species Leguminaia wheatleyi (Lea, 1862): synonymy

Monocondyloea wheatleyi Lea, 1862

Lea, 1862. Proc. Acad. Nat. Sci. 14: 176 [as ‘Monocondylœa wheatleyi’].
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 399 [34], pl. 50, fig. 307.
HOLOTYPE USNM_86331, R. Tigris, Syria.

Leguminaia (s.s.) wheatleyi — Simpson (1900), Germain (1911)
Leguminaia (s.s.) (Group of L. mardinensis) wheatleyi — Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Falkner (1994), Graf (2007), ... Jeratthitikul et al. (2025), Dai et al. (2025), Chen et al. (2025), Campbell & Lydeard (2026), Nahok et al. (2026)
Leguminaia whaetleyi [sic] — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Monocondyloea mardinensis Lea, 1865

Lea, 1865. Proc. Acad. Nat. Sci. 16: 286 [as ‘Monocondylœa mardinensis’].
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 252, pl. 30, fig. 67 [as ‘Monocondylœa mardinensis’].
FIGURED SYNTYPE USNM_86354, near Mardin, R. Tigris, Syria.

Leguminaia (s.s.) mardinensis — Simpson (1900), Germain (1911), Haas (1969)
Leguminaia mardinensis — Kobelt (1904), Thiele (1934), Pallary (1939)
Leguminaia (s.s.) (Group of L. mardinensis) mardinensis — Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Graf & Cummings (2007), Graf & Cummings (2021), Bogan et al. (2021), Jihad (2023)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia bourguignati Locard, 1883

Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 252, pl. 19 bis, figs. 11-13.

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900)
Leguminaia (s.s.) (Group of L. mardinensis) mardinensis var. chantrei — Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia bourguignati — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia chantrei Locard, 1883

Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 252, pl. 19 bis, figs. 8-10.

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900)
Leguminaia (s.s.) (Group of L. mardinensis) mardinensis var. chantrei — Simpson (1914)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia chantrei — Çek & ŞerefliŞan (2010)

Leguminaia wheatleyi var. euphratica Kobelt, 1912

(intro'd as intra-sp. of Monocondyloea wheatleyi Lea, 1862)
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 19: 17, pl. 520, fig. 2705.

Leguminaia (s.s.) (Group of L. mardinensis) wheatleyi — Simpson (1914)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia graeteri Kobelt, 1913

Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 46, pl. 533, figs. 2744-2746.
TYPE SMF_4354, A.d. Kara-su, Zufl. des Sees v. Antiochia, Syrien.

Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia graeteri — Çek & ŞerefliŞan (2010)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia nisibina Kobelt, 1913

Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 17, pl. 520, fig. 2706.

Leguminaia mardinensis — Pallary (1939)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia (mardinensis var.) naegelei Kobelt, 1913

(intro'd as intra-sp. of Monocondyloea mardinensis Lea, 1865)
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 7, pl. 515, fig. 2693.
TYPE SMF_4356, A.d. Tigris b. Mosul.

Leguminaia (mardinensis var.) naegelei — Kobelt (1913)
Leguminaia (s.s.) (Group of L. mardinensis) mardinensis — Simpson (1914)
Leguminaia mardinensis — Pallary (1939)
Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Microdondylaea haasii Kobelt, 1914

Kobelt, 1914. Icon. Land- und Süssw.-Moll. 20: 30, pl. 554, figs. 2815-2820.
TYPE SMF_4454, Euphrat Zufl. b. Biredschik [Euphrates River, Birecik].

Leguminaia wheatleyi — Haas (1940), Haas (1969), Schütt (1983), Jihad (2023)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia mardinensis var. truncata Pallary, 1939

(intro'd as intra-sp. of Monocondyloea mardinensis Lea, 1865)
Pallary, 1939. Mém. Inst. Égypte 39: 115.

Leguminaia mardinensis var. truncata — Pallary (1939)
Leguminaia cf. wheatleyi — Lopes-Lima et al. (2021)

Leguminaia mardiniensis Bourguignat, 1877

(error for Monocondyloea mardinensis Lea, 1865)
Bourguignat, 1877. Bull. Soc. Sci. Phys. Nat. Toulouse 3: 98.
Bourguignat, 1880. Mater. Moll. Acécphales Syst. Europ. 1: 3.

Leguminaia vheatleyi Locard, 1883

(error for Monocondyloea wheatleyi Lea, 1862)
‘Lea’ Locard, 1883. Arch. Mus. Hist. Nat. Lyon 3: 253.

Monocondylaea chantreyi Paetel, 1890

(error for Leguminaia chantrei Locard, 1883)
‘Brgt.’ Paetel, 1890. Cat. Conch.-Samml. 3: 174.

Leguminaia (s.s.) mardinensis var. chantrei — Simpson (1900)
Leguminaia (s.s.) (Group of L. mardinensis) mardinensis var. chantrei — Simpson (1914)

Leguminaia nisibana Simpson, 1914

(error for Leguminaia nisibina Kobelt, 1913)
‘Kobelt’ Simpson, 1914. Descr. Cat. Naiades: 452.

Potomida littoralis homsensis Schütt, 1983

(misidentified reference to Unio homsensis Lea, 1865)
‘Lea’ Schütt, 1983. Arch. Moll. 113(1/6): pl. 7 fig. 98. [in part]

Leguminaia whaetleyi Çek & ŞerefliŞan, 2010

(error for Monocondyloea wheatleyi Lea, 1862)
‘Lea’ Çek & ŞerefliŞan, 2010. J. Experimental Zool. 315: 30.

Gonidea angulata Jawad, 2021

(misidentified reference to Anodonta angulata Lea, 1838)
Jawad, 2021. Southern Iraq’s Marshes 36: ??.

Leguminaia wheatleyi — Bolotov et al. (2024)