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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Pseudanodonta Bourguignat, 1876: cladomics
     1 Recent valid species (1 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 1 (1 total)
analyzed in 64 cladograms in 40 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Chong et al., 2008 [Publ 2668], fig. 3

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11011761--
-- Pseudanodonta_complanata **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014411--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011321--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Pseudanodonta_complanata **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
11013041--
  +----------- Anodonta_anatina        
--+                                    
  |  +-------- Pseudanodonta_complanata **
  +--+                                 
     |  +----- Anodonta_exulcerata     
     +--+                              
        |  +-- Anodonta_seddonae       
        +--+                           
           +-- Anodonta_cygnea

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112431--
-- Pseudanodonta_complanata **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
11011221--
-- Pseudanodonta_complanata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015521--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
1101411--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013521--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
11013521--
-- Pseudanodonta_complanata **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta_anatina        
  |                           
--+-- Anodonta_cygnea         
  |                           
  +-- Pseudanodonta_complanata **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
1101431--
-- Pseudanodonta_complanata **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
11011621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119831--
-- Pseudanodonta_complanata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019021--
-- Pseudanodonta_complanata **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119831--
-- Pseudanodonta_complanata **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015241--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
1101321--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Pseudanodonta_complanata **

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
1101431--
-- Pseudanodonta_complanata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113831--
-- Pseudanodonta_complanata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017531--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
1101821--
-- Pseudanodonta_complanata **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014411--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017931--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
1101721--
  +-- Pseudanodonta_complanata **
  |                           
--+-- Anodonta_anatina        
  |                           
  +-- Anodonta_cygnea

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011841--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
11011821--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019441--
        +-- Anodonta_nuttalliana    
  +-----+                           
  |     +-- Pseudanodonta_complanata **
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Anodonta_cygnea         
     +--+                           
        +-- Anodonta_exulcerata

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
1101221--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
1101821--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Pseudanodonta_complanata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113831--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011721--
-- Pseudanodonta_complanata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019041--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
1101731--
  +-------- Anodonta_anatina        
--+                                 
  |  +----- Pseudanodonta_complanata **
  +--+                              
     |  +-- Anodonta_cygnea         
     +--+                           
        +-- Anodonta_exulcerata

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
11014031--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11013911--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1101821--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012300--
-- Pseudanodonta_complanata **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012300--
-- Pseudanodonta_complanata **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019021--
-- Pseudanodonta_complanata **

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
1101221--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017541--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
1101531--
     +-- Anodonta_anatina        
  +--+                           
  |  +-- Pseudanodonta_complanata **
--+                              
  |  +-- Anodonta_exulcerata     
  +--+                           
     +-- Anodonta_cygnea

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112031--
-- Pseudanodonta_complanata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017531--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta_anatina        
  |                           
--+-- Pseudanodonta_complanata **
  |                           
  +-- Anodonta_cygnea

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019441--
-- Pseudanodonta_complanata **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
1101621--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Pseudanodonta_complanata **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
11013041--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta     
--+                
  +-- Pseudanodonta **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012200--
-- Pseudanodonta_complanata **

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
1101621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
1101400--
-- Pseudanodonta_complanata **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11013911--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119631--
-- Pseudanodonta_complanata **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
1101321--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019041--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
1101221--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta
 
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