species Reginaia ebenus (Lea, 1831): synonymy

Amblema antrosa Rafinesque, 1820

Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 322.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 64.
(new name for Obovaria pachostea Rafinesque, 1820)

Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia antrosa — Haas (1969)
Amblema antrosa — Burch (1975)
Fusconaia ebena — Vidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), He & Zhuang (2013)

Obliquaria (Rotundaria) pusilla Rafinesque, 1820

Rafinesque, 1818. Amer. Mon. Mag. 3: 355 [as ‘Potamilus pusillus’]. [nomen nudum]
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 308 [also as ‘Unio pusilla’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23, 52.

Unio buxeus — Vanatta (1915)
Fusconaia pusilla — Morrison (1969)
Obliquaria pusilla — Burch (1975)
Fusconaia ebena — R.I. Johnson (1980), He & Zhuang (2013)

Obovaria obovalis Rafinesque, 1820

Rafinesque, 1818. Amer. Mon. Mag. 3: 355 [as ‘Potamilus obovalis’]. [nomen nudum]
Rafinesque, 1819. Jour. Phys. Chim. Hist. Nat. 88: 426 [as ‘U. (Obovaria) obovalis’]. [nomen nudum]
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 311 [also as ‘Unio obovalis’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23, 29, 54.
Type(s): LECTOTYPE ANSP_20244, Ohio River.

Quadrula obovalis — Vanatta (1915)
Fusconaia ebenus — Ortmann & Walker (1922)
Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia antrosa — Haas (1969)
Obovaria obovalis — Burch (1975)
Fusconaia ebena — Oesch (1984), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), He & Zhuang (2013)

Obovaria pachostea Rafinesque, 1820

Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 312 [also as ‘U. pachostea’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 55.
(renamed Amblema antrosa Rafinesque, 1820)

Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia antrosa — Haas (1969)
Plethobasus pachosteus — Morrison (1969)
Obovaria pachostea — Burch (1975)
Plethobasus cicatricosus — R.I. Johnson (1980)
Fusconaia ebena — Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), He & Zhuang (2013)

Unio ebenus Lea, 1831

Lea, 1831. Trans. Amer. Phil. Soc. 4 [O. 1]: 84 [94], pl. 9, fig. 14.
Type(s): HOLOTYPE USNM_85792, Cincinnati, Ohio.

Quadrula (sec. Fusconaia) ebenus — Simpson (1900), Simpson (1914)
Unio (s.s.) ebenus — Call (1900)
Fusconaia ebenus — Ortmann (1912), Ortmann & Walker (1922), Ortmann (1925), F.C. Baker (1928), Burch (1975), Mathiak (1979)
Unio ebenus, var. — de Gregorio (1914)
Fusconaia ebena — Utterback (1915), Gordon et al. (1979), R.I. Johnson (1980), Davis & Fuller (1981), Oesch (1984), ... Breton et al. (2011), ITIS (2012), IUCN (2012), McMurray et al. (2012), He & Zhuang (2013)
Quadrula obovalis — Vanatta (1915)
Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia antrosa — Haas (1969)
Fusconaia pusilla — Morrison (1969)
Reginaia ebenus — Campbell & Lydeard (2012), Campbell & Lydeard (2012), Inoue et al. (2013), Williams et al. (2017), Zieritz et al. (2018), ... Smith et al. (2020), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)
Reginaia ebena — Haag (2012), Pfeiffer et al. (2016), Chase et al. (2018), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Hauck et al. (2023)

Unio obovalis Say, 1834

Say, 1834. Amer. Conch. 6: [unpaginated].
(reference to Obovaria obovalis Rafinesque, 1820)

Quadrula (sec. Fusconaia) ebenus — Simpson (1900), Simpson (1914)
Fusconaia ebena — Watters et al. (2009), He & Zhuang (2013)

Unio obovata Conrad, 1834

‘Raf.’ Conrad, 1834. New Fresh-Water Shells: 69.
(error for Obovaria obovalis Rafinesque, 1820)

Pleurobema cordata — Ortmann & Walker (1922)

Unio obliqua ‘Lamarck’ of Authors

‘Lamarck’ of Authors.
Ferussac, 1835. Mag. Zool. 5(59-60): 28.
Conrad, 1837. Monogr. Fam. Unionidae 1(9): 77, pl. 43, fig. 2.
(misidentified reference to Unio obliqua Lamarck, 1819)

Quadrula (sec. Fusconaia) ebenus — Simpson (1900), Simpson (1914)
Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia antrosa — Haas (1969)
Fusconaia ebena — Parmalee & Bogan (1998), Watters et al. (2009)

Unio globatus Call, 1894

Call, 1894. Proc. Indiana Acad. Sci. 1893(3): ??.
(misidentified reference to Unio globatus Lea, 1871)

Quadrula (Fusconia [sic]) antrosa — Frierson (1927)

Unio subrotundus Call, 1894

‘Lea’ Call, 1894. Proc. Indiana Acad. Sci. 1893(3): ??.
(misidentified reference to Unio subrotundus Lea, 1831)

Quadrula (Fusconia [sic]) antrosa — Frierson (1927)

Unio pusella Lea, 1836

‘Rafinesque’ Lea, 1836. Synopsis Fam. Naiades: 59.
‘Rafinesque’ Lea, 1852. Synopsis Fam. Naiades: 58.
‘Rafinesque’ Lea, 1870. Synopsis Fam. Naiades: 92.
(error for Obliquaria (Rotundaria) pusilla Rafinesque, 1820)

Fusconaia ebena — He & Zhuang (2013)

Unio mytiliodes Swainson, 1840

Swainson, 1840. Treat. Malacol.: 292.
(error for Unio (s.s.) mytiloides Swainson, 1840)

Unio (s.s.) mytiloides Swainson, 1840

‘Rafinesque’ Swainson, 1840. Treat. Malacol.: 269-270, figs. 52-53; 292 [as ‘mytilïodes’].
(misidentified reference to Pleurobema mytiloides Rafinesque, 1820)

Quadrula (sec. Fusconaia) ebenus — Simpson (1900), Simpson (1914)
Quadrula (Fusconia [sic]) antrosa — Frierson (1927)
Fusconaia ebena — Parmalee & Bogan (1998), Watters et al. (2009)