MUSSELpdb | subfamily Velesunioninae Iredale, 1934 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily VELESUNIONINAE Iredale, 1934: cladomics
6 genera

Cladomics Summary

number of species = 17 (21 total)
number of subtaxa (rank: genus) = 6
analyzed in 65 cladograms in 38 publications
maximum number of terminals in an analysis = 10 (mean = 2.38, median = 1)
maximum number of subtaxa (genus) = 5 (mean = 1.86, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 10, (mean = 4.33 median = 3)
maximum number of subtaxa (genus) = 5 (mean = 3.07, median = 2)
proportion of cladograms that support basic monophyly = 0.67

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 26
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 10 (mean = 4.42, median = 3)
maximum number of subtaxa (genus) = 5 (mean = 3.15, median = 2)
proportion of cladograms that support rigorous monophyly = 0.65

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
10	28	34	5	622	18	1	0

  +----------------------------------- Westralunio **           
  |                                                          
  |                             +----- Diplodon_suavidicus__2
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini               
  |  |                          +--+                         
  |  |                             +-- Castaliini            
  |  |                                                       
  |  |                             +-- Hyridella_depressa    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus       
        |                                                    
        |                          +-- Cucumerunio           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis   
        |  |                                                 
        +--+  +----------------------- Echyridella           
           |  |                                              
           |  |              +-------- Velesunio__1 **          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 ** 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus **    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga **     
                 |                                           
                 |  +----------------- Velesunio__3 **          
                 |  |                                        
                 +--+  +-------------- Microdontia **           
                    |  |                                     
                    +--+           +-- Lortiella **             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi **      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 ** 
                          +--+                               
                             |  +----- Velesunio__2 **          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 ** 
                                +--+                         
                                   +-- Alathyria_pertexta__1 **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
7	14	12	5	194	7	1	39

           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae          
  |                              
--+     +----- Alathyria_jacksoni **
  |  +--+                        
  |  |  |  +-- Alathyria_profuga ** 
  |  |  +--+                     
  +--+     +-- Velesunio **         
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Graf et al., 2015 [Publ 3619], fig. 1

tT	bi	riS	oT	roT	roS	iD
7	7	5	5	13	7	1	36

  +-------------- Hyriinae          
  |                                 
--+  +----------- Westralunio **       
  |  |                              
  |  |     +----- Alathyria_jacksoni **
  +--+  +--+                        
     |  |  |  +-- Velesunio **         
     |  |  +--+                     
     +--+     +-- Alathyria_profuga ** 
        |                           
        |  +----- Alathyria_pertexta **
        +--+                        
           |  +-- Microdontia **       
           +--+                     
              +-- Lortiella **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
7	14	12	5	192	7	1	41

           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae          
  |                              
--+     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
7	14	9	5	194	7	1	38

  +----------- Echyridella       
  |                              
  +----------- Westralunio **       
  |                              
  +----------- Hyridellini       
  |                              
  |     +----- Rhipidodontini    
  +-----+                        
--+     |  +-- Castaliini        
  |     +--+                     
  |        +-- Hyriini           
  |                              
  |     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	133	3	1	39

  +----------- Hyriinae          
  |                              
  |     +----- Alathyria_pertexta **
--+  +--+                        
  |  |  |  +-- Microdontia **       
  |  |  +--+                     
  +--+     +-- Lortiella **         
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	13	11	5	14	13	1	31.5

                    +----- Diplodon_suavidicus__2
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio **           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1
  +--+              +--+                         
     |                 +-- Diplodon_gratus       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio           
        |  +-----------+                         
        |  |           +-- Hyridella_australis   
        +--+                                     
           |  +----------- Velesunio_ambiguus **    
           +--+                                  
              |  +-------- Alathyria **             
              +--+                               
                 |  +----- Velesunio_angasi **      
                 +--+                            
                    |  +-- Lortiella **             
                    +--+                         
                       +-- Microdontia **

Bolotov et al., 2016 [Publ 3743], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	24	3	1	39

  +----------- Hyriinae          
  |                              
  |     +----- Microdontia **       
--+  +--+                        
  |  |  |  +-- Alathyria_pertexta **
  |  |  +--+                     
  +--+     +-- Lortiella **         
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

tT	bi	riS	oT	roT	roS	iD
6	19	20	5	14	13	1	22.5

                 +----- Diplodon_suavidicus__1
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio **           
  |           |                               
  +-----------+     +-- Hyridella_drapeta     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa    
  |           +--+                            
  |              |  +-- Diplodon_gratus       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2
  |                                           
  |                 +-- Hyridella_australis   
  |  +--------------+                         
  |  |              +-- Cucumerunio           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus **    
        +--+                                  
           |  +-------- Microdontia **           
           +--+                               
              |  +----- Lortiella **             
              +--+                            
                 |  +-- Alathyria **             
                 +--+                         
                    +-- Velesunio_angasi **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	133	3	1	39

  +----------- Hyriinae          
  |                              
  |     +----- Velesunio **         
--+  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Microdontia **       
     +--+                        
        |  +-- Alathyria_pertexta **
        +--+                     
           +-- Lortiella **

Bolotov et al., 2016 [Publ 3743], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	24	3	1	39

  +----------- Hyriinae          
  |                              
  |     +----- Microdontia **       
--+  +--+                        
  |  |  |  +-- Lortiella **         
  |  |  +--+                     
  +--+     +-- Alathyria_pertexta **
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	4	3	31	5	1	30

  +----------------- Castaliini         
  |                                     
  +----------------- Rhipidodontini     
--+                                     
  |  +-------------- Hyridella_depressa 
  |  |                                  
  |  +-------------- Hyridella_australis
  +--+                                  
     |  +----------- Echyridella        
     |  |                               
     |  |        +-- Velesunio_ambiguus ** 
     +--+  +-----+                      
        |  |     +-- Velesunio__3 **       
        |  |                            
        +--+  +----- Velesunio__2 **       
           |  |                         
           |  |  +-- Alathyria **          
           +--+--+                      
              |  +-- Velesunio__1 **       
              |                         
              |  +-- Velesunio_angasi **   
              +--+                      
                 +-- Lortiella **

Klunzinger et al., 2021 [Publ 4086], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	4	2	2	3	1	1	32

  +----- Westralunio **
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyriinae

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	44	6	1	33

  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	43	6	1	33

  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Klunzinger et al., 2022 [Publ 4259], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	1	1	1	33

  +----- Hyriinae   
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Westralunio **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	44	6	1	33

  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	43	6	1	33

  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Baker et al., 2004 [Publ 2685], fig. 2-3

tT	bi	riS	oT	roT	roS	iD
3	3	4	2	6	5	1	32

              +----- Hyridella_depressa 
  +-----------+                         
  |           |  +-- Hyridella_australis
  |           +--+                      
  |              +-- Rhipidodontini     
--+                                     
  |              +-- Echyridella        
  |  +-----------+                      
  |  |           +-- Castaliini         
  |  |                                  
  +--+           +-- Velesunio__3 **       
     |  +--------+                      
     |  |        +-- Velesunio__4 **       
     +--+                               
        |  +-------- Velesunio__2 **       
        +--+                            
           |  +----- Velesunio_angasi **   
           +--+                         
              |  +-- Velesunio__1 **       
              +--+                      
                 +-- Alathyria **

Marshall et al., 2014 [Publ 3593], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	11	7	1	34.5

  +----------- Echyridella       
  |                              
--+        +-- Hyridellini       
  |  +-----+                     
  |  |     +-- Rhipidodontini    
  +--+                           
     |  +----- Velesunio_ambiguus **
     +--+                        
        |  +-- Velesunio **         
        +--+                     
           +-- Lortiella **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	3	1	--

        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Hoeh et al., 2002 [Publ 2653], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	3	1	--

        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	3	1	--

        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	3	1	--

        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Lortiella **     
     +--+                 
        +-- Velesunio **

Baker et al., 2003 [Publ 2690], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	6	5	1	--

           +-- Hyridella_australis
  +--------+                      
  |        +-- Rhipidodontini     
--+                               
  |  +-------- Hyridella_depressa 
  +--+                            
     |  +----- Velesunio **          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castaliini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

tT	bi	riS	oT	roT	roS	iD
2	5	0	2	29	3	1	36

     +-- Castaliini    
  +--+                 
  |  +-- Velesunio **     
--+                    
  |  +-- Echyridella   
  |  |                 
  +--+-- Lortiella **     
     |                 
     +-- Rhipidodontini

Hoeh et al., 2001 [Publ 2652], fig. 14f2

tT	bi	riS	oT	roT	roS	iD
2	5	0	2	29	3	1	35

  +----- Rhipidodontini
  |                    
  +----- Echyridella   
--+                    
  +----- Lortiella **     
  |                    
  |  +-- Castaliini    
  +--+                 
     +-- Velesunio **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	1	1	--

  +-- Hyriinae 
--+            
  +-- Velesunio **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	3	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Combosch et al., 2017 [Publ 3806], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

González & Giribet, 2015 [Publ 3632], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Hyriinae   
--+              
  +-- Westralunio **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Velesunio_ambiguus **

Combosch et al., 2017 [Publ 3806], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Alathyria_jacksoni **

González & Giribet, 2015 [Publ 3632], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

Bieler et al., 2014 [Publ 3622], fig. 28

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	5	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Sonowal et al., 2021 [Publ 4221], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Hyriinae   
--+              
  +-- Westralunio **

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	80	1	1	--

  +-- Hyriinae 
--+            
  +-- Velesunio **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Sonowal et al., 2021 [Publ 4221], fig. 3a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

  +-- Hyriini    
  |              
--+-- Hyridellini
  |              
  +-- Velesunio **

Elderkin et al., 2016 [Publ 3693], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	0	0	--

-- Velesunio_ambiguus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	139	4	1	--

  +-- Westralunio **
--+              
  +-- Hyriinae

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	2	1	--

  +-- Alathyria **
--+            
  +-- Hyriinae

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	2	1	--

  +-- Alathyria **
--+            
  +-- Hyriinae

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	4	1	--

        +-- Echyridella   
  +-----+                 
  |     +-- Westralunio **   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Hyriini

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	2	1	--

  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

González & Giribet, 2015 [Publ 3632], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

Graf, 2000 [Publ 2647], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	3	1	--

  +-------- Castaliini    
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Hyridellini   
     +--+                 
        +-- Velesunio **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	3	1	--

  +-- Velesunio **
--+            
  +-- Hyriinae

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	1	1	--

  +-- Hyriinae   
--+              
  +-- Westralunio **

Sonowal et al., 2021 [Publ 4221], fig. 3b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

  +-- Hyriini    
  |              
--+-- Velesunio **  
  |              
  +-- Hyridellini

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	4	1	--

  +-- Westralunio **
--+              
  +-- Hyriinae

González & Giribet, 2015 [Publ 3632], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

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