The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe UNIONINI Rafinesque, 1820: cladomics
14 genera Cladomics Summarynumber of species = 84 (106 total)
number of subtaxa (rank: genus) = 14
analyzed in 252 cladograms in 142 publications
maximum number of terminals in an analysis =
49 (mean = 7.14, median = 5)
maximum number of subtaxa (genus) =
11 (mean = 2.98, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 213
maximum number of included terminals = 49, (mean =
8.26 median = 6)
maximum number of subtaxa (genus) =
11 (mean = 3.34, median = 2)
proportion of cladograms that support basic monophyly
= 0.85 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 120
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 49 (mean = 10.67, median = 10)
maximum number of subtaxa (genus) =
11 (mean = 4.88, median = 5)
proportion of cladograms that support rigorous monophyly = 0.77
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Dai et al., 2025 [Publ 4440], fig. 2 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 19 | 11 | 23 | 14 | 4 | 0 | +-- Anemina
+--------------------------------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +--------------------------+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
+--+
| +-- Alasmidontina
| +--------------------------+
| | +-- Anodontina
| |
+--+ +-------------------------- Lanceolaria_gladiola
| |
| +-------------------------- Lanceolaria_oxyrhyncha
| |
+--+-------------------------- Cristaria
|
| +----------------------- Acuticostini
| |
| +----------------------- Lepidodesmini
+--+
| +-------------------- Globunio **
| |
+--+ +----------------- Aculamprotula **
| |
| | +-- Middendorffinaia **
+--+ +-----------+
| | +-- Nodularia **
| |
+--+ +----------- Cuneopsis **
| |
| | +-- Diaurora **
+--+--------+
| +-- Schistodesmus **
|
| +-------- Pseudocuneopsis **
+--+
| +----- Inversiunio **
+--+
| +-- Tchangsinaia **
+--+
+-- Unio **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 23 | 8 | 94 | 43 | 4 | 25 | +-------------------- Acuticostini
|
+-------------------- Lepidodesmini
|
| +-- Lanceolariini
--+-----------------+
| +-- Anodontini
|
| +----------------- Aculamprotula **
| |
+--+ +-------------- Middendorffinaia **
| |
+--+ +----------- Unio **
| |
+--+ +-------- Nodularia **
| |
+--+ +----- Schistodesmus **
| |
+--+----- Cuneopsis **
|
| +-- Tchangsinaia **
+--+
+-- Inversiunio **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 49 | 49 | 47 | 8 | 27 | 23 | 3 | 48 | +-------------------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +--------------------+
| | +-- Anodontini
+--+
| +-------------------- Aculamprotula **
+--+
| +----------------- Unio **
+--+
| +-------------- Middendorffinaia **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 29 | 29 | 28 | 7 | 24 | 22 | 3 | 31 | +----------------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------------+
| | +-- Anodontini
+--+
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 29 | 63 | 61 | 9 | 59 | 15 | 4 | 63 | +-------------- Chamberlainiini
|
+-----------+ +-- Contradentini
| | +--------+
| | | +-- Rectidentini
| +--+
| | +-------- Lamprotulini
| +--+
| | +----- Pseudodontini
--+ +--+
| | +-- Inversiunio **
| +--+
| +-- Gonideini
|
| +----------------------- Lepidodesmini
| |
| | +----- Anodontini
+--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Globunio **
| |
+--+ +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Pseudocuneopsis **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 6 | 2 | 2 | 1 | 35 | +----------------- Lepidodesmini
--+
| +-------------- Middendorffinaia **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 6 | 2 | 2 | 1 | 35 | +----------------- Lepidodesmini
--+
| +-------------- Middendorffinaia **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Lyubas et al., 2022 [Publ 4255], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 14 | 1 | 8 | 5 | 1 | 41 | +-- Anodontini
--+
+-- Unio **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 21 | 8 | 15 | 15 | 4 | 22 | +-------------------------- Lepidodesmini
|
| +----- Lanceolariini
--+ +-----------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Anodontini
|
| +-------------------- Aculamprotula **
+--+
| +----------------- Unio **
+--+
| +-------------- Pseudocuneopsis **
+--+
| +----------- Nodularia **
+--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Inversiunio **
Lyubas et al., 2022 [Publ 4255], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 20 | 1 | 8 | 5 | 1 | 41 | +-- Anodontini
--+
+-- Unio **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 17 | 11 | 23 | 14 | 4 | 6 | +----- Pletholophus
+--------------------------------+
| | +-- Sinanodonta
| +--+
| +-- Beringiana
--+
| +----------------------------------- Cristaria
| |
| | +-- Alasmidontina
| +--------------------------------+
+--+ +-- Anodontina
|
| +-------------------------------- Lanceolariini
| |
+--+ +----------------------------- Lepidodesmini
| |
+--+ +-------------------------- Acuticostini
| |
+--+ +----------------------- Globunio **
| |
+--+ +-------------------- Aculamprotula **
| |
+--+ +----------------- Middendorffinaia **
| |
+--+ +-------------- Nodularia **
| |
+--+ +----------- Cuneopsis **
| |
| | +-- Diaurora **
+--+--------+
| +-- Schistodesmus **
|
| +-------- Pseudocuneopsis **
+--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Inversiunio **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 7 | 73 | 24 | 4 | 25 | +-------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Pseudocuneopsis **
+--+
| +----- Tchangsinaia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 19 | 7 | 69 | 24 | 4 | 22 | +-------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-------- Nodularia **
| |
+--+-------- Pseudocuneopsis **
|
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 19 | 7 | 69 | 24 | 4 | 22 | +-------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-------- Nodularia **
| |
+--+-------- Pseudocuneopsis **
|
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 22 | 22 | 20 | 7 | 73 | 24 | 4 | 26 | +-------------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------------- Aculamprotula **
| |
+--+ +----------------- Unio **
| |
+--+ +-------------- Pseudocuneopsis **
| |
+--+ +----------- Nodularia **
| |
+--+ +-- Schistodesmus_lampreyanus__1 **
| +-----+
| | +-- Cuneopsis **
+--+
| +----- Tchangsinaia **
+--+
| +-- Schistodesmus_lampreyanus__2 **
+--+
+-- Schistodesmus_spinosus **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 18 | 7 | 53 | 15 | 4 | 21 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 19 | 7 | 20 | 6 | 4 | 26 | +----------------------- Lepidodesmini
|
| +----- Lanceolariini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Anodontini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 45 | 22 | 6 | 180 | 26 | 4 | 50 | +-------- Acuticostini
|
+-------- Alasmidonta
|
+-------- Anodonta_anatina
|
+-------- Anemina
|
+-------- Anodonta_cygnea
|
+-------- Cristaria
|
+-------- Anodonta_nuttalliana
|
+-------- Lasmigona
|
+-------- Lepidodesmini
|
+-------- Pseudanodonta
|
+-------- Pyganodon
|
--+-------- Simpsonaias
|
+-------- Strophitus
|
+-------- Schistodesmus **
|
+-------- Utterbackia
|
+-------- Lanceolariini
|
+-------- Unio **
|
+-------- Aculamprotula **
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis **
| |
| +----- Tchangsinaia **
| |
+--+----- Cuneopsis_rufescens **
|
+----- Nodularia **
|
| +-- Cuneopsis_szechenyii **
+--+
+-- Cuneopsis_heudei **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 16 | 6 | 180 | 26 | 4 | 23 | +-------------------- Acuticostini
|
--+ +----------------- Lepidodesmini
| |
| | +-- Lanceolariini
+--+--------------+
| +-- Anodontini
|
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Schistodesmus **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 3 | 3 | 3 | 2 | 36 | +-- Lanceolariini
+-----+
| +-- Anodontini
--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 3 | 3 | 3 | 2 | 35 | +-- Lanceolariini
+-----+
| +-- Anodontini
--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 6 | 179 | 26 | 4 | 25 | +-------------------- Acuticostini
|
| +----- Lepidodesmini
--+ +-----------+
| | | +-- Lanceolariini
| | +--+
+--+ +-- Anodontini
|
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 14 | 10 | 23 | 14 | 4 | 11 | +----------------------- Acuticostini
|
| +-------------------- Lepidodesmini
--+ |
| | +-- Lanceolariini
| +-----------------+
+--+ +-- Anodontini
|
| +----------------- Aculamprotula **
| |
| | +-- Middendorffinaia **
+--+ +-----------+
| | +-- Nodularia **
| |
+--+ +----------- Cuneopsis **
| |
| | +-- Diaurora **
+--+--------+
| +-- Schistodesmus **
|
| +-------- Pseudocuneopsis **
+--+
| +----- Inversiunio **
+--+
| +-- Tchangsinaia **
+--+
+-- Unio **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 14 | 10 | 23 | 14 | 4 | 11 | +-- Anemina
+-----------------------+
| +-- Amuranodonta
|
| +----- Pletholophus
| +-----------------+
--+ | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
| |
| | +-------------------- Lanceolaria_gladiola
+--+ |
| +-------------------- Acuticostini
| |
| +-------------------- Lepidodesmini
| |
| +-------------------- Lanceolaria_oxyrhyncha
| |
+--+-------------------- Cristaria
|
| +-- Alasmidontina
+-----------------+
| +-- Anodontina
|
| +----------------- Aculamprotula **
| |
| | +-- Middendorffinaia **
+--+ +-----------+
| | +-- Nodularia **
| |
+--+ +----------- Cuneopsis **
| |
| | +-- Diaurora **
+--+--------+
| +-- Schistodesmus **
|
| +-------- Pseudocuneopsis **
+--+
| +----- Inversiunio **
+--+
| +-- Tchangsinaia **
+--+
+-- Unio **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 7 | 35 | 10 | 4 | 25 | +----------------------- Lepidodesmini
|
| +----- Lanceolariini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Anodontini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 1 | 9 | 3 | 1 | 36 | +-- Anodontini
--+
+-- Unio **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 7 | 53 | 17 | 4 | 22 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 7 | 58 | 18 | 4 | 22 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 7 | 58 | 18 | 4 | 22 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Cuneopsis **
+--+
+-- Tchangsinaia **
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 21 | 19 | 5 | 28 | 14 | 4 | 26 | +----------------------------------- Anemina
|
--+ +-------------------------------- Acuticostini
| |
+--+ +----------------------------- Sinanodonta
| |
+--+ +-------------------------- Aculamprotula **
| |
+--+ +-- Cristaria
| +--------------------+
| | +-- Lanceolariini
+--+
| +-------------------- Lepidodesmini
+--+
| +----------------- Schistodesmus **
+--+
| +-------------- Nodularia **
+--+
| +----------- Cuneopsis_rufescens **
+--+
| +-------- Tchangsinaia **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_szechenyii **
+--+
+-- Cuneopsis_heudei **
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 5 | 30 | 16 | 4 | 20 | +-------------------------- Anemina
|
--+ +----------------------- Sinanodonta
| |
+--+ +-------------------- Acuticostini
| |
+--+ +-- Lepidodesmini
| +--------------+
| | +-- Cristaria
+--+
| +-------------- Lanceolariini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 7 | 35 | 13 | 4 | 23 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis **
+--+
+-- Schistodesmus **
Froufe et al., 2016 [Publ 3738], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 2 | 0 | 0 | 0 | 30 | +-- Nodularia **
--+
+-- Unio **
Lopes-Lima et al., 2024 [Publ 4376], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 1 | 0 | 0 | 0 | 33 | Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 6 | 39 | 19 | 4 | 19 | +-------------- Lepidodesmini
|
| +----- Anodontini
+--------+
| | +-- Acuticostini
--+ +--+
| +-- Lanceolariini
|
| +----------- Aculamprotula **
| |
+--+ +-------- Unio **
| |
+--+ +-- Nodularia **
| +--+
| | +-- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 8 | 10 | 8 | 3 | 29 | +----------------------------- Acuticostini
|
--+ +-- Lanceolaria_lanceolata
| +-----------------------+
| | +-- Lanceolaria_grayii
+--+
| +----------------------- Lanceolaria_gladiola
| |
+--+ +-------------------- Anodontini
| |
+--+ +----------------- Unio **
| |
+--+ +-------------- Aculamprotula **
| |
+--+ +----------- Cuneopsis **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Schistodesmus **
+--+
| +----- Tchangsinaia **
+--+
| +-- Inversiunio **
+--+
+-- Pseudocuneopsis **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 10 | 2 | 4 | 3 | 1 | 32 | +----- Anodontini
--+
| +-- Nodularia **
+--+
+-- Unio **
Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 17 | 16 | 3 | 8 | 3 | 1 | 28 | +-------- Aculamprotula **
--+
| +----- Anodontini
+--+
| +-- Schistodesmus **
+--+
+-- Nodularia **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 9 | 8 | 10 | 8 | 3 | 26 | +----------------- Acuticostini
|
| +-------------- Lanceolaria_grayii
--+ |
| +-------------- Lanceolaria_gladiola
| |
+--+-------------- Lanceolaria_lanceolata
|
| +----------- Anodontini
| |
| | +-------- Unio **
+--+ |
| +-------- Aculamprotula **
| |
+--+ +----- Cuneopsis **
| |
| | +-- Nodularia **
| +--+
+--+ +-- Schistodesmus **
|
| +-- Tchangsinaia **
| |
+--+-- Inversiunio **
|
+-- Pseudocuneopsis **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 14 | 6 | 55 | 22 | 3 | 26 | +-- Anemina
+-----------------------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +-----------------------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +-------------------------- Cristaria
| |
+--+ +-- Anodontina
| +--------------------+
| | +-- Alasmidontina
+--+
| +-------------------- Lanceolariini
+--+
| +----------------- Lepidodesmini
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Inversiunio **
+--+
| +----- Nodularia **
+--+
| +-- Schistodesmus **
+--+
+-- Tchangsinaia **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 7 | 33 | 13 | 4 | 24 | +----------------------- Lepidodesmini
|
| +----- Anodontini
--+ +--------------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis **
+--+
+-- Schistodesmus **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 14 | 6 | 55 | 22 | 3 | 26 | +-- Anemina
+-----------------------------+
| +-- Buldowskia
|
--+ +----- Sinanodonta
| +-----------------------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +-------------------------- Cristaria
| |
+--+ +-- Anodontina
| +--------------------+
| | +-- Alasmidontina
+--+
| +-------------------- Lepidodesmini
+--+
| +----------------- Lanceolariini
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Nodularia **
+--+
| +----- Inversiunio **
+--+
| +-- Schistodesmus **
+--+
+-- Tchangsinaia **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 13 | 6 | 55 | 22 | 3 | 26 | +-- Anemina
+-----------------------------+
| +-- Buldowskia
|
--+ +----- Pletholophus
| +-----------------------+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +-------------------------- Cristaria
| |
+--+ +-- Anodontina
| +--------------------+
| | +-- Alasmidontina
+--+
| +-------------------- Lanceolariini
+--+
| +----------------- Lepidodesmini
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Inversiunio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Hou et al., 2024 [Publ 4400], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 4 | 2 | 0 | 0 | 27 | +----- Cuneopsis_rufescens **
+--------+
| | +-- Middendorffinaia **
| +--+
--+ +-- Pseudocuneopsis **
|
| +----------- Tchangsinaia **
| |
+--+ +-- Cuneopsis_kiangsiensis **
| +-----+
| | +-- Cuneopsis_celtiformis **
+--+
| +----- Cuneopsis_szechenyii **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 8 | 10 | 8 | 3 | 28 | +-------------------------- Acuticostini
|
--+ +----------------------- Lanceolariini
| |
+--+ +-------------------- Anodontini
| |
+--+ +----------------- Unio **
| |
+--+ +-------------- Aculamprotula **
| |
+--+ +----------- Cuneopsis **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Schistodesmus **
+--+
| +----- Tchangsinaia **
+--+
| +-- Inversiunio **
+--+
+-- Pseudocuneopsis **
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 4 | 0 | 0 | 0 | 26 | +-------- Unio **
--+
| +----- Inversiunio **
+--+
| +-- Middendorffinaia **
+--+
+-- Nodularia **
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 7 | 0 | 0 | 0 | 19.5 | +----------------- Aculamprotula **
--+
| +-------------- Nodularia **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Inversiunio **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 16 | 14 | 7 | 4 | 4 | 2 | 32 | +----- Aculamprotula **
+--------+
| | +-- Acuticostini
| +--+
--+ +-- Lanceolariini
|
| +----------- Unio **
| |
+--+ +-------- Pseudocuneopsis **
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Wu et al., 2024 [Publ 4379], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 4 | 0 | 0 | 0 | 26 | +----- Nodularia **
|
--+ +-- Cuneopsis **
| |
+--+-- Tchangsinaia **
|
+-- Pseudocuneopsis **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 7 | 5 | 5 | 2 | 22 | +----------------------- Anodontini
--+
| +-------------------- Lanceolariini
+--+
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis **
+--+
+-- Schistodesmus **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 6 | 44 | 21 | 4 | 25 | +-- Lepidodesmini
+--+
| +-- Lanceolariini
+-----------+
| | +-- Acuticostini
| +--+
--+ +-- Anodontini
|
| +-------------- Middendorffinaia **
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 11 | 5 | 0 | 0 | 0 | 23 | +----------- Nodularia **
--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis **
+--+
| +-- Unio **
+--+
+-- Tchangsinaia **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 7 | 41 | 15 | 4 | 25 | +----------------------- Lepidodesmini
|
--+ +-------------------- Lanceolariini
| |
+--+ +-- Acuticostini
| +--------------+
| | +-- Anodontini
+--+
| +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-- Schistodesmus **
| +-----+
| | +-- Cuneopsis **
+--+
| +----- Pseudocuneopsis **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Prié & Puillandre, 2014 [Publ 3529], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 1 | 6 | 3 | 1 | 29 | +-- Anodontini
--+
+-- Unio **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 6 | 80 | 15 | 4 | 23 | +-------------------- Acuticostini
|
| +----- Lanceolariini
--+ +-----------+
| | | +-- Lepidodesmini
| | +--+
+--+ +-- Anodontini
|
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Nodularia **
+--+
| +----- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 6 | 29 | 15 | 4 | 24 | +-------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis **
+--+
+-- Schistodesmus **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 15 | 13 | 5 | 6 | 5 | 3 | 29 | +-------------- Aculamprotula **
|
--+ +-- Acuticostini
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 5 | 45 | 15 | 4 | 23 | +----------------- Lepidodesmini
|
| +----- Lanceolariini
--+ +--------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Anodontini
|
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 8 | 4 | 4 | 3 | 3 | 25 | +-------- Lanceolariini
|
--+ +----- Unio **
| |
+--+ +-- Schistodesmus **
| |
+--+-- Cuneopsis **
|
+-- Nodularia **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 8 | 6 | 69 | 17 | 2 | 27 | +-- Lanceolariini
+--------------+
| +-- Anodontini
--+
| +-------------- Unio **
+--+
| +----------- Aculamprotula **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 7 | 10 | 5 | 4 | 26 | +----------------------------- Anodontina
|
--+ +-- Acuticostini
| +-----------------------+
| | +-- Cristariina
+--+
| +----------------------- Lepidodesmini
+--+
| +-------------------- Lanceolariini
+--+
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Middendorffinaia **
+--+
| +-------- Nodularia **
+--+
| +----- Cuneopsis **
+--+
| +-- Inversiunio **
+--+
+-- Schistodesmus **
Hou et al., 2025 [Publ 4401], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 1 | 0 | 0 | 0 | 29 | Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 6 | 80 | 15 | 4 | 23 | +----------------------- Acuticostini
|
--+ +-- Lanceolariini
| +-----------------+
| | +-- Anodontini
+--+
| +----------------- Lepidodesmini
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Unio **
+--+
| +-------- Nodularia **
+--+
| +----- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 13 | 10 | 7 | 10 | 5 | 4 | 27 | +----------------- Acuticostini
|
+----------------- Cristariina
|
+----------------- Anodontina
--+
| +-------------- Lanceolariini
| |
| | +-- Aculamprotula **
| +-----------+
+--+ +-- Lepidodesmini
|
| +----------- Unio **
| |
+--+ +-------- Middendorffinaia **
| |
+--+ +----- Cuneopsis **
| |
+--+----- Nodularia **
|
| +-- Inversiunio **
+--+
+-- Schistodesmus **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 6 | 129 | 17 | 2 | 26 | +-- Lanceolariini
+--------------+
| +-- Anodontini
--+
| +-------------- Unio **
+--+
| +----------- Aculamprotula **
+--+
| +-------- Nodularia **
+--+
| +----- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 34 | 12 | 3 | 24 | +----------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------+
| | +-- Anodontini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 28 | 11 | 3 | 24.5 | +----------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------+
| | +-- Anodontini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Hou et al., 2025 [Publ 4432], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 4 | 1 | 1 | 1 | 26 | +----------- Lepidodesmini
--+
| +-------- Tchangsinaia **
+--+
| +----- Pseudocuneopsis **
+--+
| +-- Middendorffinaia **
+--+
+-- Cuneopsis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 6 | 6 | 129 | 17 | 2 | 24 | +----------- Unio **
|
+--+ +-------- Aculamprotula **
| | |
| | +-------- Schistodesmus **
| +--+
| +-------- Nodularia **
| |
| | +----- Cuneopsis_szechenyii **
--+ +--+
| | +-- Tchangsinaia **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-------- Lanceolariini
| |
| +-------- Cristariina
+-----+
| +----- Anodonta_anatina
| |
| +----- Pseudanodonta
+--+
+----- Alasmidontina
|
| +-- Anodonta_cygnea
+--+
+-- Anodonta_nuttalliana
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 6 | 66 | 16 | 2 | 26 | +-- Lanceolariini
+-----------------+
| +-- Anodontini
--+
| +----------------- Unio **
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Nodularia **
+--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis_szechenyii **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis_rufescens **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 7 | 19 | 7 | 1 | 21 | +-------------------- Anodontini
--+
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Cuneopsis **
+--+
+-- Tchangsinaia **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 1 | 0 | 0 | 0 | 30 | Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 12 | 10 | 6 | 3 | 2 | 2 | 29 | +-------------- Unio **
|
| +----- Aculamprotula **
--+ +-----+
| | | +-- Anodontini
| | +--+
+--+ +-- Lepidodesmini
|
| +-------- Middendorffinaia **
+--+
| +----- Inversiunio **
+--+
| +-- Cuneopsis **
+--+
+-- Nodularia **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 0 | 6 | 115 | 17 | 2 | 23 | +-- Unio_crassus **
|
+-- Unio_pictorum **
|
+-- Unio_tumidus **
|
+-- Aculamprotula **
|
+-- Cuneopsis_szechenyii **
+--+
| +-- Tchangsinaia **
| |
| +-- Cuneopsis_rufescens **
| |
| +-- Nodularia_douglasiae **
| |
| +-- Nodularia_nuxpersicae **
| |
| +-- Schistodesmus **
|
| +-- Alasmidonta
| |
| +-- Anodonta_anatina
--+ |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
| |
| +-- Lasmigona
| |
| +-- Pseudanodonta
| |
| +-- Pyganodon
| |
| +-- Simpsonaias
| |
| +-- Strophitus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii__1
|
+-- Lanceolaria_grayii__2
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 6 | 66 | 16 | 2 | 26 | +-- Lanceolariini
+-----------------+
| +-- Anodontini
--+
| +----------------- Unio **
+--+
| +-------------- Aculamprotula **
+--+
| +----------- Nodularia **
+--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis_szechenyii **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis_rufescens **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 13 | 13 | 3 | 24 | +-------------------- Anodontini
--+
| +----------------- Lanceolariini
+--+
| +-------------- Lepidodesmini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 38 | 12 | 3 | 24 | +----------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------+
| | +-- Anodontini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Schistodesmus **
+--+
+-- Tchangsinaia **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 5 | 34 | 12 | 3 | 24 | +----------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------+
| | +-- Anodontini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Schistodesmus **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 4 | 33 | 9 | 1 | 21 | +----------- Anodontini
--+
| +-------- Aculamprotula **
+--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 7 | 45 | 9 | 3 | 23 | +----------------------- Lepidodesmini
|
--+ +-- Lanceolariini
| +-----------------+
| | +-- Anodontini
+--+
| +----------------- Aculamprotula **
+--+
| +-------------- Unio **
+--+
| +----------- Pseudocuneopsis **
+--+
| +-------- Nodularia **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis **
+--+
+-- Schistodesmus **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 5 | 1 | 8 | 3 | 1 | 29 | +-- Anodontini
--+
+-- Unio **
Khalloufi et al., 2011 [Publ 2748], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 1 | 2 | 0 | 0 | 27 | Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 5 | 23 | 9 | 2 | 25 | +----------------- Anodontini
--+
| +-------------- Lanceolariini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Tchangsinaia **
+--+
+-- Cuneopsis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 2 | 2 | 0 | 0 | 24 | +-- Middendorffinaia **
--+
+-- Nodularia **
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 3 | 3 | 2 | 1 | 22 | +-------- Anodontini
--+
| +----- Unio **
+--+
| +-- Middendorffinaia **
+--+
+-- Nodularia **
Marrone et al., 2019 [Publ 3907], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 1 | 0 | 0 | 0 | 28 | Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 12 | 13 | 4 | 25 | 13 | 4 | 27 | +-------------- Anodontini
|
| +-- Tchangsinaia **
--+ +--------+
| | +-- Nodularia **
| |
+--+ +-- Schistodesmus **
| +-----+
| | +-- Acuticostini
+--+
| +----- Inversiunio **
+--+
| +-- Lepidodesmini
+--+
+-- Lanceolariini
Klishko et al., 2018 [Publ 3945], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 2 | 0 | 0 | 0 | 24 | +-- Nodularia **
--+
+-- Unio **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 46 | 10 | 4 | 24 | +-------------------- Lepidodesmini
|
| +----- Anodontini
--+ +-----------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Lanceolariini
|
| +-------------- Aculamprotula **
| |
+--+ +----------- Unio **
| |
+--+ +-- Pseudocuneopsis **
| +-----+
| | +-- Nodularia **
+--+
| +----- Schistodesmus **
+--+
| +-- Cuneopsis **
+--+
+-- Tchangsinaia **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 0 | 0 | 0 | 25 | Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 22 | 5 | 5 | 27 | 15 | 4 | 43 | +----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1
|
+----- Cristaria_plicata__2
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula
|
+----- Lepidodesmini
|
| +-- Aculamprotula **
+--+
| +-- Acuticostini
|
| +-- Cuneopsis **
+--+
| +-- Nodularia **
|
| +-- Schistodesmus **
+--+
+-- Inversiunio **
Araujo et al., 2009 [Publ 2589], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 1 | 4 | 1 | 1 | 26 | +-- Anodontini
--+
+-- Unio **
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 11 | 9 | 4 | 14 | 6 | 3 | 25 | +----- Aculamprotula **
+--+
| | +-- Tchangsinaia **
| +--+
--+ +-- Nodularia **
|
| +----- Schistodesmus **
+--+
| +-- Lanceolariini
+--+
+-- Acuticostini
Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 8 | 4 | 2 | 4 | 1 | 1 | 25 | +----------- Nodularia **
|
| +-- Unio_gibbus **
--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodontini
+--+
| +----- Unio_pictorum__1 **
| |
| +----- Unio_pictorum__2 **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Choi et al., 2020 [Publ 4169], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 3 | 0 | 0 | 25 | Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 25 | 11 | 2 | 21 | +----------------- Anodontini
--+
| +-------------- Lanceolariini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 1 | 8 | 3 | 1 | 27 | +-- Anodontini
--+
+-- Unio **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 2 | 0 | 0 | 19 | +-- Nodularia **
--+
+-- Aculamprotula **
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 7 | 5 | 3 | 22 | +-------------------- Anodontini
|
--+ +-- Acuticostini
| +--------------+
| | +-- Lanceolariini
+--+
| +-------------- Tchangsinaia **
+--+
| +----------- Cuneopsis_heudei **
+--+
| +-------- Nodularia **
+--+
| +----- Pseudocuneopsis **
+--+
| +-- Cuneopsis_rufescens **
+--+
+-- Schistodesmus **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 4 | 24 | 10 | 3 | 24 | +----- Anodontini
+-----+
| | +-- Acuticostini
| +--+
--+ +-- Lanceolariini
|
| +-------- Aculamprotula **
+--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 24 | 11 | 2 | 21 | +----------------- Anodontini
--+
| +-------------- Lanceolariini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Lopes-Lima et al., 2020 [Publ 4080], fig. 12 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 2 | 0 | 0 | 20 | +-- Nodularia **
--+
+-- Inversiunio **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 30 | 9 | 4 | 23 | +----------------- Lepidodesmini
|
| +----- Lanceolariini
--+ +--------+
| | | +-- Acuticostini
| | +--+
+--+ +-- Anodontini
|
| +----------- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Choi et al., 2020 [Publ 4169], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 3 | 0 | 0 | 0 | 21 | +----- Nodularia **
--+
| +-- Cuneopsis **
+--+
+-- Unio **
Dai et al., 2024 [Publ 4358], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 3 | 0 | 0 | 0 | 22 | +----- Pseudocuneopsis **
--+
| +-- Schistodesmus **
+--+
+-- Nodularia **
Klishko et al., 2017 [Publ 3946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20 | +-- Nodularia **
--+
+-- Unio **
Araujo et al., 2009 [Publ 2589], fig. 7C iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 4 | 1 | 1 | 25 | +-- Anodontini
--+
+-- Unio **
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 3 | 16 | 7 | 1 | 18 | +-------- Anodontini
--+
| +----- Aculamprotula **
+--+
| +-- Unio **
+--+
+-- Nodularia **
Reis et al., 2013 [Publ 3580], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 8 | 3 | 1 | 24 | +-- Anodontini
--+
+-- Unio **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 5 | 5 | 32 | 10 | 4 | 24 | +----------------- Anodontini
|
| +-- Acuticostini
--+ +-----------+
| | +-- Lanceolariini
| |
+--+ +-- Lepidodesmini
| +--------+
| | +-- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 18 | 5 | 2 | 24 | +----------- Anodontini
--+
| +-------- Lanceolariini
+--+
| +----- Schistodesmus **
+--+
| +-- Aculamprotula **
+--+
+-- Unio **
Mabuchi et al., 2021 [Publ 4233], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 23 | +-- Tchangsinaia **
--+
+-- Nodularia **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 15 | 5 | 2 | 23 | +-------- Anodontini
--+
| +----- Lanceolariini
+--+
| +-- Aculamprotula **
+--+
+-- Unio **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 17 | 5 | 2 | 22 | +----------- Anodontini
--+
| +-------- Lanceolariini
+--+
| +----- Aculamprotula **
+--+
| +-- Schistodesmus **
+--+
+-- Unio **
Wu et al., 2020 [Publ 4121], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 29 | 0 | 0 | 19 | +-- Unio **
--+
+-- Aculamprotula **
Prie et al., 2012 [Publ 3487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 1 | 0 | 0 | 23 | Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 7 | 5 | 1 | 25 | +-- Unio **
--+
+-- Anodontini
Wu et al., 2020 [Publ 4121], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 29 | 0 | 0 | 19 | +-- Unio **
--+
+-- Aculamprotula **
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 6 | 4 | 3 | 13 | 3 | 2 | 20 | +-------- Aculamprotula **
--+
| +----- Acuticostini
+--+
| +-- Nodularia **
+--+
+-- Cuneopsis **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 13 | 11 | 3 | 36 | 8 | 2 | 30 | +-------- Aculamprotula **
|
--+ +-- Schistodesmus **
| +--+
| | +-- Unio **
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 36 | 8 | 2 | 22 | +----------- Anodontini
--+
| +-------- Lanceolariini
+--+
| +----- Aculamprotula **
+--+
| +-- Schistodesmus **
+--+
+-- Unio **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 2 | 78 | 12 | 2 | 22 | +-- Unio **
+--+
| +-- Nodularia **
--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2020 [Publ 4147], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 9 | 0 | 0 | 19 | +-- Unio **
--+
+-- Aculamprotula **
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 16 | 7 | 1 | 19 | +-------- Anodontini
--+
| +----- Aculamprotula **
+--+
| +-- Unio **
+--+
+-- Nodularia **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 26 | 6 | 2 | 19 | +----------- Anodontini
--+
| +-------- Acuticostini
+--+
| +----- Aculamprotula **
+--+
| +-- Unio **
+--+
+-- Nodularia **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 5 | 5 | 2 | 23 | +-------- Lanceolariini
--+
| +----- Anodontini
+--+
| +-- Aculamprotula **
+--+
+-- Unio **
Bashê, 2022 [Publ 4265], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 1 | 1 | 0 | 0 | 26 | Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 17 | 5 | 2 | 24 | +----------- Anodontini
--+
| +-------- Lanceolariini
+--+
| +----- Schistodesmus **
+--+
| +-- Aculamprotula **
+--+
+-- Unio **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 31 | 14 | 2 | 21 | +-- Lanceolariini
+-----+
| +-- Anodontini
--+
| +----- Middendorffinaia **
+--+
| +-- Nodularia **
+--+
+-- Unio **
Nagel et al., 1998 [Publ 1946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 1 | 0 | 0 | 0 | 22 | Soroka, 2005 [Publ 2726], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 1 | 0 | 0 | 24 | Soroka, 2010 [Publ 3404], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 5 | 5 | 1 | 24 | +----- Anodonta_anatina
|
+----- Anodonta_cygnea
--+
+----- Pseudanodonta
|
| +-- Cristariina
+--+
+-- Unio **
Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 13 | 4 | 1 | 22 | +----- Anodontini
--+
| +-- Aculamprotula **
+--+
+-- Unio **
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 12 | 2 | 2 | 20 | +-------- Lanceolariini
--+
| +----- Aculamprotula **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Soroka, 2010 [Publ 3404], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 5 | 5 | 1 | 24 | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Unio **
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 5 | 2 | 17 | 9 | 2 | 22 | +-------- Anodontini
--+
| +----- Lanceolariini
+--+
| +-- Nodularia **
+--+
+-- Inversiunio **
Nagel et al., 1998 [Publ 1946], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 5 | 3 | 1 | 22 | +-- Anodontini
--+
+-- Unio **
Wu et al., 2024 [Publ 4388], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 5 | 0 | 0 | 23 | Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 11 | 7 | 2 | 20 | +-------------- Anodontini
--+
| +----------- Lanceolariini
+--+
| +-------- Aculamprotula **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Unio **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 20 | 4 | 1 | 24 | +-- Unio **
--+
+-- Anodontini
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 21 | 1 | 1 | 21 | +----- Acuticostini
--+
| +-- Nodularia **
+--+
+-- Inversiunio **
Källersjö et al., 2005 [Publ 2684], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 1 | 4 | 3 | 1 | 21 | Kilikowska et al., 2020 [Publ 4335], fig. 4a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 23 | Soroka, 2010 [Publ 3404], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 1 | 4 | 4 | 1 | 23 | +----- Cristariina
--+
| +-- Unio **
+--+
+-- Anodontina
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 16 | 4 | 1 | 24 | +-- Unio **
--+
+-- Anodontini
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 2 | 2 | 31 | 12 | 2 | 20 | +-------- Anodontini
--+
| +----- Schistodesmus **
+--+
| +-- Lepidodesmini
+--+
+-- Aculamprotula **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 4 | 3 | 1 | 21 | Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 9 | 0 | 2 | 12 | 1 | 1 | 28 | +-- Aculamprotula **
|
+-- Lamprotula
|
+-- Lampsilini
|
+-- Potomida
|
--+-- Unio_pictorum **
|
+-- Anodontini
|
+-- Unio_ravoisieri **
|
+-- Lamellidentini
|
+-- Pleurobemini
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 2 | 2 | 1 | 21 | +-- Anodontini
--+
+-- Unio **
Bogatov et al., 2018 [Publ 3859], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 1 | 3 | 2 | 1 | 23 | +-------- Unio_crassus **
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Anodontini
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 54 | 5 | 2 | 22 | +----------- Lanceolariini
--+
| +-------- Anodontini
+--+
| +----- Unio **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 3 | 3 | 10 | 6 | 3 | 22 | +----------- Anodontini
|
--+ +-- Nodularia **
| +-----+
| | +-- Cuneopsis **
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticostini
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula **
Bogatov et al., 2018 [Publ 3859], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 1 | 3 | 2 | 1 | 23 | +-------- Unio_tumidus **
--+
| +----- Anodontini
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3b-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 23 | +----- Anodontina
--+
| +-- Cristariina
+--+
+-- Unio **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 2 | 13 | 1 | 1 | 24 | +----------- Aculamprotula **
--+
| +-------- Unio_pictorum **
+--+
| +----- Anodontini
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidentini
Soroka, 2010 [Publ 3404], fig. 8b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 23 | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Unio **
Soroka, 2010 [Publ 3404], fig. 8a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 23 | +-- Unio **
--+
+-- Anodontini
Soroka, 2010 [Publ 3404], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 4 | 4 | 1 | 23 | +-- Anodonta_anatina
+-----+
| +-- Unio **
--+
| +----- Cristariina
+--+
| +-- Anodonta_cygnea
+--+
+-- Pseudanodonta
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 10 | 3 | 1 | 22 | +----- Anodontini
--+
| +-- Aculamprotula **
+--+
+-- Unio **
Soroka, 2010 [Publ 3404], fig. 3a-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 3 | 1 | 23 | +----- Anodontina
--+
| +-- Cristariina
+--+
+-- Unio **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 15 | 6 | 2 | 21 | Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 54 | 5 | 2 | 22 | +----------- Anodontini
--+
| +-------- Lanceolariini
+--+
| +----- Unio **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Zieritz et al., 2012 [Publ 3673], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 5 | 4 | 1 | 21 | Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 2 | 3 | 11 | 1 | 1 | 23 | +-- Schistodesmus **
+--+
| +-- Aculamprotula **
--+
| +-- Anodontini
+--+
+-- Unio **
Soroka, 2010 [Publ 3404], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 5 | 5 | 1 | 23 | +-------- Anodonta_anatina
|
--+ +----- Cristariina
| |
+--+----- Unio **
|
| +-- Anodonta_cygnea
+--+
+-- Pseudanodonta
Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 10 | 5 | 1 | 21 | +-- Unio **
--+
+-- Anodontini
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 16 | 4 | 1 | 24 | +-- Unio **
--+
+-- Anodontini
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 10 | 3 | 1 | 22 | +----- Anodontini
--+
| +-- Aculamprotula **
+--+
+-- Unio **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 0 | 2 | 12 | 1 | 1 | 23 | +-- Anodontini
|
+-- Unio_ravoisieri **
--+
+-- Unio_pictorum **
|
+-- Aculamprotula **
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 2 | 2 | 13 | 4 | 2 | 20 | +-------- Anodontini
--+
| +----- Aculamprotula **
+--+
| +-- Lepidodesmini
+--+
+-- Unio **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 14 | 5 | 3 | 31 | 11 | 4 | 32 | +-------- Strophitus
|
+-------- Schistodesmus **
|
+-------- Lepidodesmini
|
+-------- Acuticostini
|
+-------- Cristariina
--+
+-------- Lanceolariini
|
| +-- Nodularia **
+-----+
| +-- Tchangsinaia **
|
| +----- Utterbackia
| |
+--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 7 | 5 | 3 | 31 | 11 | 4 | 25 | +----------- Anodontini
|
| +----- Acuticostini
--+ +--+
| | | +-- Nodularia **
| | +--+
+--+ +-- Tchangsinaia **
|
| +----- Lanceolariini
+--+
| +-- Schistodesmus **
+--+
+-- Lepidodesmini
Froufe et al., 2017 [Publ 3802], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 7 | 4 | 1 | 23 | +-- Unio **
--+
+-- Anodontini
Soroka, 2010 [Publ 3404], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 5 | 5 | 1 | 23 | +-- Unio **
--+
+-- Anodontini
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 9 | 7 | 2 | 22 | 6 | 1 | 28 | +-------------- Nodularia **
|
--+ +----------- Anodonta_cygnea
| |
+--+ +-------- Unio_pictorum__2 **
| |
+--+ +-- Unio_pictorum__1 **
| +--+
| | +-- Unio_caffer **
+--+
| +-- Anodonta
+--+
+-- Alasmidontina
Soroka, 2010 [Publ 3404], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 2 | 2 | 1 | 23 | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Unio **
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 10 | 0 | 2 | 23 | 8 | 1 | 28 | +----- Unio **
--+
| +-- Aculamprotula **
+--+
+-- Anodontini
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 4 | 2 | -- | +----- Lanceolariini
--+
| +-- Inversiunio **
+--+
+-- Nodularia **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 8 | 2 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 17 | 15 | 2 | -- | +----- Anodontini
--+
| +-- Lanceolariini
+--+
+-- Unio **
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 17 | 0 | 2 | 20 | 4 | 1 | 35 | +----- Anodontina
|
| +-- Gonideini
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula **
|
+-- Unio **
Lopes-Lima et al., 2020 [Publ 4080], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 10 | 8 | 1 | -- | +-- Nodularia **
--+
+-- Lanceolariini
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 7 | 2 | -- | +-------- Anodontini
--+
| +----- Lepidodesmini
+--+
| +-- Schistodesmus **
+--+
+-- Unio **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 16 | 2 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 3 | 1 | -- | +----- Anodontini
--+
| +-- Unio **
+--+
+-- Aculamprotula **
Wu et al., 2025 [Publ 4442], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 33 | 33 | 1 | -- | +-- Cuneopsis **
--+
+-- Anodontini
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 36 | 14 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 44 | 4 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 45 | 4 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Soroka, 2010 [Publ 3404], fig. 8b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 1 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 2 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 28 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 3 | 1 | -- | Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 44 | 4 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 2 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 32 | 3 | 1 | -- | Huang et al., 2018 [Publ 3828], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 27 | 0 | 0 | -- | +-- Unio **
--+
+-- Aculamprotula **
Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 28 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 28 | 6 | 1 | -- | Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 1 | 29 | 3 | 1 | 25 | +-- Unio_pictorum **
|
--+-- Unio_tumidus **
|
+-- Anodontini
Soroka, 2010 [Publ 3404], fig. 8a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 1 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Tomilova et al., 2020 [Publ 4097], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 6 | 1 | -- | +-- Unio **
|
--+-- Cristariina
|
+-- Anodontina
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 45 | 4 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 10 | 6 | 2 | 21 | +-------- Anodontini
--+
| +----- Nodularia **
+--+
| +-- Lanceolariini
+--+
+-- Inversiunio **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 28 | 6 | 1 | -- | Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 15 | 15 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 23 | 8 | 1 | -- | +----- Anodontini
--+
| +-- Aculamprotula **
+--+
+-- Unio **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 3 | 1 | -- | +----- Anodontini
--+
| +-- Unio **
+--+
+-- Aculamprotula **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 1 | 29 | 3 | 1 | 25 | +----- Unio_pictorum **
--+
| +-- Unio_tumidus **
+--+
+-- Anodontini
Bolotov et al., 2015 [Publ 3646], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 3 | 1 | -- | Combosch et al., 2017 [Publ 3806], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | Nagel et al., 1998 [Publ 1946], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 2 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 17 | 1 | -- | +----------------- Anodonta_cygnea
|
+----------------- Pyganodon
|
+----------------- Alasminota
|
+----------------- Anodonta_anatina
|
+----------------- Unio **
|
+----------------- Cristariina
|
| +-- Anodonta_nuttalliana
--+--------------+
| +-- Anodonta_californiensis
|
| +-- Pseudodontoideus
| |
+--------------+-- Alasmidonta
| |
| +-- Lasmigona
|
| +-------------- Pseudanodonta
| |
| | +----- Strophitus
| +--------+
+--+ | +-- Prolasmidonta
| +--+
| +-- Platynaias
|
| +----------- Anodonta_oregonensis__4
+--+
| +-------- Anodonta_oregonensis__3
+--+
| +----- Anodonta_oregonensis__2
+--+
| +-- Anodonta_oregonensis__1
+--+
+-- Anodonta_kennerlyi
Combosch et al., 2017 [Publ 3806], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
González & Giribet, 2015 [Publ 3632], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- | +-- Unio **
--+
+-- Anodontini
González & Giribet, 2015 [Publ 3632], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Graf et al., 2015 [Publ 3619], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 4 | 1 | -- | Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 3 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 6 | 1 | -- | +-- Tchangsinaia **
--+
+-- Anodontini
González & Giribet, 2015 [Publ 3632], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Araujo et al., 2009 [Publ 2395], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 28 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 2 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Lopes-Lima et al., 2018 [Publ 3829], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Wang et al., 2016 [Publ 3714], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Klunzinger et al., 2021 [Publ 4086], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 28 | 1 | -- | +-- Schistodesmus **
--+
+-- Anodontini
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Bolotov et al., 2015 [Publ 3646], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Vikhrev et al., 2018 [Publ 3812], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- | Taylor et al., 2007 [Publ 2696], fig. 1-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 6 | 1 | -- | +-- Unio **
--+
+-- Anodontini
Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Graf et al., 2014 [Publ 3591], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Anodontini
--+
+-- Unio **
|
