **
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 4 | 22 | 19 | 2 | 27 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 15 | 4 | 2 | 2 | 1 | 50 | +----------- Gonideinae
|
--+ +-------- Acuticostini **
| |
+--+ +----- Lanceolaria_grayii **
| |
| +----- Lanceolaria_gladiola **
+--+
+----- Lanceolaria_lanceolata **
|
| +-- Anodontini **
+--+
+-- Unionini **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 4 | 26 | 23 | 2 | 27 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 4 | 22 | 19 | 2 | 27 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 5 | 5 | 12 | 8 | 1 | 60 | +-------- Gonideinae
|
| +----- Lanceolaria_grayii__1 **
| |
| +----- Lanceolaria_grayii__2 **
| |
| +----- Cristaria_plicata__1 **
| |
| +----- Cristaria_plicata__2 **
| |
| +----- Sinanodonta_lauta **
| |
| +----- Beringiana_japonica **
--+ |
| +----- Beringiana_fukuharai__1 **
| |
| +----- Beringiana_beringiana **
| |
| +----- Beringiana_fukuharai__2 **
| |
| +----- Beringiana_fukuharai__3 **
| |
| +----- Buldowskia_iwakawai__1 **
| |
| +----- Buldowskia_iwakawai__2 **
| |
| +----- Sinanodonta_tumens__1 **
+--+
+----- Sinanodonta_tumens__2 **
|
+----- Pletholophus **
|
+----- Anemina **
|
+----- Lanceolaria_lanceolata **
|
+----- Cristaria_plicata__3 **
|
+----- Lanceolaria_gladiola **
|
+----- Sinanodonta_angula **
|
+----- Lepidodesmini **
|
| +-- Aculamprotula **
+--+
| +-- Acuticostini **
|
| +-- Cuneopsis **
+--+
| +-- Nodularia **
|
| +-- Schistodesmus **
+--+
+-- Inversiunio **
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 21 | 21 | 19 | 4 | 2 | 2 | 1 | 47 | +----------- Gonideinae
--+
| +-------- Acuticostini **
+--+
| +----- Lanceolariini **
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 21 | 21 | 22 | 5 | 12 | 8 | 1 | 56 | +----------------- Gonideinae
|
--+ +-------------- Anodontini **
| |
| | +-- Tchangsinaia **
+--+ +--------+
| | +-- Nodularia **
| |
+--+ +-- Schistodesmus **
| +-----+
| | +-- Acuticostini **
+--+
| +----- Inversiunio **
+--+
| +-- Lepidodesmini **
+--+
+-- Lanceolariini **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 21 | 21 | 19 | 4 | 17 | 14 | 2 | 29.5 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 0 | 1 | 84 | 83 | 2 | 58 | +----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 2 | 6 | 6 | 1 | 53 | +----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 3 | 17 | 13 | 2 | 49 | +-------- Gonideinae
--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Sano et al., 2022 [Publ 4311], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 1 | 0 | 0 | 0 | 54 | Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 4 | 36 | 34 | 3 | 34 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Parreysiinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Smith et al., 2018 [Publ 3790], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 17 | 1 | 0 | 0 | 0 | 57 | Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 10 | 2 | 0 | 0 | 0 | 55 | +----------------- Anodonta_cygnea **
|
+----------------- Pyganodon **
|
+----------------- Alasminota **
|
+----------------- Anodonta_anatina **
|
+----------------- Unionini **
|
+----------------- Cristariina **
|
| +-- Anodonta_nuttalliana **
--+--------------+
| +-- Anodonta_californiensis **
|
| +-- Pseudodontoideus **
| |
+--------------+-- Alasmidonta **
| |
| +-- Lasmigona **
|
| +-------------- Pseudanodonta **
| |
| | +----- Strophitus **
| +--------+
+--+ | +-- Prolasmidonta **
| +--+
| +-- Platynaias **
|
| +----------- Anodonta_oregonensis__4 **
+--+
| +-------- Anodonta_oregonensis__3 **
+--+
| +----- Anodonta_oregonensis__2 **
+--+
| +-- Anodonta_oregonensis__1 **
+--+
+-- Anodonta_kennerlyi **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 2 | 24 | 14 | 2 | 33 | +-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 14 | 11 | 2 | 35 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 13 | 11 | 2 | 35 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 2 | 12 | 11 | 2 | 36 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 3 | 0 | 0 | 0 | 34 | +----- Anodontini **
--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 13 | 2 | 1 | 0 | 0 | 50 | +-- Anodontini **
--+
+-- Unionini **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 16 | 3 | 66 | 66 | 3 | 45 | +----------------- Parreysiinae
|
| +-------------- Pseudodontini
--+ |
| | +-- Rectidentini
| | +--------+
+--+ | +-- Contradentini
| |
| | +----- Unionini **
| | +--+
| | | | +-- Lanceolariini **
+--+ | +--+
| | +-- Anodontini **
| |
| | +-- Lamprotula
| | |
| | +-- Potomida
+--+ |
| +-- Pronodularia
| |
| +-- Gonidea
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
+-----+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
+-- Cambarunio
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 16 | 2 | 5 | 3 | 1 | 47 | +----------- Gonideinae
--+
| +-------- Aculamprotula **
+--+
| +----- Anodontini **
+--+
| +-- Schistodesmus **
+--+
+-- Nodularia **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 5 | 36 | 34 | 2 | 34 | +----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 4 | 14 | 14 | 2 | 32 | +-- Ambleminae
+--------+
| +-- Gonideinae
--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 3 | 14 | 14 | 2 | 34 | +-- Ambleminae
+-----+
| +-- Gonideinae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 3 | 2 | 0 | 0 | 39 | +----- Anodontini **
--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 2 | 0 | 0 | 0 | 51 | +-- Unionini **
--+
+-- Anodontini **
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 5 | 5 | 3 | 1 | 51 | +-------------- Parreysiinae
|
--+ +----------- Anodontina **
| |
+--+ +-- Acuticostini **
| +-----+
| | +-- Cristariina **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 5 | 22 | 20 | 3 | 40 | +-- Ambleminae
+--------------------+
| +-- Gonideinae
--+
| +-------------------- Parreysiinae
| |
+--+ +----------------- Anodontini **
| |
| | +-- Acuticostini **
+--+ +-----------+
| | +-- Lanceolariini **
| |
+--+ +-- Lepidodesmini **
| +--------+
| | +-- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Zanatta et al., 2007 [Publ 2698], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 10 | 1 | 0 | 0 | 0 | 54 | Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 15 | 2 | 22 | 17 | 1 | 42 | +-- Unionini **
--+
+-- Anodontini **
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 11 | 5 | 5 | 3 | 1 | 51 | +-------------------- Parreysiinae
|
| +----------------- Acuticostini **
--+ |
| +----------------- Cristariina **
| |
| +----------------- Anodontina **
+--+
| +-------------- Lanceolariini **
| |
| | +-- Aculamprotula **
| +-----------+
+--+ +-- Lepidodesmini **
|
| +----------- Unio **
| |
+--+ +-------- Middendorffinaia **
| |
+--+ +----- Cuneopsis **
| |
+--+----- Nodularia **
|
| +-- Inversiunio **
+--+
+-- Schistodesmus **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 17 | 15 | 4 | 1 | 1 | 1 | 37 | +-- Gonideinae
+--------------+
| +-- Lepidodesmini **
--+
| +-------------- Aculamprotula **
| |
+--+ +-- Acuticostini **
| +--------+
| | +-- Lanceolariini **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 3 | 0 | 0 | 0 | 40 | +----- Aculamprotula **
+--------+
| | +-- Acuticostini **
| +--+
--+ +-- Lanceolariini **
|
| +----------- Unio **
| |
+--+ +-------- Pseudocuneopsis **
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Zanatta et al., 2007 [Publ 2698], fig. 1B iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 13 | 1 | 0 | 0 | 0 | 54 | Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 5 | 21 | 19 | 3 | 37 | +-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Hoeh, 1990 [Publ 2717], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 1 | 0 | 0 | 0 | 53 | Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 34 | 32 | 3 | 19 | 18 | 2 | 72 | +----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Schistodesmus **
| +-----+
| | | +-- Lepidodesmini **
--+ | +--+
| | +-- Aculamprotula **
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina **
|
| +-- Utterbackia_peninsularis **
| +-----+
| | +-- Potamilus_alatus
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosulinae
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Hoeh, 1990 [Publ 2717], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 2 | 1 | 0 | 0 | 0 | 55 | Hoeh, 1990 [Publ 2717], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 1 | 0 | 0 | 0 | 53 | Froufe et al., 2016 [Publ 3738], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 1 | 0 | 0 | 0 | 52 | Lopes-Lima et al., 2024 [Publ 4376], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 1 | 0 | 0 | 0 | 54 | Prié & Puillandre, 2014 [Publ 3529], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 2 | 3 | 2 | 2 | 50 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Inoue et al., 2014 [Publ 3635], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 13 | 1 | 6 | 5 | 2 | 51 | +-------- Amblemini
|
+-------- Pleurobemini
--+
+-------- Lampsilini
|
| +----- Quadrulini
+--+
| +-- Gonideinae
+--+
+-- Anodontini **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 15 | 12 | 4 | 1 | 1 | 1 | 48 | +-- Lanceolariini **
+-----+
| +-- Unionini **
--+
| +----- Acuticostini **
+--+
| +-- Anodontini **
+--+
+-- Gonideinae
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 5 | 5 | 20 | 18 | 2 | 47 | +-- Ptychorhynchus
|
+-- Chamberlainiini
|
+--------+-- Lamprotulini
| |
| +-- Sinosolenaia
| |
| +-- Ambleminae
|
| +-------- Strophitus **
--+ |
| +-------- Schistodesmus **
| |
| +-------- Lepidodesmini **
| |
| +-------- Acuticostini **
| |
| +-------- Cristariina **
+--+
+-------- Lanceolariini **
|
| +-- Nodularia **
+-----+
| +-- Tchangsinaia **
|
| +----- Utterbackia **
| |
+--+----- Utterbackiana **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 5 | 20 | 18 | 2 | 50 | +----------- Sinosolenaia
+--+
| | +-------- Lamprotulini
| +--+
| | +----- Chamberlainiini
| +--+
| | +-- Ptychorhynchus
--+ +--+
| +-- Ambleminae
|
| +----------- Anodontini **
| |
| | +----- Acuticostini **
+--+ +--+
| | | +-- Nodularia **
| | +--+
+--+ +-- Tchangsinaia **
|
| +----- Lanceolariini **
+--+
| +-- Schistodesmus **
+--+
+-- Lepidodesmini **
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 19 | 17 | 4 | 8 | 5 | 1 | 52 | +----------------- Anodontini **
|
| +----------- Chamberlainiini
| +--+
--+ | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Lamprotula_caveata
| | +--+
| | | +-- Gibbosulinae
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +----- Aculamprotula **
| +--+
| | | +-- Tchangsinaia **
| | +--+
+-----+ +-- Nodularia **
|
| +----- Schistodesmus **
+--+
| +-- Lanceolariini **
+--+
+-- Acuticostini **
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 1 | 0 | 0 | 0 | 51 | Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 14 | 3 | 8 | 4 | 1 | 51 | +-------- Gonideinae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Douda et al., 2024 [Publ 4364], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 12 | 1 | 1 | 0 | 0 | 52 | Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 2 | 9 | 9 | 2 | 38 | +-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Hou et al., 2024 [Publ 4400], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 1 | 2 | 2 | 1 | 49 | +-- Gonideinae
--+
+-- Unionini **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 3 | 28 | 8 | 2 | 37 | +-------------- Ambleminae
|
--+ +----------- Gonideinae
| |
+--+ +-------- Aculamprotula **
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Unio **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 3 | 28 | 8 | 2 | 37 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 1 | 0 | 0 | 0 | 43 | Sano et al., 2022 [Publ 4311], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 12 | 1 | 0 | 0 | 0 | 52 | Xiang et al., 2025 [Publ 4433], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 2 | 1 | 1 | 1 | 38 | +----- Ambleminae
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wu et al., 2024 [Publ 4379], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 1 | 0 | 0 | 0 | 53 | Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 11 | 1 | 25 | 20 | 2 | 48 | +-------------- Quadrulina
|
| +----- Lamprotulini
--+ +-----+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Anodontini **
|
| +-------- Megalonaias
+--+
| +----- Pleurobemini
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 11 | 1 | 0 | 0 | 0 | 50 | Froufe et al., 2017 [Publ 3802], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 9 | 1 | 0 | 0 | 0 | 47 | Bogan et al., 2023 [Publ 4295], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 9 | 1 | 1 | 0 | 0 | 51 | Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 2 | 9 | 9 | 2 | 39 | +-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 4 | 2 | 2 | 1 | 45 | +----------- Gonideinae
--+
| +-------- Anodontini **
+--+
| +----- Unionini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 3 | 1 | 1 | 1 | 53 | +----------------- Parreysiinae
|
--+ +-------------- Unio **
| |
| | +----- Aculamprotula **
+--+ +-----+
| | | +-- Anodontini **
| | +--+
+--+ +-- Lepidodesmini **
|
| +-------- Middendorffinaia **
+--+
| +----- Inversiunio **
+--+
| +-- Cuneopsis **
+--+
+-- Nodularia **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 8 | 2 | 5 | 2 | 1 | 46 | +----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 8 | 1 | 2 | 0 | 0 | 51 | Hou et al., 2025 [Publ 4401], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 1 | 0 | 0 | 0 | 53 | Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 3 | 20 | 18 | 3 | 40 | +----------- Parreysiinae
|
--+ +-- Ambleminae
| +-----+
| | +-- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Unionini **
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 3 | 4 | 4 | 2 | 44 | +-- Gonideinae
+-----+
| +-- Ambleminae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 2 | 8 | 6 | 1 | 41 | +----- Ambleminae
--+
| +-- Lepidodesmini **
+--+
+-- Anodontini **
Hou et al., 2025 [Publ 4432], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 2 | 0 | 0 | 0 | 46 | +-- Lepidodesmini **
--+
+-- Unionini **
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 2 | 15 | 9 | 2 | 40 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Unio **
+--+
| +-- Aculamprotula **
+--+
+-- Anodontini **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 12 | 7 | 2 | 40 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 2 | 2 | 2 | 1 | 36 | +----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 2 | 5 | 3 | 2 | 43 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 5 | 1 | 23 | 21 | 1 | 51.5 | +-- Anodontini **
--+
+-- Ambleminae
Lopes-Lima et al., 2020 [Publ 4080], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 2 | 2 | 0 | 0 | 48 | +-- Unionini **
--+
+-- Lanceolariini **
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 2 | 15 | 9 | 2 | 40 | +-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 13 | 10 | 2 | 40 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 7 | 3 | 10 | 8 | 2 | 39 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 7 | 2 | 1 | 0 | 0 | 44 | +-- Anodontini **
--+
+-- Unionini **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 1 | 0 | 0 | 0 | 50 | Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 12 | 7 | 2 | 40 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 0 | 0 | 0 | 38 | +----- Lanceolariini **
--+
| +-- Anodontini **
+--+
+-- Unionini **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 11 | 10 | 2 | 41 | +----- Anodontini **
--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 18 | 11 | 3 | 9 | 8 | 2 | 54 | +----------- Cristariina **
|
| +-------- Lanceolariini **
| |
--+ +-------- Anodonta **
| |
| +-------- Unionini **
| |
+--+ +-- Alasmidontina **
+-----+
| +-- Pseudanodonta **
|
| +----- Ambleminae
| |
+--+ +-- Velesunioninae
| |
+--+-- Lamprotulini
|
+-- Chamberlainiini
Khalloufi et al., 2011 [Publ 2748], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 1 | 2 | 2 | 2 | 47 | +----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 4 | 4 | 4 | 1 | 48 | +-------------- Gonideinae
|
--+ +----------- Anodontini **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Cuneopsis **
+--+
| +----- Lanceolaria_grayii **
| |
+--+----- Acuticostini **
|
| +-- Lanceolaria_gladiola **
+--+
+-- Aculamprotula **
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 1 | 29 | 29 | 1 | 51 | +-- Anodontini **
--+
+-- Ambleminae
Lopes-Lima et al., 2020 [Publ 4080], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 1 | 2 | 0 | 0 | 51 | Reis et al., 2013 [Publ 3580], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 2 | 5 | 3 | 2 | 45 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 2 | 0 | 0 | 0 | 45 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 2 | 16 | 16 | 3 | 53 | +-------------------------------- Parreysiinae
|
| +-------------- Nodularia **
| |
| +--------------+ +----------- Anodonta_cygnea **
--+ | | |
| | +--+ +-------- Unio_pictorum__2 **
| | | |
| | +--+ +-- Unio_pictorum__1 **
| | | +--+
| | | | +-- Unio_caffer **
+--+ +--+
| | +-- Anodonta **
| +--+
| +-- Alasmidontina **
|
| +-------------------------- Lamprotulini
| |
+--+ +----------------------- Quadrulini
| |
+--+ +-------------------- Gonideini
| |
| | +----- Eurynia
+--+ +-----------+
| | | +-- Cambarunio
| | +--+
| | +-- Ptychobranchus
+--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema
+--+
| +----------- Truncilla
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma
+--+
| +----- Ligumia
+--+
| +-- Obliquaria
+--+
+-- Ortmanniana
Soroka, 2010 [Publ 3404], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 5 | 2 | 0 | 0 | 0 | 45 | +----- Anodonta_anatina **
|
+----- Anodonta_cygnea **
--+
+----- Pseudanodonta **
|
| +-- Cristariina **
+--+
+-- Unionini **
King et al., 1999 [Publ 2699], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 1 | 0 | 0 | 0 | 49 | Araujo et al., 2009 [Publ 2589], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 7 | 2 | 3 | 2 | 2 | 47 | +-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 18 | 16 | 3 | 10 | 10 | 2 | 55 | +----------------- Sinanodonta **
|
--+ +-------------- Cristaria **
| |
+--+ +-- Gonideinae
| +--------+
| | +-- Ambleminae
+--+
| +-------- Aculamprotula **
+--+
| +----- Acuticostini **
+--+
| +-- Nodularia **
+--+
+-- Cuneopsis **
Marrone et al., 2019 [Publ 3907], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 1 | 0 | 0 | 0 | 47 | Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 2 | 5 | 5 | 1 | 43 | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 8 | 3 | 4 | 4 | 1 | 49 | +----------- Gonideinae
--+
| +-------- Anodontini **
+--+
| +----- Nodularia **
+--+
| +-- Lanceolariini **
+--+
+-- Inversiunio **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 25 | 19 | 2 | 22 | 17 | 1 | 63 | +----- Unionini **
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Kondakov et al., 2018 [Publ 3914], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 0 | 0 | 0 | 48 | Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 2 | 9 | 7 | 2 | 40 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 2 | 0 | 0 | 0 | 46 | +-- Unionini **
--+
+-- Anodontini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 2 | 0 | 0 | 50 | Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 2 | 3 | 2 | 2 | 50 | +----------------- Ambleminae
|
--+ +-------------- Gonideinae
| |
+--+ +----------- Nodularia **
| |
| | +-- Unio_gibbus **
+--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodontini **
+--+
| +----- Unio_pictorum__1 **
| |
| +----- Unio_pictorum__2 **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 25 | 23 | 2 | 22 | 17 | 1 | 63 | +----- Unionini **
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Tomilova et al., 2020 [Publ 4097], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 2 | 0 | 0 | 0 | 43 | +-- Unionini **
|
--+-- Cristariina **
|
+-- Anodontina **
Vikhrev et al., 2017 [Publ 3810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 2 | 0 | 0 | 48 | Klishko et al., 2018 [Publ 3945], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 1 | 0 | 0 | 0 | 44 | Bespalaya et al., 2018 [Publ 3811], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 7 | 1 | 2 | 0 | 0 | 49 | Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 3 | 49 | 42 | 3 | 42 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Parreysiinae
+--+
| +----- Lanceolariini **
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 3 | 49 | 42 | 3 | 42 | +----------- Ambleminae
|
--+----------- Gonideinae
|
| +-------- Parreysiinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 2 | 0 | 0 | 0 | 45 | +-------- Anodonta_anatina **
|
--+ +----- Cristariina **
| |
+--+----- Unionini **
|
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Bauer et al., 2021 [Publ 4365], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 1 | 0 | 0 | 0 | 48 | Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 12 | 11 | 2 | 39 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 0 | 0 | 0 | 45 | +-- Anodonta_anatina **
+-----+
| +-- Unionini **
--+
| +----- Cristariina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Choi et al., 2020 [Publ 4169], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 3 | 3 | 1 | 49 | +-- Parreysiinae
--+
+-- Unionini **
Nagel et al., 1998 [Publ 1946], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 2 | 2 | 1 | 41 | +-------- Gonideini
--+
| +----- Lamprotulini
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 9 | 8 | 2 | 39 | +-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Inoue et al., 2014 [Publ 3635], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 2 | 1 | 1 | 1 | 1 | 50 | +-- Ambleminae
--+
+-- Anodontini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 2 | 0 | 0 | 45 | Soroka, 2010 [Publ 3404], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 4 | 2 | 0 | 0 | 0 | 44 | +----- Cristariina **
--+
| +-- Unionini **
+--+
+-- Anodontina **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 0 | 0 | 0 | 47 | Zieritz et al., 2012 [Publ 3673], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 8 | 4 | 2 | 1 | 0 | 0 | 45 | +----- Anodontina **
|
--+ +-- Cristariina **
| |
+--+-- Margaritiferinae
|
+-- Unionini **
Araujo et al., 2009 [Publ 2589], fig. 7C iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 3 | 2 | 2 | 46 | +-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 12 | 11 | 2 | 39 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Froufe et al., 2017 [Publ 3802], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 3 | 2 | 1 | 41 | +----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Hoeh et al., 1995 [Publ 3391], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 4 | 1 | 0 | 0 | 0 | 47 | Smith et al., 2023 [Publ 4291], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 0 | 0 | 0 | 52 | Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 16 | 15 | 2 | 39 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 12 | 10 | 2 | 7 | 5 | 2 | 50 | +-- Anodontina **
+--------+
| +-- Utterbackia **
|
--+ +-- Gonideinae
| +-----+
| | +-- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Pyganodon **
+--+
+-- Cristariina **
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 9 | 7 | 2 | 43 | +-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Anodontini **
+--+
| +----- Aculamprotula **
+--+
| +-- Lepidodesmini **
+--+
+-- Unio **
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 9 | 7 | 3 | 4 | 3 | 1 | 51 | +-------- Anodontini **
--+
| +----- Gonideinae
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Choi et al., 2020 [Publ 4169], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 1 | 0 | 0 | 0 | 47 | Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 2 | 1 | 29 | 29 | 1 | 47 | +-- Anodontini **
--+
+-- Ambleminae
Soroka, 2010 [Publ 3404], fig. 8a iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 40 | +-- Unionini **
--+
+-- Anodontini **
Klishko et al., 2017 [Publ 3946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 0 | 0 | 0 | 44 | Lopes-Lima et al., 2020 [Publ 4080], fig. 12 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 2 | 0 | 0 | 50 | Hewitt et al., 2018 [Publ 3758], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 1 | 3 | 2 | 1 | 48 | +-- Ambleminae
--+
+-- Anodontini **
Dai et al., 2024 [Publ 4358], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 0 | 0 | 0 | 47 | Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 40 | 18 | 3 | 48 | +----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 41 | 19 | 3 | 48 | +-- Unionini **
+--------+
| +-- Anodontini **
--+
| +-------- Contradentini
+--+
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 21 | 18 | 1 | 48 | +-- Anodontini **
--+
+-- Ambleminae
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 7 | 7 | 2 | 40 | +-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Soroka, 2010 [Publ 3404], fig. 3a-M iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 46 | +----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini **
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 0 | 0 | 0 | 43 | King et al., 1999 [Publ 2699], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 1 | 0 | 0 | 0 | 48 | Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 16 | 14 | 3 | 10 | 10 | 2 | 53 | +-------- Anodontini **
|
--+ +-- Lanceolariini **
| +--+
| | +-- Unionini **
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Soroka, 2010 [Publ 3404], fig. 8b iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 42 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 2 | 0 | 0 | 46 | Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 40 | 18 | 3 | 48 | +----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 7 | 7 | 2 | 40 | +-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Källersjö et al., 2005 [Publ 2684], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 2 | 2 | 1 | 0 | 0 | 44 | +-- Margaritiferinae
|
--+-- Anodontini **
|
+-- Unionini **
Riccardi et al., 2020 [Publ 4089], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 0 | 0 | 0 | 42 | Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 18 | 1 | 2 | 16 | 12 | 2 | 58.5 | +----- Anodontina **
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula **
|
+-- Unio **
Bogan et al., 2023 [Publ 4295], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 1 | 0 | 0 | 49 | Bolotov et al., 2016 [Publ 3732], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 1 | 2 | 0 | 0 | 49 | Källersjö et al., 2005 [Publ 2684], fig. 1B-F iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 4 | 2 | 1 | 0 | 0 | 45 | +-- Margaritiferinae
|
--+-- Anodontini **
|
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 3b-M iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 44 | +----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 1 | 96 | 94 | 2 | 48 | +-- Chamberlainiini
+-----+
| +-- Anodontini **
--+
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 96 | 94 | 2 | 47 | +-------- Chamberlainiini
|
--+-------- Anodontini **
|
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 41 | 19 | 3 | 48 | +-------- Pseudodontini
|
| +----- Parreysiinae
--+ |
| +----- Contradentini
| |
+--+ +-- Unionini **
+--+
| +-- Anodontini **
|
| +-- Gonideini
+--+
+-- Ambleminae
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 22 | 18 | 1 | 48.5 | +-- Anodontini **
--+
+-- Ambleminae
Prie et al., 2012 [Publ 3487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 1 | 0 | 0 | 45 | Lopes-Lima et al., 2020 [Publ 4080], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 2 | 0 | 0 | 45 | Wu et al., 2020 [Publ 4121], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 29 | 6 | 3 | 43 | +-- Ambleminae
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 15 | 10 | 2 | 26 | 13 | 3 | 53 | +-------------------- Amblemini
|
+-------------------- Parreysiinae
|
+-------------------- Eurynia
|
+-------------------- Fusconaia
|
+-------------------- Gonideinae
|
+-------------------- Pleurobema
|
+-------------------- Quadrulini
--+
| +----------- Ortmanniana
+--------+
| | +-------- Glebula
| +--+
| | +----- Potamilus
| +--+
| | +-- Ligumia
| +--+
| +-- Toxolasma
|
| +----------------- Cyrtonaias
| |
+--+ +-------------- Unionini **
| |
| | +----------- Anodontini **
+--+ |
| | +----- Iridininae
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaeinae
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 6 | 4 | 2 | 41 | +-- Gonideinae
+--+
| +-- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 19 | 12 | 2 | 29 | 13 | 3 | 57 | +-------- Anodontini **
|
--+ +----- Gonideinae
| |
+--+----- Unionini **
|
| +-- Margaritiferinae
+--+
+-- Ambleminae
Bogan et al., 2023 [Publ 4295], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 1 | 0 | 0 | 49 | Soroka, 2010 [Publ 3404], fig. 9a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 0 | 0 | 0 | 48 | Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 26 | 13 | 3 | 45 | +----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Anodontini **
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 6 | 4 | 2 | 41 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 18 | 1 | 2 | 17 | 13 | 2 | 59 | +----- Anodontina **
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unionini **
Nagel et al., 1998 [Publ 1946], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 0 | 0 | 0 | 47 | Wu et al., 2020 [Publ 4147], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 9 | 6 | 2 | 43 | +----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Nagel et al., 1996 [Publ 3400], fig. 351 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 0 | 0 | 0 | 47 | Bashê, 2022 [Publ 4265], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 1 | 0 | 0 | 48 | Amaro et al., 2017 [Publ 3791], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 11 | 8 | 2 | 42 | +-------- Anodontini **
|
--+ +-- Pleurobemini
| +--+
| | +-- Gonideinae
+--+
| +-- Lampsilini
+--+
+-- Quadrulini
Wu et al., 2020 [Publ 4121], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 29 | 6 | 3 | 43 | +-- Ambleminae
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Bogatov et al., 2018 [Publ 3859], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 1 | 0 | 0 | 45 | +-------- Unio_crassus **
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Anodontini **
Bogatov et al., 2018 [Publ 3859], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 1 | 0 | 0 | 45 | +-------- Unio_tumidus **
--+
| +----- Anodontini **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 26 | 13 | 3 | 43 | +-------- Parreysiinae
|
--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideinae
+--+
+-- Ambleminae
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 26 | 13 | 3 | 43 | +----------------------------- Gonideinae
|
--+ +-------------------------- Parreysiinae
| |
+--+ +----------------------- Fusconaia
| |
| +----------------------- Amblemini
+--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema
| |
| | +-------------- Eurynia
+--+ |
| | +----- Cyrtonaias
| +--------+
+--+ | +-- Unionini **
| +--+
| +-- Anodontini **
|
| +----------- Ortmanniana
+--+
| +-------- Glebula
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Toxolasma
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 2 | 26 | 13 | 3 | 44 | +-------- Gonideinae
|
--+ +----- Ambleminae
| |
+--+ +-- Unio_pictorum **
| |
+--+-- Unio_tumidus **
|
+-- Anodontini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 2 | 2 | 1 | 41 | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 45 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Mabuchi et al., 2021 [Publ 4233], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 0 | 0 | 0 | 45 | Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 43 | +-- Anodontini **
--+
+-- Unionini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 2 | 2 | 1 | 41 | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Cyr et al., 2007 [Publ 2745], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 0 | 0 | 0 | 47 | Cyr et al., 2007 [Publ 2745], fig. 2C iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 0 | 0 | 0 | 47 | Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 20 | 20 | 2 | 47 | +-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Acuticostini **
+--+
+-- Unionini **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 6 | 4 | 2 | 42 | +-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Soroka, 2005 [Publ 2726], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 1 | 1 | 1 | 44 | +-- Gonideinae
--+
+-- Unionini **
Sayenko et al., 2017 [Publ 3760], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 47 | Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 6 | 5 | 2 | 42 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 10 | 9 | 3 | 46 | +-------------- Parreysiinae
|
--+ +----------- Ambleminae
| |
+--+ +-------- Gonideinae
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Aculamprotula **
+--+
| +-- Anodontini **
+--+
+-- Unio **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 0 | 2 | 11 | 5 | 3 | 45 | +----- Pleurobemini
|
+----- Lampsilini
--+
| +-- Anodontini **
| |
| +-- Unio_ravoisieri **
+--+
+-- Unio_pictorum **
|
+-- Aculamprotula **
Bulakhova et al., 2023 [Publ 4325], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 48 | Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 9 | 0 | 2 | 11 | 5 | 3 | 50 | +-- Aculamprotula **
|
+-- Lamprotula
|
+-- Lampsilini
|
+-- Potomida
|
--+-- Unio_pictorum **
|
+-- Anodontini **
|
+-- Unio_ravoisieri **
|
+-- Parreysiinae
|
+-- Pleurobemini
Cyr et al., 2007 [Publ 2745], fig. 2D iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 48 | Soroka, 2010 [Publ 3404], fig. 9c iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 0 | 0 | 0 | 49 | Lopes-Lima et al., 2021 [Publ 4217], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 44 | Mezhzherin et al., 2014 [Publ 3705], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 44 | Klishko et al., 2014 [Publ 3666], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 45 | Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 15 | 2 | 2 | 15 | 12 | 2 | 55 | +-------- Gonideinae
|
| +-- Elliptio
| |
| +-- Fusconaia
| |
--+ +--+-- Amblemini
| | |
| | +-- Pleurobema
| | |
| | +-- Unionini **
| |
| | +-- Pyganodon_cataracta **
+--+ |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Lampsilis_teres
| |
| +-- Lampsilis_straminea
+--+
+-- Megalonaias
|
+-- Obliquaria
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 5 | 5 | 2 | 42 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Cyr et al., 2007 [Publ 2745], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 1 | 1 | 1 | 48 | +-- Ambleminae
--+
+-- Anodontini **
Klishko et al., 2016 [Publ 3733], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 45 | Burzynski & Soroka, 2018 [Publ 3862], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 42 | Froufe et al., 2017 [Publ 3802], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 42 | Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 14 | 8 | 2 | 45 | +----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 5 | 3 | 2 | 12 | 6 | 3 | 47 | +-------------------------- Lamellidens_mainwaringi
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilini
+--+
| +-------------- Pleurobemini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio_pictorum **
+--+
| +----- Anodontini **
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens_corrianus
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 32 | 20 | 3 | 45 | +----------------- Parreysiinae
|
--+ +-------------- Contradentini
| |
+--+ +----------- Pseudodontini
| |
+--+ +-- Anodontini **
| +-----+
| | +-- Unionini **
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Lopes-Lima et al., 2020 [Publ 4080], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 2 | 0 | 0 | 47 | Amaro et al., 2017 [Publ 3791], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 8 | 5 | 2 | 42 | +-------- Anodontini **
--+
| +----- Lampsilini
+--+
| +-- Quadrulini
+--+
+-- Gonideinae
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 32 | 20 | 3 | 45 | +-------------- Parreysiinae
|
| +-- Pseudodontini
--+ +--------+
| | +-- Contradentini
| |
+--+ +-- Anodontini **
| +-----+
| | +-- Unionini **
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Amaro et al., 2017 [Publ 3791], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 1 | 6 | 6 | 2 | 46 | +----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 9b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 48 | Nagel et al., 1998 [Publ 1946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 1 | 0 | 0 | 0 | 44 | Mezhzherin et al., 2014 [Publ 3705], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 42 | Amaro et al., 2017 [Publ 3791], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 8 | 5 | 2 | 42 | +----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Wu et al., 2024 [Publ 4388], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 5 | 5 | 1 | 48 | +-- Unionini **
--+
+-- Gonideinae
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 12 | 9 | 2 | 45.5 | Nagel et al., 1998 [Publ 1946], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 3 | 2 | 1 | 44.5 | +-------- Lamprotulini
--+
| +----- Gonideini
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2016 [Publ 3757], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 43 | Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 46 | +-- Anodontini **
--+
+-- Ambleminae
Soroka, 2008 [Publ 3695], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 1 | 1 | 1 | 43 | +-- Ambleminae
--+
+-- Anodontini **
Froufe et al., 2015 [Publ 3664], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 9 | 9 | 2 | 43 | +-- Gonideini
+-----+
| +-- Lamprotulini
--+
| +----- Chamberlainiini
+--+
| +-- Ambleminae
+--+
+-- Anodontini **
Kilikowska et al., 2020 [Publ 4335], fig. 4a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 45 | Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 2 | 1 | 1 | 46 | +-- Gonideinae
--+
+-- Anodontini **
Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 27 | 1 | 1 | 45 | +----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Krebs et al., 2019 [Publ 4092], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 47 | Soroka, 2008 [Publ 3695], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 1 | 1 | 1 | 43 | +-- Ambleminae
--+
+-- Anodontini **
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 12 | 9 | 2 | 45.5 | +----------- Pseudodontini
--+
| +-------- Gonidea
+--+
| +----- Inversidens
+--+
| +-- Anodontini **
+--+
+-- Ambleminae
Soroka, 2010 [Publ 3404], fig. 8b-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 0 | 0 | 0 | 43 | +-- Anodontini **
--+
+-- Unionini **
Froufe et al., 2014 [Publ 3592], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 45 | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 37 | 27 | 3 | 44 | +-------- Parreysiinae
|
--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Froufe et al., 2015 [Publ 3664], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 5 | 5 | 2 | 43 | +----- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Soroka, 2010 [Publ 3404], fig. 8a-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 0 | 0 | 0 | 43 | +-- Anodontini **
--+
+-- Unionini **
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 44 | 42 | 1 | 43 | +-- Anodontini **
--+
+-- Gonideinae
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 17 | 13 | 3 | 43 | +-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Parreysiinae
+--+
+-- Anodontini **
Hinzmann et al., 2013 [Publ 3541], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 43 | Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 27 | 1 | 1 | 45 | +----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Wen et al., 2017 [Publ 3795], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 43 | +-- Anodontini **
--+
+-- Ambleminae
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 45 | 43 | 1 | 43 | +-- Anodontini **
--+
+-- Gonideinae
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 12 | 9 | 2 | 45.5 | +----------- Pseudodontini
--+
| +-------- Gonidea
+--+
| +----- Inversidens
+--+
| +-- Anodontini **
+--+
+-- Ambleminae
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 26 | 24 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 10 | 1 | 1 | 46.5 | +-- Unionini **
|
--+-- Anodontini **
|
+-- Ambleminae
Soroka, 2010 [Publ 3404], fig. 5a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 2 | 1 | -- | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Huang et al., 2013 [Publ 3629], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 4 | 2 | -- | +----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Taylor et al., 2007 [Publ 2696], fig. 1-3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 0 | 0 | -- | +-- Anodontini **
--+
+-- Unionini **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 24 | 22 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Guerra et al., 2017 [Publ 3772], fig. 2a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 4 | 2 | -- | +-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Soroka, 2010 [Publ 3404], fig. 4-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- | Walker et al., 2006 [Publ 2666], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 12 | 8 | 2 | -- | +----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Hoeh et al., 2002 [Publ 2673], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 6 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Amaro et al., 2017 [Publ 3791], fig. 3A-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 5 | 4 | 2 | -- | +-------- Quadrulini
--+
| +----- Gonideinae
+--+
| +-- Lampsilini
+--+
+-- Anodontini **
Amaro et al., 2017 [Publ 3791], fig. 3B-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 5 | 4 | 2 | -- | +-------- Quadrulini
--+
| +----- Lampsilini
+--+
| +-- Anodontini **
+--+
+-- Gonideinae
Mock et al., 2004 [Publ 2686], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- | Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 15 | 4 | 2 | 47 | +-------- Unionini **
--+
| +----- Pleurobemini
+--+
| +-- Anodontini **
+--+
+-- Lampsilini
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 1 | 1 | -- | +----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Hoeh et al., 1996 [Publ 2694], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 2 | 1 | 1 | -- | +-- None__1
+--+
| +-- None__2
--+
| +-- Ambleminae
+--+
+-- Anodontini **
Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 1 | 1 | -- | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Hoeh et al., 2002 [Publ 2673], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 6 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Soroka, 2010 [Publ 3404], fig. 5b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- | Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 8 | 0 | 1 | 6 | 6 | 2 | 50 | +-------------- Cristariina **
--+
| +----------- Alasmidontina **
+--+
| +-------- Lampsilis
+--+
| +----- Gonideinae
+--+
| +-- Venustaconcha
+--+
+-- Quadrulini
Krebs, 2004 [Publ 2692], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 3 | 3 | 1 | -- | +-------- Quadrulini
--+
| +----- Anodontini **
+--+
| +-- Lampsilini
+--+
+-- Pleurobemini
Mock et al., 2004 [Publ 2686], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- | Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 1 | 4 | 4 | 2 | 46 | +----- Gonideinae
|
--+ +-- Cristariina **
| |
| +-- Lampsilini
+--+
+-- Alasmidontina **
|
+-- Quadrulini
Huang et al., 2018 [Publ 3828], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 27 | 4 | 3 | -- | +-------- Gonideinae
--+
| +----- Unionini **
+--+
| +-- Ambleminae
+--+
+-- Parreysiinae
Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 5 | 5 | 2 | -- | +----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Gonideinae
Walker et al., 2006 [Publ 2666], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 8 | 2 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 1 | 1 | -- | +----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Zhang et al., 2013 [Publ 4214], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 1 | 2 | 1 | 1 | 47 | +-------- Cristaria **
--+
| +----- Sinanodonta **
+--+
| +-- Margaritiferinae
+--+
+-- Gonideinae
Krebs, 2004 [Publ 2692], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 2 | 2 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Curole & Kocher, 2005 [Publ 2682], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 10 | 9 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Xia et al., 2016 [Publ 3712], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 1 | 1 | 1 | 1 | 45 | +----- Sinanodonta **
--+
| +-- Cristaria **
+--+
+-- Gonideinae
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 42 | 3 | -- | +----------------- Ambleminae
|
| +-------- Parreysiinae
| |
| +-----+ +-- Anodontini **
--+ | | +--+
| | | | +-- Unionini **
| | +--+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Discomya
|
| +----- Ctenodesmini
| +-----+
| | | +-- Rectidentini
| | +--+
| | +-- Contradentini
+--+
| +-- Schepmaniini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Caudiculatus
+--+
| +-- Potomida
+--+
+-- Gonideini
Guerra et al., 2017 [Publ 3772], fig. 2b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 4 | 2 | -- | +-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 1 | 1 | -- | +----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 27 | 25 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 12 | 1 | 1 | -- | +-- Anodontini **
--+
+-- Unionini **
Bolotov et al., 2015 [Publ 3646], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Combosch et al., 2017 [Publ 3806], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 2 | 2 | -- | +----- Unionini **
--+
| +-- Parreysiinae
+--+
+-- Ambleminae
Combosch et al., 2017 [Publ 3806], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 2 | 2 | -- | +----- Unionini **
--+
| +-- Ambleminae
+--+
+-- Parreysiinae
González & Giribet, 2015 [Publ 3632], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Hewitt et al., 2019 [Publ 3962], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 8 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Inoue et al., 2013 [Publ 3528], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 16 | 2 | -- | +-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Bieler et al., 2014 [Publ 3622], fig. 28 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Lydeard et al., 2000 [Publ 2654], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 1 | -- | Wang et al., 2016 [Publ 3714], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 2 | 1 | -- | +-- Unionini **
--+
+-- Gonideinae
Vikhrev et al., 2018 [Publ 3812], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- | +-- Unionini **
--+
+-- Ambleminae
Graf et al., 2014 [Publ 3591], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 9 | 1 | -- | Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 55 | 1 | -- | Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 54 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
González & Giribet, 2015 [Publ 3632], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Peng et al., 2012 [Publ 3689], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 2 | 1 | -- | +-- Anodontini **
--+
+-- Gonideinae
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 37 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Lydeard et al., 2000 [Publ 2654], fig. 2c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 1 | -- | Huff et al., 2004 [Publ 2697], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Pieri et al., 2018 [Publ 3867], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 19 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Lopes-Lima et al., 2018 [Publ 3829], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 2 | 2 | -- | +----- Gonideinae
--+
| +-- Ambleminae
+--+
+-- Unionini **
Bolotov et al., 2015 [Publ 3646], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 2 | 1 | -- | +-- Pleurobemini
|
--+-- Lampsilini
|
+-- Unionini **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 2 | 2 | -- | +----- Gonideinae
--+
| +-- Ambleminae
+--+
+-- Unionini **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 3 | 2 | -- | +-- Anodontini **
|
--+-- Ambleminae
|
+-- Gonideinae
Graf et al., 2015 [Publ 3619], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Campbell, 2000 [Publ 2724], fig. 1-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 41 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
González & Giribet, 2015 [Publ 3632], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Hoeh et al., 1998 [Publ 2695], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 1 | 1 | -- | Araujo et al., 2009 [Publ 2395], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 16 | 2 | -- | +----- Ambleminae
--+
| +-- Lanceolariini **
+--+
+-- Gonideinae
Mulvey et al., 1997 [Publ 3390], fig. 1_3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 4 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Turner et al., 2000 [Publ 2722], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 1 | -- | +-- Quadrulini
+-----+
| +-- Anodontini **
--+
| +----- Lampsilini
+--+
| +-- Amblemini
+--+
+-- Pleurobemini
Lydeard et al., 2000 [Publ 2654], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 1 | -- | Klunzinger et al., 2021 [Publ 4086], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Curole & Kocher, 2002 [Publ 2712], fig. 1a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- | +----- Quadrulini
--+
| +-- Anodontini **
+--+
+-- Lampsilini
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- | +----- Lampsilini
--+
| +-- Anodontini **
+--+
+-- Quadrulini
Inoue et al., 2013 [Publ 3579], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Zanatta & Harris, 2013 [Publ 3530], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 4 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Lydeard et al., 2000 [Publ 2654], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 12 | 1 | -- | Metzger et al., 2018 [Publ 3952], fig. 1C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 2 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 40 | 1 | -- | +-- Anodontini **
--+
+-- Ambleminae
Hoeh et al., 1998 [Publ 2695], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 1 | 1 | -- | Curole & Kocher, 2005 [Publ 2682], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 9 | 1 | -- | Jiang et al., 2010 [Publ 3438], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 3 | 2 | -- | +-- Ambleminae
|
--+-- Anodontini **
|
+-- Gonideinae
|
