MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Thu 19 March 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Truncilla Rafinesque, 1819: cladomics
4 Recent valid species (4 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 4 (4 total)
analyzed in 39 cladograms in 25 publications
maximum number of terminals in an analysis = 4 (mean = 1.62, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 10
maximum number of included terminals = 4, (mean = 3.4 median = 4)
proportion of cladograms that support basic monophyly = 1
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 627 | 89 | 26 | 0 |
+-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
| | | +--+
+--+ | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
| | | +----------------------- Pachynaias
+--+ | |
| | | +-------------------- Sagittunio
| | +--------------+ |
| | | | | +-------------- Ellipsaria
| | | | | +--+
| | | | | | | +----------- Potamilus_capax
| | | +--+ | +--+
| | | | | | +-------- Truncilla_macrodon **
| | | | | +--+
| | | | | | +----- Truncilla_truncata **
+--+ | | | +--+
| | +--+ | +-- Truncilla_cognata **
| | | +--+
| | | +-- Truncilla_donaciformis **
| | |
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
+--+ +--+ +-- Cyprogenia_aberti__2
| | +-----+
| | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Smith et al., 2020 [Publ 4129], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 24 | 16 | 7 | 4 |
+----------------------- Obliquaria | --+ +-------------------- Cyrtonaias | | | | +-- Pachynaias +--+ +--------------+ | | +-- Lampsilis | | +--+ +-- Atlanticoncha | +-----------+ | | +-- Potamilus +--+ | +----------- Ellipsaria +--+ | +-------- Truncilla_truncata ** +--+ | +----- Truncilla_donaciformis ** +--+ | +-- Truncilla_cognata ** +--+ +-- Truncilla_macrodon **
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 4 | 29 | 22 | 17 | 2 |
+----------------- Toxolasma | | +-------------- Potamilus | | | | +----- Medionidus | +--------+ | | | +-- Lemiox --+ | +--+ | | +-- Dromus | | | | +----- Ellipsaria | | | | +--------+ +-- Truncilla_cognata ** | | | | | | | +-- Truncilla_donaciformis ** +--+ +--+ | +-- Truncilla_macrodon ** | | | +-- Truncilla_truncata ** | | +----------- Cambarunio | | | +----------- Leaunio | | | +----------- Villosa | | | | +-- Sagittunio | +--------+ +--+ +-- Lampsilis_ovata | | +----- Venustaconcha +-----+ | | +-- Epioblasma | +--+ | +-- Obovaria | | +-------- Lampsilis_teres +--+ | +----- Lampsilis_siliquoidea +--+ | +-- Lampsilis_hydiana +--+ +-- Lampsilis_straminea
Saxon et al., 2023 [Publ 4338], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 1 | 1 | 1 | 2 |
+----------- Ellipsaria --+ | +-------- Truncilla_truncata ** +--+ | +----- Truncilla_macrodon ** +--+ | +-- Truncilla_cognata ** +--+ +-- Truncilla_donaciformis **
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 37 | 32 | 19 | 4 |
+-------------------------- Obliquaria | | +-- Medionidus | +--+ | | +-- Lemiox | +-----------------+ --+ | | +-- Cyprogenia | | +--+ | | +-- Ptychobranchus | | | | +----------------- Potamilus | | +--+ +--+ | | +-------------- Atlanticoncha | | +--+ | | | +----------- Ellipsaria | | +--+ | | | +-------- Truncilla_truncata ** | | +--+ | | | +----- Truncilla_cognata ** +--+ +--+ | | +-- Truncilla_macrodon ** | +--+ | +-- Truncilla_donaciformis ** | | +-- Hamiota | +-----------+ | | +-- Villosa | | +-----+ +----------- Ligumia | | | | +----- Venustaconcha +--+ +--+ | | | +-- Obovaria | | +--+ +--+ +-- Epioblasma | | +----- Ortmanniana_pectorosa +--+ | +-- Ortmanniana_ligamentina +--+ +-- Lampsilis
Smith et al., 2020 [Publ 4129], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 24 | 16 | 7 | 4 |
+----------------------- Obliquaria | --+ +-------------------- Cyrtonaias | | | | +-- Pachynaias +--+ +--------------+ | | +-- Lampsilis | | +--+ +-- Atlanticoncha | +-----------+ | | +-- Potamilus +--+ | +----------- Ellipsaria +--+ | +-------- Truncilla_truncata ** +--+ | +----- Truncilla_donaciformis ** +--+ | +-- Truncilla_cognata ** +--+ +-- Truncilla_macrodon **
Inoue et al., 2020 [Publ 3941], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 38 | 23 | 6 | 3 |
+----------------------------------- Toxolasma | --+ +-------------------------------- Ellipsaria | | | | +----- Truncilla_cognata ** +--+ +-----------------------+ | | | +-- Truncilla_macrodon ** | | +--+ | | +-- Truncilla_donaciformis ** +--+ | +-------------------------- Potamilus | | | | +----------------------- Leaunio | | | +--+ | +----------------- Lampsilis_bracteata | | | | | +--+ +-- Lampsilis_ornata | | | | +-----------+ +--+ | | | +-- Lampsilis_fasciola | | +--+ | | | +----------- Lampsilis_ovata | | +--+ | | | +-------- Lampsilis_cardium__3 | | +--+ +--+ | +----- Lampsilis_cardium__2 | +--+ | | +-- Lampsilis_satura | +--+ | +-- Lampsilis_cardium__1 | | +-------------- Lampsilis_teres | | | | +-- Lampsilis_higginsii +-----+ +--------+ | | +-- Ortmanniana | | +--+ +-- Lampsilis_radiata | +-----+ | | +-- Lampsilis_siliquoidea | | +--+ +-- Lampsilis_hydiana | +--+ | | +-- Lampsilis_bergmanni +--+ | +-- Lampsilis_virescens +--+ +-- Lampsilis_straminea
Inoue et al., 2020 [Publ 3941], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 39 | 24 | 6 | 1 |
+----------------------------------- Toxolasma | | +-------------------------------- Potamilus --+ | | | +-------- Ellipsaria | | +--------------------+ +--+ | | +----- Truncilla_macrodon ** | | +--+ | | | +-- Truncilla_donaciformis ** | | +--+ | | +-- Truncilla_cognata ** +--+ | +-------------------------- Leaunio | | | | +-------------------- Lampsilis_bracteata | | | | | +--+ +----------------- Lampsilis_fasciola | | | | | +--+ | +--+ +-------------- Lampsilis_ornata | | | | | | +--+ +-- Lampsilis_ovata | | | +--------+ | | | | +-- Lampsilis_cariosa | | +--+ | | | +-------- Lampsilis_cardium__3 +--+ +--+ | | +----- Lampsilis_cardium__2 | +--+ | | +-- Lampsilis_satura | +--+ | +-- Lampsilis_cardium__1 | | +-- Lampsilis_higginsii | +--------------+ | | +-- Ortmanniana +-----+ | +-------------- Lampsilis_teres | | +--+ +-- Lampsilis_radiata | +--------+ | | +-- Lampsilis_siliquoidea +--+ | +-------- Lampsilis_bergmanni +--+ | +----- Lampsilis_hydiana +--+ | +-- Lampsilis_virescens +--+ +-- Lampsilis_straminea
Boyer et al., 2011 [Publ 2718], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 33 | 14 | 9 | -- |
+-------------- Toxolasma | | +----------- Obliquaria --+ | | | +-------- Obovaria | | | | | +-------- Lampsilis_cardium +--+ | | +-------- Potamilus | | | | +-- Epioblasma | +-----+ | | +-- Lampsilis_siliquoidea +--+ | +-- Ortmanniana +-----+ | +-- Lampsilis_higginsii | | +-- Ligumia +-----+ | +-- Lampsilis_teres | | +----- Ellipsaria +--+ | +-- Truncilla_truncata ** +--+ +-- Truncilla_donaciformis **
Boyer et al., 2011 [Publ 2718], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 36 | 14 | 9 | -- |
+----------------- Potamilus | | +----- Ellipsaria --+ +--------+ | | | +-- Truncilla_donaciformis ** | | +--+ | | +-- Truncilla_truncata ** +--+ | +----------- Lampsilis_cardium | | | | +----- Ligumia | +-----+ +--+ | +-- Obovaria | +--+ | +-- Epioblasma | | +-------- Lampsilis_teres +--+ | +----- Lampsilis_siliquoidea +--+ | +-- Ortmanniana +--+ +-- Lampsilis_higginsii
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 5 | 5 | -- |
+-- Cyrtonaias
+-----+
| +-- Glebula
|
--+ +-- Lampsilis
| +--+
| | +-- Leaunio
+--+
| +-- Truncilla_macrodon **
+--+
+-- Potamilus
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla_truncata **
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Wu et al., 2023 [Publ 4339], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 18 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Truncilla_truncata ** +--+ | +-- Lampsilis +--+ +-- Potamilus
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- |
+----------------------- Ellipsaria | --+ +-------------------- Truncilla_truncata ** | | +--+ +----------------- Obliquaria | | | | +-------------- Potamilus +--+ | | | +----- Lampsilis_ornata | | +-----+ | | | | +-- Lampsilis_cardium +--+ | +--+ | | +-- Lampsilis_ovata | | | | +-------- Lampsilis_teres | +--+ +--+ | +----- Lampsilis_siliquoidea | +--+ | | +-- Ortmanniana | +--+ | +-- Lampsilis_higginsii | | +----- Hamiota | | +-----+ +-- Epioblasma | | +--+-- Cambarunio | +-- Ligumia
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- |
+----- Ellipsaria | +----- Potamilus_leptodon | +----- Medionidus | +----- Potamilus_alatus | +----- Ptychobranchus --+ +----- Truncilla_truncata ** | +----- Venustaconcha | +----- Cambarunio | +----- Leaunio | | +-- Ortmanniana +--+ +-- Lampsilis
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Truncilla_truncata ** --+ +-- Lampsilis
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 39 | 2 | 2 | -- |
+----- Potamilus --+ | +-- Lampsilis +--+ +-- Truncilla_truncata **
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 197 | 13 | 11 | -- |
+-------------------- Potamilus_leptodon | --+ +----------------- Potamilus_alatus | | | | +-- Medionidus +--+ +-----------+ | | +-- Ptychobranchus | | +--+ +-- Ellipsaria | +--------+ | | +-- Truncilla_truncata ** +--+ | +-------- Leaunio | | +--+ +-- Venustaconcha | +--+ | | +-- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- |
+-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla_truncata **
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla_truncata **
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- |
+----------------- Ptychobranchus --+ | +-------------- Truncilla_truncata ** +--+ | +----------- Ortmanniana +--+ | +-------- Lampsilis +--+ | +----- Cambarunio +--+ | +-- Ligumia +--+ +-- Epioblasma
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- |
+----- Truncilla_truncata ** | +----- Ptychobranchus --+ | +-- Ortmanniana | | | +-- Cambarunio +--+ +-- Ligumia | +-- Epioblasma
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- |
+-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla_truncata **
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- |
+-------------------- Truncilla_truncata ** | --+ +----------------- Ligumia | | | | +-- Ptychobranchus +--+ +-----------+ | | +-- Sagittunio | | +--+ +----------- Lampsilis_siliquoidea | | | | +----- Ortmanniana +--+ +--+ | | | +-- Lampsilis_cardium | | +--+ +--+ +-- Lampsilis_fasciola | | +----- Epioblasma +--+ | +-- Cambarunio +--+ +-- Leaunio
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla_truncata **
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Breton et al., 2011 [Publ 3376], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Ellipsaria | | +--+ +----------- Truncilla_macrodon ** | | +--+ +-- Lemiox | +-----+ | | +-- Potamilus +--+ | +----- Lampsilis +--+ | +-- Cambarunio +--+ +-- Venustaconcha
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- |
+----------- Ligumia | +----------- Truncilla_truncata ** | | +-- Ptychobranchus +--------+ --+ +-- Sagittunio | | +-- Lampsilis_siliquoidea | +-----+ | | +-- Epioblasma | | +--+ +----- Ortmanniana | | | +----- Cambarunio +--+ +----- Leaunio | | +-- Lampsilis_cardium +--+ +-- Lampsilis_fasciola
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- |
+----------------- Ptychobranchus --+ | +-------------- Truncilla_truncata ** +--+ | +----------- Ortmanniana +--+ | +-------- Lampsilis +--+ | +----- Cambarunio +--+ | +-- Ligumia +--+ +-- Epioblasma
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Potamilus +--+ | +-- Lampsilis +--+ +-- Truncilla_truncata **
Chase et al., 2018 [Publ 3847], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 38 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Venustaconcha | | +--+ +-- Lemiox | +--------+ | | +-- Lampsilis +--+ | +-------- Potamilus +--+ | +----- Cambarunio +--+ | +-- Ellipsaria +--+ +-- Truncilla_macrodon **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Lampsilis --+ +-- Truncilla_truncata **
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- |
+-------------- Potamilus_leptodon | | +----------- Potamilus_alatus --+ | | | +-------- Venustaconcha | | | +--+ +-------- Leaunio | | | | +-- Ellipsaria | +-----+ +--+ +-- Truncilla_truncata ** | | +-- Medionidus +-----+ | +-- Ptychobranchus | | +----- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- |
+-------------- Obliquaria | +-------------- Truncilla_truncata ** --+ +-------------- Ptychobranchus | | +----------- Ligumia +--+ | +-------- Cambarunio +--+ | +----- Epioblasma +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Potamilus +--+ | +-- Lampsilis +--+ +-- Truncilla_truncata **
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Lampsilis --+ +-- Truncilla_truncata **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- |
+----------- Truncilla_truncata ** | --+ +-- Lampsilis | +-----+ | | +-- Epioblasma +--+ | +----- Ligumia +--+ | +-- Obliquaria +--+ +-- Ortmanniana
Inoue et al., 2020 [Publ 3941], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 15 | 1 | -- |
+-- Truncilla_macrodon ** --+ +-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- |
+----------------------- Cyrtonaias | --+ +-------------------- Ptychobranchus | | +--+ +----------------- Ligumia | | +--+ +-------------- Potamilus | | +--+ +----------- Pachynaias | | | | +-- Epioblasma +--+ +-----+ | | +-- Truncilla_truncata ** | | +--+ +-- Mesonaias | +--+ | | +-- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- |
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla_truncata **
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
