MUSSELpdb | valid gen. Cristaria cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Cristaria Schumacher, 1817: cladomics
7 Recent valid species (10 total species)

Cladomics Summary

number of species = 7 (10 total)
analyzed in 123 cladograms in 83 publications
maximum number of terminals in an analysis = 7 (mean = 1.39, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 7, (mean = 3.53 median = 3)
proportion of cladograms that support basic monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Dai et al., 2026 [Publ 4757], fig. 3

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	7	6	9	27	23	6	0

                    +----- Amuranodonta         
  +-----------------+                           
  |                 |  +-- Anemina              
  |                 +--+                        
  |                    +-- Buldowskia           
--+                                             
  |                 +----- Pletholophus         
  |  +--------------+                           
  |  |              |  +-- Beringiana           
  |  |              +--+                        
  +--+                 +-- Sinanodonta          
     |                                          
     |  +----------------- Cristaria_wangyucheni **
     |  |                                       
     +--+  +-------------- Cristaria_bellua **     
        |  |                                    
        +--+  +----------- Cristaria_clessini **   
           |  |                                 
           +--+        +-- Cristaria_plicata **    
              |  +-----+                        
              |  |     +-- Cristaria_truncata **   
              +--+                              
                 |  +----- Cristaria_radiata__2 ** 
                 |  |                           
                 +--+----- Cristaria_radiata__3 ** 
                    |                           
                    |  +-- Cristaria_liboi **      
                    +--+                        
                       +-- Cristaria_radiata__1 **

Chen et al., 2026 [Publ 4805], fig. 4

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	28	27	8	5

        +----------- Cristaria_clessini **
        |                              
  +-----+  +-------- Cristaria_bellua **  
  |     |  |                           
  |     +--+     +-- Cristaria_plicata ** 
  |        |  +--+                     
  |        |  |  +-- Cristaria_truncata **
  |        +--+                        
--+           |  +-- Cristaria_radiata ** 
  |           +--+                     
  |              +-- Cristaria_liboi **   
  |                                    
  |  +-------------- Beringiana        
  |  |                                 
  |  |        +----- Mimo              
  +--+  +-----+                        
     |  |     |  +-- Acudonta          
     |  |     +--+                     
     +--+        +-- Sinanodonta       
        |                              
        |  +-------- Pletholophus      
        +--+                           
           |  +----- Anemina           
           +--+                        
              |  +-- Amuranodonta      
              +--+                     
                 +-- Buldowskia

Wu et al., 2025 [Publ 4442], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	29	27	7	5

                 +-- Cristaria_clessini **         
           +-----+                              
           |     +-- Cristaria_bellua **           
           |                                    
  +--------+     +-- Cristaria_plicata **          
  |        |  +--+                              
  |        |  |  +-- Cristaria_truncata **         
  |        +--+                                 
  |           |  +-- Cristaria_radiata **          
  |           +--+                              
--+              +-- Cristaria_liboi **            
  |                                             
  |           +----- Beringiana                 
  |  +--------+                                 
  |  |        |  +-- Acudonta                   
  |  |        +--+                              
  |  |           +-- Sinanodonta                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus     
        |                                       
        |  +-------- Pletholophus_honglinhensis 
        +--+                                    
           |  +----- Anemina                    
           +--+                                 
              |  +-- Amuranodonta               
              +--+                              
                 +-- Buldowskia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	51	8	5	8

           +----- Anemina           
  +--------+                        
  |        |  +-- Amuranodonta      
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |           +-- Beringiana        
  |  +--------+                     
  |  |        +-- Sinanodonta       
  +--+                              
     |  +-------- Cristaria_radiata ** 
     |  |                           
     +--+     +-- Cristaria_truncata **
        |  +--+                     
        |  |  +-- Cristaria_plicata ** 
        +--+                        
           |  +-- Cristaria_bellua **  
           +--+                     
              +-- Cristaria_clessini **

Dai et al., 2024 [Publ 4385], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	27	23	6	5

           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	3	2	2	7

           +-- Sinanodonta       
  +--------+                     
  |        +-- Pletholophus      
--+                              
  |  +-------- Cristaria_radiata ** 
  +--+                           
     |  +----- Cristaria_clessini **
     +--+                        
        |  +-- Cristaria_plicata ** 
        +--+                     
           +-- Cristaria_truncata **

Dai et al., 2024 [Publ 4385], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	27	23	6	5

           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Xiang et al., 2025 [Publ 4433], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	24	23	7	5

  +----------------- Pletholophus      
  |                                    
  |           +----- Beringiana        
--+  +--------+                        
  |  |        |  +-- Mimo              
  |  |        +--+                     
  |  |           +-- Sinanodonta       
  +--+                                 
     |        +----- Anemina           
     |  +-----+                        
     |  |     |  +-- Amuranodonta      
     |  |     +--+                     
     +--+        +-- Buldowskia        
        |                              
        |  +-------- Cristaria_bellua **  
        +--+                           
           |  +----- Cristaria_clessini **
           +--+                        
              |  +-- Cristaria_plicata ** 
              +--+                     
                 +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	27	23	6	5

     +-------- Cristaria_bellua **  
  +--+                           
  |  |  +----- Cristaria_clessini **
  |  +--+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
--+        +-- Cristaria_truncata **
  |                              
  |     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  +--+     +-- Buldowskia        
     |                           
     |  +----- Pletholophus      
     +--+                        
        |  +-- Beringiana        
        +--+                     
           +-- Sinanodonta

Wu et al., 2025 [Publ 4442], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	14	12	7	6

        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	4	2	1	6

  +-------- Sinanodonta       
--+                           
  |  +----- Cristaria_clessini **
  +--+                        
     |  +-- Cristaria_plicata ** 
     +--+                     
        +-- Cristaria_radiata **

Wang et al., 2025 [Publ 4488], fig. 3B

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	62	12	7	6

        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	10	8	3	--

  +-------- Anemina          
  |                          
--+     +-- Cristaria_plicata **
  |  +--+                    
  |  |  +-- Cristaria_radiata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	17	0	2	123	4	2	22

  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata **    
  |                        
  +-- Cristaria_radiata **    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Klishko et al., 2016 [Publ 3733], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	2	2	2	--

     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Beringiana

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	120	22	6	--

        +-- Cristaria_plicata ** 
  +-----+                     
  |     +-- Cristaria_clessini **
  |                           
  |  +----- Anemina           
  +--+                        
--+  |  +-- Amuranodonta      
  |  +--+                     
  |     +-- Buldowskia        
  |                           
  |  +----- Pletholophus      
  +--+                        
     |  +-- Beringiana        
     +--+                     
        +-- Sinanodonta

Klishko et al., 2014 [Publ 3666], fig. 8

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	2	2	2	--

     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	137	4	2	--

     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	137	4	2	--

     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Wu et al., 2016 [Publ 3711], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	2	2	--

-- Cristaria_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	22	0	3	33	9	5	36

  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1 **   
  |                             
  +----- Cristaria_plicata__2 **   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3 **   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	5	5	--

-- Cristaria_plicata **

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	4	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wang et al., 2018 [Publ 3846], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	4	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	43	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	5	5	--

-- Cristaria_plicata **

Chen et al., 2012 [Publ 3624], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	6	0	0	--

-- Cristaria_plicata **

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	43	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	2	2	--

  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	10	5	--

-- Cristaria_plicata **

Riccardi et al., 2020 [Publ 4089], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Sheng et al., 2014 [Publ 3704], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Cristaria_plicata **

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	5	2	--

  +-- Anemina          
  |                    
--+-- Cristaria_plicata **
  |                    
  +-- Sinanodonta

Sano et al., 2022 [Publ 4311], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	17	4	--

        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	2	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	5	5	--

  +----------- Cristaria_plicata **
  |                             
--+        +-- Anemina          
  |  +-----+                    
  |  |     +-- Amuranodonta     
  +--+                          
     |  +----- Pletholophus     
     +--+                       
        |  +-- Sinanodonta      
        +--+                    
           +-- Beringiana

Fonseca et al., 2016 [Publ 3775], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	5	5	--

  +----------- Cristaria_plicata **
  |                             
--+        +-- Anemina          
  |  +-----+                    
  |  |     +-- Amuranodonta     
  +--+                          
     |  +----- Pletholophus     
     +--+                       
        |  +-- Sinanodonta      
        +--+                    
           +-- Beringiana

Wei et al., 2016 [Publ 3665], fig. 0

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Cristaria_plicata **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Smith et al., 2018 [Publ 3790], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Cristaria_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	3	2	--

  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	0	0	--

-- Cristaria_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	5	3	--

     +-- Anemina          
  +--+                    
  |  +-- Cristaria_plicata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	2	2	--

  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Wu et al., 2023 [Publ 4339], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	3	2	--

  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	2	2	--

  +----- Cristaria_radiata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	1	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Cristaria_plicata **

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Xue et al., 2016 [Publ 3659], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	3	2	--

-- Cristaria_plicata **

Huang et al., 2018 [Publ 3828], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	2	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	10	5	--

-- Cristaria_plicata **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	7	2	--

-- Cristaria_plicata **

Cha et al., 2018 [Publ 3916], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Xia et al., 2016 [Publ 3712], fig. 7B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	1	1	--

-- Cristaria_plicata **

Sano et al., 2022 [Publ 4311], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	11	4	--

        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia

Sano et al., 2017 [Publ 3799], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	7	4	--

        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Sinanodonta      
     +--+                    
        +-- Beringiana

Zhou et al., 2007 [Publ 2658], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	1	1	--

-- Cristaria_plicata **

Zhang et al., 2013 [Publ 4214], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	1	1	--

-- Cristaria_plicata **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	4	2	--

  +-------------- Cristaria_plicata **       
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 
     |  |                                 
     |  |  +----- Beringiana_japonica__1  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3  
        |  |                              
        |  +----- Beringiana_japonica__4  
        |  |                              
        |  +----- Beringiana_japonica__5  
        |  |                              
        |  +----- Sinanodonta_lauta__1    
        |  |                              
        |  +----- Beringiana_japonica__6  
        |  |                              
        |  +----- Sinanodonta_lauta__2    
        |  |                              
        |  +----- Sinanodonta_calipygos__1
        |  |                              
        +--+----- Beringiana_japonica__7  
           |                              
           +----- Sinanodonta_lauta__4    
           |                              
           +----- Beringiana_japonica__8  
           |                              
           +----- Beringiana_japonica__9  
           |                              
           +----- Beringiana_japonica__10 
           |                              
           +----- Beringiana_japonica__11 
           |                              
           +----- Beringiana_japonica__12 
           |                              
           +----- Beringiana_japonica__13 
           |                              
           +----- Beringiana_japonica__14 
           |                              
           |  +-- Sinanodonta_lauta__3    
           +--+                           
              +-- Sinanodonta_calipygos__2

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	2	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	4	2	--

  +----- Cristaria_truncata **
--+                        
  |  +-- Sinanodonta       
  +--+                     
     +-- Pletholophus

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	3	2	--

  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	3	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	4	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	82	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Zhou et al., 2016 [Publ 3710], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	1	1	--

-- Cristaria_plicata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Pletholophus

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Cristaria_plicata **

Guerra et al., 2017 [Publ 3772], fig. 2a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	1	1	--

-- Cristaria_plicata **

Jiang et al., 2010 [Publ 3438], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	0	0	--

-- Cristaria_plicata **

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	2	2	--

  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Bolotov et al., 2020 [Publ 4084], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	8	5	--

     +----- Amuranodonta     
  +--+                       
  |  |  +-- Anemina          
  |  +--+                    
--+     +-- Buldowskia       
  |                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	200	8	3	--

  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Breton et al., 2011 [Publ 3376], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	0	0	--

-- Cristaria_plicata **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	5	5	--

-- Cristaria_plicata **

Xiang et al., 2025 [Publ 4433], fig. 7

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	6	4	--

  +----------- Cristaria_plicata **
--+                             
  |  +-------- Anemina          
  +--+                          
     |  +----- Beringiana       
     +--+                       
        |  +-- Mimo             
        +--+                    
           +-- Sinanodonta

Peng et al., 2012 [Publ 3689], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	0	0	--

-- Cristaria_plicata **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	31	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	1	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 4_6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	2	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	11	2	--

  +----- Pletholophus      
--+                        
  |  +-- Cristaria_truncata **
  +--+                     
     +-- Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	5	5	--

-- Cristaria_plicata **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	2	2	--

-- Cristaria_plicata **

Wen et al., 2017 [Publ 3795], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	1	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Chen et al., 2020 [Publ 4170], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	2	--

-- Cristaria_plicata **

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	8	3	--

  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	200	8	3	--

-- Cristaria_plicata **

Sano et al., 2017 [Publ 3799], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	8	4	--

        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Beringiana       
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Wang et al., 2018 [Publ 3865], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	3	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	2	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	7	2	--

-- Cristaria_plicata **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	4	2	--

  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Guerra et al., 2017 [Publ 3772], fig. 2b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	1	1	--

  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	5	3	--

  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Huang et al., 2013 [Publ 3629], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	1	1	--

  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	10	5	--

-- Cristaria_plicata **

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