Bivalvia | Unionida | Hyrioidea | Hyriidae | Velesunioninae

genus Velesunio Iredale, 1934: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 63 cladograms in 37 publications
maximum number of terminals in an analysis = 3 (mean = 1.27, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 13
maximum number of included terminals = 3, (mean = 2.31 median = 2)
proportion of cladograms that support basic monophyly = 0.08

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Velesunio cladogram distances
PCoA Ordination

NJ Tree of Velesunio cladogram distances
NJ Tree

Cladograms

Lawrie et al., 2026 [Publ 4843], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.667 3 9 14 3 15 8 4 **
Cladomics cladogram Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.500 3 8 10 3 630 7 4 11.0
Cladomics cladogram Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1

Lawrie et al., 2026 [Publ 4843], fig. 2

cst iT tT bi riS oT roT roS iD
0.600 3 9 13 3 6 6 3 11.0
Cladomics cladogram Velesunio 9 Velesunio ambiguus Alathyria jacksoni 4 Alathyria profuga Velesunio 5 Velesunio 8 Microdontia Lortiella Velesunio angasi Velesunio 6 Velesunio 7 Velesunio 4 Alathyria jacksoni 1 Alathyria jacksoni 2 Alathyria jacksoni 3 Alathyria pertexta 1 Velesunio 1 Velesunio 2 Velesunio 3 Alathyria pertexta 2

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.750 3 5 4 3 33 2 2 10.0
Cladomics cladogram Echyridella Velesunio ambiguus Velesunio 3 Velesunio 2 Alathyria Velesunio 1 Velesunio angasi Lortiella

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.667 2 5 0 2 19 4 4 15.5
Cladomics cladogram Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 2 5 0 2 19 4 4 13.5
Cladomics cladogram Cucumerunio Hyridella Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.667 2 3 4 2 7 1 1 12.0
Cladomics cladogram Echyridella Castalia Velesunio 3 Velesunio 4 Velesunio 2 Velesunio angasi Velesunio 1 Alathyria

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 45 1 1 13.0
Cladomics cladogram Velesunio ambiguus Velesunio angasi Lortiella Echyridella Hyridella Castalia Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 45 1 1 13.0
Cladomics cladogram Velesunio ambiguus Velesunio angasi Lortiella Echyridella Hyridella Castalia Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 44 1 1 13.0
Cladomics cladogram Velesunio ambiguus Velesunio angasi Lortiella Echyridella Hyridella Castalia Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 44 1 1 13.0
Cladomics cladogram Velesunio ambiguus Velesunio angasi Lortiella Echyridella Hyridella Castalia Diplodon

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 12 1 1 14.5
Cladomics cladogram Velesunio ambiguus Velesunio Lortiella Diplodon Cucumerunio Hyridella

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 6 0 0 --
Cladomics cladogram Velesunio angasi Velesunio ambiguus Echyridella Castalia

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 19 6 4 --
Cladomics cladogram Westralunio Alathyria jacksoni Velesunio ambiguus Alathyria profuga Alathyria pertexta Microdontia Lortiella

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 200 6 4 --
Cladomics cladogram Alathyria jacksoni Alathyria profuga Velesunio ambiguus Alathyria pertexta Lortiella Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 198 6 4 --
Cladomics cladogram Velesunio ambiguus Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 200 6 4 --
Cladomics cladogram Velesunio ambiguus Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 22 4 4 --
Cladomics cladogram Velesunio ambiguus Alathyria Lortiella Microdontia

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 29 5 3 --
Cladomics cladogram Microdontia Lortiella Alathyria pertexta Velesunio ambiguus Alathyria jacksoni Alathyria profuga

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 29 5 3 --
Cladomics cladogram Microdontia Alathyria pertexta Lortiella Velesunio ambiguus Alathyria jacksoni Alathyria profuga

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 138 5 3 --
Cladomics cladogram Alathyria pertexta Microdontia Lortiella Velesunio ambiguus Alathyria jacksoni Alathyria profuga

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 138 5 3 --
Cladomics cladogram Velesunio ambiguus Alathyria jacksoni Alathyria profuga Microdontia Alathyria pertexta Lortiella

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram Velesunio angasi Lortiella

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 30 1 1 --
Cladomics cladogram Castalia Velesunio angasi

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Lortiella Velesunio angasi

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Velesunio angasi Lortiella

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 30 1 1 --
Cladomics cladogram Castalia Velesunio angasi

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Velesunio angasi Lortiella

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 5 2 1 --
Cladomics cladogram Velesunio ambiguus Cucumerunio

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 1 --
Cladomics cladogram Velesunio ambiguus Westralunio

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Hyridella Velesunio

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Velesunio ambiguus Diplodon Castalia Hyridella australis 2 Hyridella australis 1 Echyridella Hyridella depressa

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella Diplodon Castalia

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella Castalia Triplodon

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella Castalia Triplodon

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Margaritifera Velesunio ambiguus Lamellidens Coelatura Acuticosta Scriptolamprotula Unio Nodularia Cristaria Sinanodonta Strophitus Platynaias Pyganodon Quadrula Eurynia Amblema Lampsilis Truncilla Sinohyriopsis Gonidea Pseudodon Sundadontina Physunio Pronodularia Potomida Lamprotula

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Velesunio ambiguus

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Margaritifera Aspatharia

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella Triplodon Castalia

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Velesunio ambiguus

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Velesunio ambiguus Aspatharia Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Velesunio ambiguus Unio Aspatharia Margaritifera

Elderkin et al., 2016 [Publ 3693], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Etheria Aspatharia Velesunio ambiguus

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram Velesunio ambiguus Triplodon Hyridella

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella Velesunio ambiguus

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Triplodon Hyridella Velesunio ambiguus

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Triplodon Velesunio ambiguus Hyridella

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Velesunio ambiguus Hyridella