Bivalvia | Unionida | Hyrioidea | Hyriidae | Velesunioninae

genus Velesunio Iredale, 1934: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 56 cladograms in 35 publications
maximum number of terminals in an analysis = 3 (mean = 1.23, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 11
maximum number of included terminals = 3, (mean = 2.18 median = 2)
proportion of cladograms that support basic monophyly = 0.09

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Velesunio cladogram distances
PCoA Ordination

NJ Tree of Velesunio cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 8 10 5 627 7 4 0 
              +-------- Velesunio__1 **         
  +-----------+                              
  |           |  +----- Alathyria_jacksoni__1
  |           +--+                           
  |              |  +-- Velesunio_ambiguus **   
--+              +--+                        
  |                 +-- Alathyria_profuga    
  |                                          
  |  +----------------- Velesunio__3 **         
  |  |                                       
  +--+  +-------------- Microdontia          
     |  |                                    
     +--+           +-- Lortiella            
        |  +--------+                        
        |  |        +-- Velesunio_angasi **     
        +--+                                 
           |  +-------- Alathyria_pertexta__2
           +--+                              
              |  +----- Velesunio__2 **         
              +--+                           
                 |  +-- Alathyria_jacksoni__2
                 +--+                        
                    +-- Alathyria_pertexta__1

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
3 5 4 5 33 2 2 6 
-- 2889

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
2 5 0 2 18 4 4 11.5 
  +----------- Velesunio_ambiguus **
--+                              
  |  +-------- Alathyria         
  +--+                           
     |  +----- Velesunio_angasi **  
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
2 3 0 5 7 1 1 8 
           +-- Velesunio__3 **    
  +--------+                   
  |        +-- Velesunio__4 **    
--+                            
  |  +-------- Velesunio__2 **    
  +--+                         
     |  +----- Velesunio_angasi **
     +--+                      
        |  +-- Velesunio__1 **    
        +--+                   
           +-- Alathyria

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
2 5 0 2 18 4 4 9.5 
  +----------- Velesunio_ambiguus **
--+                              
  |  +-------- Microdontia       
  +--+                           
     |  +----- Lortiella         
     +--+                        
        |  +-- Alathyria         
        +--+                     
           +-- Velesunio_angasi **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 3 0 2 45 1 1 9 
  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 6 0 0 -- 
  +-- Velesunio_angasi **  
--+                     
  +-- Velesunio_ambiguus **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 12 1 1 10.5 
  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio **         
  +--+                     
     +-- Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 44 1 1 9 
  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 44 1 1 9 
  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 45 1 1 9 
  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 6 4 -- 
  +----------- Westralunio       
  |                              
  |     +----- Alathyria_jacksoni
--+  +--+                        
  |  |  |  +-- Velesunio_ambiguus **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Microdontia       
        +--+                     
           +-- Lortiella

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Lortiella       
--+                   
  +-- Velesunio_angasi **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
-- Velesunio_angasi **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Velesunio_ambiguus **

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Velesunio_ambiguus **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Velesunio_ambiguus **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 3 -- 
     +----- Microdontia       
  +--+                        
  |  |  +-- Alathyria_pertexta
  |  +--+                     
--+     +-- Lortiella         
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Velesunio_ambiguus **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Velesunio_ambiguus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 0 0 -- 
-- Velesunio_ambiguus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 0 0 -- 
-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Velesunio_ambiguus **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 3 -- 
     +----- Microdontia       
  +--+                        
  |  |  +-- Lortiella         
  |  +--+                     
--+     +-- Alathyria_pertexta
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 2 1 -- 
  +-- Velesunio_ambiguus **
--+                     
  +-- Westralunio

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
-- Velesunio_angasi **

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 200 6 4 -- 
     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 198 6 4 -- 
     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 2 1 -- 
-- Velesunio_ambiguus **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Velesunio_ambiguus **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Velesunio_ambiguus **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Velesunio_ambiguus **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Velesunio_ambiguus **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Velesunio_ambiguus **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Velesunio_ambiguus **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Velesunio_ambiguus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 0 0 -- 
-- Velesunio_ambiguus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 138 5 3 -- 
     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Microdontia       
  +--+                        
     |  +-- Alathyria_pertexta
     +--+                     
        +-- Lortiella

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 138 5 3 -- 
     +----- Alathyria_pertexta
  +--+                        
  |  |  +-- Microdontia       
  |  +--+                     
--+     +-- Lortiella         
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Velesunio **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Velesunio_ambiguus **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Velesunio_ambiguus **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 200 6 4 -- 
     +----- Alathyria_jacksoni
  +--+                        
  |  |  +-- Alathyria_profuga 
  |  +--+                     
--+     +-- Velesunio_ambiguus **
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Velesunio_ambiguus **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Velesunio_ambiguus **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Velesunio_ambiguus **