Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina

genus Inversiunio Habe, 1991: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 26 cladograms in 14 publications
maximum number of terminals in an analysis = 4 (mean = 2.12, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 17
maximum number of included terminals = 4, (mean = 2.71 median = 3)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Inversiunio cladogram distances
PCoA Ordination

NJ Tree of Inversiunio cladogram distances
NJ Tree

Cladograms

Sano et al., 2020 [Publ 4138], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
2 2 0 6 32 4 3 -- 
  +----- Schistodesmus              
  |                                 
  |  +-- Inversiunio_reinianus__1 **   
--+  |                              
  |  +-- Inversiunio_reinianus__2 **   
  |  |                              
  |  +-- Inversiunio_jokohamensis__1 **
  +--+                              
     +-- Inversiunio_jokohamensis__2 **
     |                              
     +-- Inversiunio_jokohamensis__3 **
     |                              
     +-- Inversiunio_jokohamensis__4 **

Chen et al., 2023 [Publ 4302], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Inversiunio_yanagawensis **
--+                           
  +-- Inversiunio_reinianus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
4 4 2 4 118 14 4 -- 
  +----------------- Nodularia               
  |                                          
--+  +-------------- Schistodesmus           
  |  |                                       
  +--+-------------- Cuneopsis               
     |                                       
     |  +----------- Tchangsinaia            
     +--+                                    
        |  +-------- Inversiunio_jokohamensis **
        +--+                                 
           |  +----- Inversiunio_verrucosus **  
           +--+                              
              |  +-- Inversiunio_reinianus **   
              +--+                           
                 +-- Inversiunio_yanagawensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iTtTbiriSoTroTroSiD
4 4 2 5 4 2 1 -- 
  +----------- Nodularia                
  |                                     
--+  +-------- Inversiunio_jokohamensis ** 
  |  |                                  
  +--+  +----- Inversiunio_verrucosus__1 **
     |  |                               
     +--+----- Inversiunio_verrucosus__2 **
        |                               
        |  +-- Inversiunio_reinianus **    
        +--+                            
           +-- Inversiunio_yanagawensis **

Sayenko et al., 2021 [Publ 4189], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 5 9 6 1 -- 
  +-------- Inversiunio_jokohamensis **   
  |                                    
--+  +----- Inversiunio_yanagawensis__2 **
  |  |                                 
  +--+----- Inversiunio_verrucosus **     
     |                                 
     |  +-- Inversiunio_reinianus **      
     +--+                              
        +-- Inversiunio_yanagawensis__1 **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
3 3 4 6 18 1 1 -- 
  +----------------- Nodularia                  
--+                                             
  |  +-------------- Inversiunio_jokohamensis **   
  +--+                                          
     |  +----------- Inversiunio_yanagawensis__1 **
     +--+                                       
        |  +-------- Inversiunio_yanagawensis__2 **
        +--+                                    
           |  +----- Inversiunio_yanagawensis__3 **
           +--+                                 
              |  +-- Inversiunio_yanagawensis__4 **
              +--+                              
                 +-- Inversiunio_reinianus **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 2 4 65 4 3 -- 
        +----- Nodularia                  
  +-----+                                 
  |     |  +-- Schistodesmus              
  |     +--+                              
--+        +-- Tchangsinaia               
  |                                       
  |  +-------- Inversiunio_jokohamensis **   
  +--+                                    
     |  +----- Inversiunio_yanagawensis__2 **
     +--+                                 
        |  +-- Inversiunio_reinianus **      
        +--+                              
           +-- Inversiunio_yanagawensis__1 **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 4 65 4 3 -- 
        +----- Nodularia                  
  +-----+                                 
  |     |  +-- Tchangsinaia               
  |     +--+                              
--+        +-- Schistodesmus              
  |                                       
  |  +-------- Inversiunio_jokohamensis **   
  +--+                                    
     |  +----- Inversiunio_yanagawensis__2 **
     +--+                                 
        |  +-- Inversiunio_reinianus **      
        +--+                              
           +-- Inversiunio_yanagawensis__1 **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 5 30 5 3 -- 
  +-------- Inversiunio_jokohamensis **   
  |                                    
--+  +----- Inversiunio_yanagawensis__3 **
  |  |                                 
  +--+  +-- Inversiunio_yanagawensis__1 **
     |  |                              
     +--+-- Inversiunio_yanagawensis__2 **
        |                              
        +-- Inversiunio_reinianus **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 3 36 12 5 -- 
  +-------------------- Pseudocuneopsis         
--+                                             
  |  +----------------- Nodularia               
  +--+                                          
     |  +-------------- Schistodesmus           
     +--+                                       
        |  +----------- Cuneopsis               
        +--+                                    
           |  +-------- Tchangsinaia            
           +--+                                 
              |  +----- Inversiunio_jokohamensis **
              +--+                              
                 |  +-- Inversiunio_reinianus **   
                 +--+                           
                    +-- Inversiunio_yanagawensis **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 2 4 65 4 3 -- 
  +-------------- Nodularia                  
  |                                          
--+           +-- Schistodesmus              
  |  +--------+                              
  |  |        +-- Tchangsinaia               
  +--+                                       
     |  +-------- Inversiunio_jokohamensis **   
     +--+                                    
        |  +----- Inversiunio_yanagawensis__2 **
        +--+                                 
           |  +-- Inversiunio_reinianus **      
           +--+                              
              +-- Inversiunio_yanagawensis__1 **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Inversiunio_yanagawensis **
--+                           
  +-- Inversiunio_reinianus **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +-- Inversiunio_yanagawensis **
--+                           
  +-- Inversiunio_reinianus **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 22 1 1 -- 
  +----- Nodularia               
--+                              
  |  +-- Inversiunio_jokohamensis **
  +--+                           
     +-- Inversiunio_reinianus **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +----- Tchangsinaia            
--+                              
  |  +-- Inversiunio_yanagawensis **
  +--+                           
     +-- Inversiunio_reinianus **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
           +----- Tchangsinaia            
  +--------+                              
  |        |  +-- Inversiunio_yanagawensis **
  |        +--+                           
--+           +-- Inversiunio_reinianus **   
  |                                       
  |  +----------- Pseudocuneopsis         
  +--+                                    
     |  +-------- Cuneopsis               
     +--+                                 
        |  +----- Nodularia               
        +--+                              
           |  +-- Diaurora                
           +--+                           
              +-- Schistodesmus

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
        +----- Tchangsinaia            
  +-----+                              
  |     |  +-- Inversiunio_yanagawensis **
  |     +--+                           
  |        +-- Inversiunio_reinianus **   
--+                                    
  |        +-- Cuneopsis               
  |  +-----+                           
  |  |     +-- Pseudocuneopsis         
  +--+                                 
     |  +----- Nodularia               
     +--+                              
        |  +-- Diaurora                
        +--+                           
           +-- Schistodesmus

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
-- Inversiunio_reinianus **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 9 5 -- 
  +----------- Cuneopsis               
  |                                    
--+        +-- Nodularia               
  |  +-----+                           
  |  |     +-- Schistodesmus           
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Inversiunio_yanagawensis **
        +--+                           
           +-- Pseudocuneopsis

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Inversiunio_yanagawensis **
--+                           
  +-- Nodularia

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 10 5 -- 
  +----- Cuneopsis               
  |                              
  |  +-- Nodularia               
  +--+                           
--+  +-- Schistodesmus           
  |                              
  |  +-- Tchangsinaia            
  |  |                           
  +--+-- Inversiunio_yanagawensis **
     |                           
     +-- Pseudocuneopsis

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 11 10 5 -- 
  +-------------- Nodularia      
--+                              
  |  +----------- Pseudocuneopsis
  +--+                           
     |  +-------- Schistodesmus  
     +--+                        
        |  +----- Cuneopsis      
        +--+                     
           |  +-- Tchangsinaia   
           +--+                  
              +-- Inversiunio **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 8 3 -- 
  +-------- Nodularia               
--+                                 
  |  +----- Cuneopsis               
  +--+                              
     |  +-- Inversiunio_jokohamensis **
     +--+                           
        +-- Schistodesmus

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 10 5 -- 
  +----------- Cuneopsis               
  |                                    
--+        +-- Nodularia               
  |  +-----+                           
  |  |     +-- Schistodesmus           
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Inversiunio_yanagawensis **
        +--+                           
           +-- Pseudocuneopsis

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 7 2 -- 
  +----- Inversiunio_jokohamensis **
--+                              
  |  +-- Cuneopsis               
  +--+                           
     +-- Nodularia

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 8 3 -- 
  +----- Cuneopsis               
  |                              
--+----- Nodularia               
  |                              
  |  +-- Inversiunio_jokohamensis **
  +--+                           
     +-- Schistodesmus