species Ptychobranchus foremanianus (Lea, 1842): synonymy

Unio foremanianus Lea, 1842

Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 247 [85], pl. 27, fig. 64.
Lea, 1843. Proc. Amer. Phil. Soc. 2: 224.
LECTOTYPE [AS HOLOTYPE] USNM_84439 (by R.I. Johnson, 1974: 58), Coosa R., Alabama.

Ptychobranchus foremanianus — Simpson (1900), Ortmann (1912), Williams et al. (2008), Bogan (2010), Haag (2012), Williams et al. (2017), Graf & Cummings (2021), Bieler (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)
Ptychobranchus foremanianum — Simpson (1914)
Ptychobranchus (s.s.) foremanianum — Frierson (1927), Haas (1969)
Ptychobranchus greeni — Burch (1975), Parmalee & Bogan (1998)

Unio velatus Conrad, 1854

Conrad, 1853. Proc. Acad. Nat. Sci. 6: 259.
Conrad, 1854. Jour. Acad. Nat. Sci. 2(4): 298, pl. 27, fig. 6.

Ptychobranchus foremanianus — Simpson (1900), Williams et al. (2008)
Ptychobranchus foremanianum — Simpson (1914)
Ptychobranchus (s.s.) foremanianum — Frierson (1927), Haas (1969)
Ptychobranchus greeni — Burch (1975), Parmalee & Bogan (1998)

Unio woodwardius Lea, 1857

Lea, 1857. Proc. Acad. Nat. Sci. 9: 170.
Lea, 1859. Jour. Acad. Nat. Sci. 4 [O. 7]: 199, pl. 23, fig. 82; pl. 29, fig. 103 [as ‘woodwardianus’].

Ptychobranchus foremanianus — Simpson (1900), Williams et al. (2008), Bieler (2021)
Ptychobranchus foremanianum — Simpson (1914)
Ptychobranchus (s.s.) foremanianum — Frierson (1927), Haas (1969)
Ptychobranchus greeni — Burch (1975), Parmalee & Bogan (1998)

Unio woodwardianus ‘Lea’ of Authors

(error for Unio woodwardius Lea, 1857)
‘Lea’ of Authors.
Lea, 1859. Jour. Acad. Nat. Sci. 4 [O. 7]: 199.
Lea, 1870. Synopsis Fam. Naiades: 36.
B.H. Wright, 1888. Check List N. Amer. Unionidae etc.: [6].

Ptychobranchus foremanianus — Simpson (1900), Bieler (2021)
Ptychobranchus foremanianum — Simpson (1914)
Ptychobranchus (s.s.) foremanianum — Frierson (1927), Haas (1969)
Ptychobranchus greeni — Burch (1975), Parmalee & Bogan (1998)

Unio trinacrus Lea, 1861

Lea, 1861. Proc. Acad. Nat. Sci. 13: 59.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 8]: 86 [90], pl. 12, fig. 235.
HOLOTYPE USNM_84445, Coosa R., Alabama.

Ptychobranchus trinacrus — Simpson (1900)
Ptychobranchus trinacrium [sic] — Simpson (1914)
Ptychobranchus (s.s.) foremanianum — Frierson (1927), Haas (1969)
Ptychobranchus greeni — Burch (1975), Parmalee & Bogan (1998)
Ptychobranchus foremanianus — Williams et al. (2008)

Unio trinacris ‘Lea’ of Authors

(error for Unio trinacrus Lea, 1861)
‘Lea’ of Authors.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 8]: pl. 12, fig. 235.
Sowerby, 1868. Conch. Icon. 16: pl. 81, sp. 428.

Ptychobranchus trinacrus — Simpson (1900)

Ptychobranchus trinacrium Simpson, 1914

(error for Unio trinacrus Lea, 1861)
‘Lea’ Simpson, 1914. Descr. Cat. Naiades: 337.