species Leguminaia wheatleyi (Lea, 1862): synonymy

Monocondyloea wheatleyi Lea, 1862

Lea, 1862. Proc. Acad. Nat. Sci. 14: 176 [as ‘Monocondylœa wheatleyi’].
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 399 [34], pl. 50, fig. 307.
Type(s): HOLOTYPE USNM_86331, R. Tigris, Syria.

Leguminaia (s.s.) wheatleyi — Simpson (1900), Germain (1911), Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Falkner (1994), Graf (2007), ... Dai et al. (2024), Bolotov et al. (2024), Jeratthitikul et al. (2025), Dai et al. (2025), Chen et al. (2025)
Leguminaia whaetleyi [sic] — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Monocondyloea mardinensis Lea, 1865

Lea, 1865. Proc. Acad. Nat. Sci. 16: 286 [as ‘Monocondylœa mardinensis’].
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 252, pl. 30, fig. 67 [as ‘Monocondylœa mardinensis’].
Type(s): FIGURED SYNTYPE USNM_86354, near Mardin, R. Tigris, Syria.

Leguminaia (s.s.) mardinensis — Simpson (1900), Germain (1911), Simpson (1914), Haas (1969)
Leguminaia mardinensis — Kobelt (1904), Thiele (1934), Pallary (1939)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Graf & Cummings (2007), Graf & Cummings (2021), Bogan et al. (2021), Jihad (2023)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia bourguignati Locard, 1883

Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 252, pl. 19 bis, figs. 11-13.

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900), Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia bourguignati — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia chantrei Locard, 1883

Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 252, pl. 19 bis, figs. 8-10.

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900), Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia chantrei — Çek & ŞerefliŞan (2010)

Leguminaia wheatleyi var. euphratica Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 19: 17, pl. 520, fig. 2705.
(intro'd as intra-sp. of Monocondyloea wheatleyi Lea, 1862)

Leguminaia (s.s.) wheatleyi — Simpson (1914)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia graeteri Kobelt, 1913

Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 46, pl. 533, figs. 2744-2746.
Type(s): TYPE SMF_4354, A.d. Kara-su, Zufl. des Sees v. Antiochia, Syrien.

Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia graeteri — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia nisibina Kobelt, 1913

Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 17, pl. 520, fig. 2706.

Leguminaia nisibana [sic] — Simpson (1914)
Leguminaia mardinensis — Pallary (1939)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia (mardinensis var.) naegelei Kobelt, 1913

Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 7, pl. 515, fig. 2693.
(intro'd as intra-sp. of Monocondyloea mardinensis Lea, 1865)
Type(s): TYPE SMF_4356, A.d. Tigris b. Mosul.

Leguminaia (mardinensis var.) naegelei — Kobelt (1913)
Leguminaia (s.s.) mardinensis — Simpson (1914)
Leguminaia mardinensis — Pallary (1939)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Microdondylaea haasii Kobelt, 1914

Kobelt, 1914. Icon. Land- und Süssw.-Moll. 20: 30, pl. 554, figs. 2815-2820.
Type(s): TYPE SMF_4454, Euphrat Zufl. b. Biredschik [Euphrates River, Birecik].

Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Jihad (2023)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia mardinensis var. truncata Pallary, 1939

Pallary, 1939. Mém. Inst. Égypte 39: 115.
(intro'd as intra-sp. of Monocondyloea mardinensis Lea, 1865)

Leguminaia mardinensis var. truncata — Pallary (1939)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia mardiniensis Bourguignat, 1876

Bourguignat, 1876. Bull. Soc. Sci. Phys. Nat. Toulouse 3: 98.
Bourguignat, 1880. Mater. Moll. Acécphales Syst. Europ. 1: 3.
(error for Monocondyloea mardinensis Lea, 1865)

Leguminaia vheatleyi Locard, 1883

‘Lea’ Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 253.
(error for Monocondyloea wheatleyi Lea, 1862)

Monocondylaea chantreyi Paetel, 1890

‘Brgt.’ Paetel, 1890. Cat. Conch.-Samml. 3: 174.
(error for Leguminaia chantrei Locard, 1883)

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900), Simpson (1914)

Leguminaia nisibana Simpson, 1914

‘Kobelt’ Simpson, 1914. Descr. Cat. Naiades: 452.
(error for Leguminaia nisibina Kobelt, 1913)

Potomida littoralis homsensis Schütt, 1983

‘Lea’ Schütt, 1983. Arch. Moll. 113(1/6): pl. 7 fig. 98. [in part]
(misidentified reference to Unio homsensis Lea, 1865)

Leguminaia whaetleyi Çek & ŞerefliŞan, 2010

‘Lea’ Çek & ŞerefliŞan, 2010. J. Experimental Zool. 315: 30.
(error for Monocondyloea wheatleyi Lea, 1862)

Gonidea angulata Jawad, 2021

Jawad, 2021. Southern Iraq’s Marshes 36: ??.
(misidentified reference to Anodonta angulata Lea, 1838)

Leguminaia wheatleyi — Bolotov et al. (2024)