The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Ptychobranchus Simpson, 1900: cladomics
6 Recent valid species (8 total species) Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently
asked rhetorical questions) page. Abbreviations for the various statistics are explained on
the FARQ! Cladomics Summarynumber of species = 6 (8 total)
analyzed in 31 cladograms in 18 publications
maximum number of terminals in an analysis =
5 (mean = 1.13, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 1
maximum number of included terminals = 5, (mean =
5 median = 5)
proportion of cladograms that support basic monophyly
= 1
Roe, 2013 [Publ 3571], fig. 4 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 6 | 6 | 5 | 0 | +----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Medionidus
| |
+--+ +-- Cyprogenia
| +--------------+
| | +-- Dromus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Ptychobranchus_jonesi **
+--+
| +-------- Ptychobranchus_greenii **
+--+
| +----- Ptychobranchus_occidentalis__2 **
+--+
| +-- Ptychobranchus_occidentalis__1 **
+--+
+-- Ptychobranchus_fasciolaris **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- | +-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus_fasciolaris **
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +-- Villosa
|
+-- Ellipsaria
|
--+-- Lampsilis
|
+-- Potamilus
|
+-- Ptychobranchus_subtentus **
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- | +-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus_fasciolaris **
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus_fasciolaris **
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 13 | 11 | -- | +-------------- Potamilus_leptodon
|
| +----------- Potamilus_alatus
--+ |
| | +-------- Venustaconcha
| | |
+--+ +-------- Leaunio
| |
| | +-- Ellipsaria
| +-----+
+--+ +-- Truncilla
|
| +-- Medionidus
+-----+
| +-- Ptychobranchus_subtentus **
|
| +----- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 10 | 7 | -- | +----- Ptychobranchus_subtentus **
+--------+
| | +-- Potamilus_fragilis
| +--+
| +-- Medionidus
--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 8 | 6 | -- | +-------------- Potamilus_fragilis
|
--+ +----------- Ptychobranchus_subtentus **
| |
+--+ +-------- Medionidus
| |
+--+ +----- Potamilus_purpuratus
| |
+--+----- Lampsilis
|
| +-- Ellipsaria
+--+
+-- Villosa
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +-------------- Obliquaria
|
+-------------- Truncilla
--+
+-------------- Ptychobranchus_fasciolaris **
|
| +----------- Ligumia
+--+
| +-------- Cambarunio
+--+
| +----- Epioblasma
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Serb, 2006 [Publ 2678], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 13 | 11 | -- | +----- Ellipsaria
|
+----- Potamilus_leptodon
|
+----- Medionidus
|
+----- Potamilus_alatus
|
+----- Ptychobranchus_subtentus **
--+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- | +----------------------- Cyrtonaias
|
--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +----------------- Ligumia
| |
+--+ +-------------- Potamilus
| |
+--+ +----------- Pachynaias
| |
| | +-- Epioblasma
+--+ +-----+
| | +-- Truncilla
| |
+--+ +-- Mesonaias
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- | +-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- | +----------- Ligumia
|
+----------- Truncilla
|
| +-- Ptychobranchus_fasciolaris **
+--------+
--+ +-- Sagittunio
|
| +-- Lampsilis_siliquoidea
| +-----+
| | +-- Epioblasma
| |
+--+ +----- Ortmanniana
| |
| +----- Cambarunio
+--+
+----- Leaunio
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +----------------- Medionidus
--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus_subtentus **
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis
+--+
| +-- Villosa
+--+
+-- Ellipsaria
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 8 | 6 | -- | +-------------------- Obliquaria
--+
| +----------------- Medionidus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Potamilus_fragilis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ellipsaria
+--+
| +-- Villosa
+--+
+-- Lampsilis
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- | +-- Cambarunio
--+
+-- Ptychobranchus_fasciolaris **
Serb, 2006 [Publ 2678], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------- Obliquaria
|
| +-- Ptychobranchus_fasciolaris **
+-----------+
--+ +-- Lampsilis
|
| +-- Cyprogenia_aberti__6
| +--------+
| | +-- Dromus
+--+
| +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_aberti__4
| |
| | +-- Cyprogenia_stegaria__1
+--+ |
| +-- Cyprogenia_stegaria__2
| |
+--+-- Cyprogenia_aberti__1
|
+-- Cyprogenia_aberti__2
|
+-- Cyprogenia_aberti__3
Chapman et al., 2008 [Publ 2744], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- | +----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Serb & Barnhart, 2008 [Publ 2674], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__5
| |
+--+ +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- | +-------------------- Truncilla
|
--+ +----------------- Ligumia
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----------+
| | +-- Sagittunio
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_fasciola
|
| +----- Epioblasma
+--+
| +-- Cambarunio
+--+
+-- Leaunio
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- | +----- Truncilla
|
+----- Ptychobranchus_fasciolaris **
--+
| +-- Ortmanniana
| |
| +-- Cambarunio
+--+
+-- Ligumia
|
+-- Epioblasma
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- | +-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus_fasciolaris **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Chapman et al., 2008 [Publ 2667], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 196 | 13 | 11 | -- | +-------------------- Potamilus_leptodon
|
--+ +----------------- Potamilus_alatus
| |
| | +-- Medionidus
+--+ +-----------+
| | +-- Ptychobranchus_subtentus **
| |
+--+ +-- Ellipsaria
| +--------+
| | +-- Truncilla
+--+
| +-------- Leaunio
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Serb & Barnhart, 2008 [Publ 2674], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------------- Lampsilis
|
--+ +----------------------- Obliquaria
| |
+--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__4
| |
+--+ +----------- Cyprogenia_aberti__3
| |
+--+ +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Serb, 2006 [Publ 2678], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
|
