MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Hamiota Roe & Hartfield, 2005: cladomics
4 Recent valid species (4 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 4 (4 total)
analyzed in 10 cladograms in 8 publications
maximum number of terminals in an analysis = 4 (mean = 2.8, median = 4)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 4, (mean = 4 median = 4)
proportion of cladograms that support basic monophyly = 1
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Roe et al., 2002 [Publ 2689], fig. 3a
** Top Monophyly Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 4 | 10 | 6 | 6 | 4 | 0 |
+-------------- Ligumia | +-------------- Obliquaria | --+-------------- Lampsilis | | +----------- Villosa | | | | +----- Hamiota_australis__2 ** +--+ +--+ | | | +-- Hamiota_subangulata ** | | +--+ | | +-- Hamiota_australis__1 ** +--+ | +----- Hamiota_altilis__3 ** | | | | +-- Hamiota_perovalis__1 ** +--+ | | +-- Hamiota_perovalis__2 ** | | | +-- Hamiota_altilis__1 ** +--+ +-- Hamiota_altilis__2 ** | +-- Hamiota_altilis__4 ** | +-- Hamiota_altilis__5 **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 5 | 628 | 89 | 26 | 6 |
+-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
| | | +--+
+--+ | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +-- Hamiota_australis **
| | +--+
| | | +-- Hamiota_subangulata **
| | +-----------------------------------+
+--+ | | +-- Hamiota_altilis **
| | | |
| | +--+-- Hamiota_perovalis__1 **
| | |
| | +-- Hamiota_perovalis__2 **
| |
| | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
+--+ | | | | +-- Potamilus_capax
| | | | +--+
| | | | +-- Truncilla
| | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 52 | 44 | 22 | 6 |
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
| | +----------------- Leaunio
+--+ |
| | +----- Venustaconcha
| | +-----+
| | | | +-- Obovaria
| | | +--+
| | +--+ +-- Epioblasma
+--+ | |
| | | +-------- Lampsilis_siliquoidea
| | +--+
| | | +----- Ortmanniana_ligamentina
| | +--+
| | | +-- Cambarunio
| | +--+
+--+ +-- Ligumia
|
| +-- Hamiota_australis **
| +--+
| | +-- Hamiota_subangulata **
| +-----+
| | | +-- Hamiota_altilis **
| | +--+
| | +-- Hamiota_perovalis **
+--+
| +-- Lampsilis_fasciola
| +-----+
| | +-- Ortmanniana_pectorosa
+--+
| +----- Lampsilis_ornata
+--+
| +-- Lampsilis_ovata
+--+
+-- Lampsilis_cardium
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 6 | 8 | 98 | 39 | 22 | 5 |
+----- Obliquaria
+-----------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Sagittunio
| |
| +-------------- Leaunio
| |
| +-------------- Potamilus
| |
| | +-- Ellipsaria
| +-----------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +--------+
--+ | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +--------+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
| | +-------- Ortmanniana_pectorosa
| +-----+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_ovata
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+-----+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+-----+ +----- Medionidus_conradicus
| | |
| | | +-- Lemiox
| +--+--+
| | +-- Medionidus_acutissimus
| |
| | +-- Dromus
| +--+
| +-- Cyprogenia
|
| +----- Hamiota_australis__2 **
| +-----+
| | | +-- Hamiota_australis__1 **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| +-- Hamiota_altilis__3 **
| +-----+
| | +-- Hamiota_perovalis__2 **
+--+
| +----- Hamiota_altilis__2 **
+--+
| +-- Hamiota_altilis__1 **
+--+
+-- Hamiota_perovalis__1 **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 53 | 44 | 22 | 6 |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
| | | +-- Glebula
--+ | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
| | | | +-- Dromus
+--+ | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
| | +-- Lemiox
+--+
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
| | +-----------------------+
+--+ | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
| | +--------------+
+--+ | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
| |
| | +-- Ligumia
+--+ +--------+
| | +-- Cambarunio
| +--------+
| | | +-------- Lampsilis_teres
| | | |
| | +--+ +-- Hamiota_australis **
| | | +--+
| | | | +-- Hamiota_subangulata **
+--+ +--+
| | +-- Hamiota_altilis **
| +--+
| +-- Hamiota_perovalis **
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 5 | 8 | 98 | 39 | 22 | 5 |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
| | +-----+
+--+ | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
| | |
+--+ | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
| | | | +-------- Epioblasma
+--+ | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Ligumia
+--+ +--------+
| | +-- Cambarunio
| +--+
| | | +-------- Lampsilis_teres
| | +--+
| | | +----- Ortmanniana_ligamentina
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Leaunio
+--+
| +-- Lampsilis_cardium
| +--+
| | +-- Lampsilis_ovata
| +-----+
| | | +-- Ortmanniana_pectorosa
| | +--+
| | +-- Lampsilis_ornata
| |
+--+ +----- Hamiota_australis__2 **
| +--+
| | | +-- Hamiota_australis__1 **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| +----- Hamiota_altilis__2 **
| |
| | +-- Hamiota_altilis__1 **
+--+--+
| +-- Hamiota_perovalis__1 **
|
| +-- Hamiota_altilis__3 **
+--+
+-- Hamiota_perovalis__2 **
Chapman et al., 2008 [Publ 2667], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- |
+-------------------------- Toxolasma | --+ +----------------------- Obliquaria | | | | +-- Cyrtonaias +--+ +-----------------+ | | +-- Glebula | | +--+ +-- Ptychobranchus | +--------------+ | | +-- Lemiox | | +--+ +-- Venustaconcha | +-----------+ | | +-- Obovaria +--+ | +----------- Hamiota_subangulata ** +--+ | +-------- Lampsilis_ovata +--+ | +----- Ortmanniana +--+ | +-- Lampsilis_straminea +--+ +-- Lampsilis_hydiana
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- |
+----------------------- Obliquaria | | +-- Medionidus | +--+ | | +-- Lemiox --+ +--------------+ | | | +-- Cyprogenia | | +--+ | | +-- Ptychobranchus | | +--+ +-------- Potamilus | +--------+ | | | +----- Atlanticoncha | | +--+ | | | +-- Ellipsaria | | +--+ +--+ +-- Truncilla | | +-- Hamiota_subangulata ** | +-----------+ | | +-- Villosa | | +--+ +----------- Ligumia | | | | +----- Venustaconcha +--+ +--+ | | | +-- Obovaria | | +--+ +--+ +-- Epioblasma | | +----- Ortmanniana_pectorosa +--+ | +-- Ortmanniana_ligamentina +--+ +-- Lampsilis
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- |
+----------------------- Ellipsaria | --+ +-------------------- Truncilla | | +--+ +----------------- Obliquaria | | | | +-------------- Potamilus +--+ | | | +----- Lampsilis_ornata | | +-----+ | | | | +-- Lampsilis_cardium +--+ | +--+ | | +-- Lampsilis_ovata | | | | +-------- Lampsilis_teres | +--+ +--+ | +----- Lampsilis_siliquoidea | +--+ | | +-- Ortmanniana | +--+ | +-- Lampsilis_higginsii | | +----- Hamiota_altilis ** | | +-----+ +-- Epioblasma | | +--+-- Cambarunio | +-- Ligumia
Chapman et al., 2008 [Publ 2744], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- |
+-------------------------- Toxolasma | --+ +----------------------- Obliquaria | | | | +-- Cyrtonaias +--+ +-----------------+ | | +-- Glebula | | +--+ +-- Ptychobranchus | +--------------+ | | +-- Lemiox | | +--+ +-- Venustaconcha | +-----------+ | | +-- Obovaria +--+ | +----------- Hamiota_subangulata ** +--+ | +-------- Lampsilis_ovata +--+ | +----- Ortmanniana +--+ | +-- Lampsilis_straminea +--+ +-- Lampsilis_hydiana
