MUSSELpdb
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Data last updated
Mon 2 March 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Cyrtonaias Crosse & Fischer, 1894: cladomics
1 Recent valid species (1 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 1 (1 total)
analyzed in 40 cladograms in 22 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)
Cladograms
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- |
+-- Cyrtonaias_berlandierii **
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Campbell et al., 2008 [Publ 2675], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- |
-- Cyrtonaias_berlandierii **
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias_berlandierii **
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- |
+-- Cyrtonaias_berlandierii **
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- |
+-- Cyrtonaias_berlandierii ** --+ +-- Glebula
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- |
+-- Glebula --+ +-- Cyrtonaias_berlandierii **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
-- Cyrtonaias_berlandierii **
Freeman et al., 2025 [Publ 4443], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 11 | 3 | 3 | -- |
+-------- Toxolasma --+ | +----- Obliquaria +--+ | +-- Glebula +--+ +-- Cyrtonaias_berlandierii **
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- |
+----------- Cyrtonaias_berlandierii ** --+ | +-------- Ortmanniana +--+ | +----- Lampsilis +--+ | +-- Potamilus +--+ +-- Ligumia
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+-- Cyrtonaias_berlandierii **
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Ortmanniana
Freeman et al., 2025 [Publ 4443], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 11 | 3 | 3 | -- |
+-- Glebula --+ +-- Cyrtonaias_berlandierii **
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 4 | 4 | -- |
-- Cyrtonaias_berlandierii **
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- |
+----------------------- Cyrtonaias_berlandierii ** | --+ +-------------------- Ptychobranchus | | +--+ +----------------- Ligumia | | +--+ +-------------- Potamilus | | +--+ +----------- Pachynaias | | | | +-- Epioblasma +--+ +-----+ | | +-- Truncilla | | +--+ +-- Mesonaias | +--+ | | +-- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 5 | 5 | -- |
+-- Cyrtonaias_berlandierii **
+-----+
| +-- Glebula
|
--+ +-- Lampsilis
| +--+
| | +-- Leaunio
+--+
| +-- Truncilla
+--+
+-- Potamilus
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias_berlandierii **
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Walker et al., 2006 [Publ 2666], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 13 | 4 | 4 | -- |
+----------- Potamilus --+ | +-------- Cyrtonaias_berlandierii ** +--+ | +----- Ligumia +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Ouyang et al., 2015 [Publ 4493], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- |
-- Cyrtonaias_berlandierii **
Campbell et al., 2008 [Publ 2675], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- |
-- Cyrtonaias_berlandierii **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- |
-- Cyrtonaias_berlandierii **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- |
+----------- Potamilus --+ | +-------- Cyrtonaias_berlandierii ** +--+ | +----- Ligumia +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Chapman et al., 2008 [Publ 2744], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- |
+-------------------------- Toxolasma | --+ +----------------------- Obliquaria | | | | +-- Cyrtonaias_berlandierii ** +--+ +-----------------+ | | +-- Glebula | | +--+ +-- Ptychobranchus | +--------------+ | | +-- Lemiox | | +--+ +-- Venustaconcha | +-----------+ | | +-- Obovaria +--+ | +----------- Hamiota +--+ | +-------- Lampsilis_ovata +--+ | +----- Ortmanniana +--+ | +-- Lampsilis_straminea +--+ +-- Lampsilis_hydiana
Smith et al., 2020 [Publ 4129], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- |
+-------------- Obliquaria | --+ +----------- Cyrtonaias_berlandierii ** | | | | +-- Pachynaias +--+ +-----+ | | +-- Lampsilis | | +--+ +-- Ellipsaria | +--+ | | +-- Truncilla +--+ | +-- Atlanticoncha +--+ +-- Potamilus
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 33 | 5 | 5 | -- |
+-- Glebula --+ +-- Cyrtonaias_berlandierii **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
-- Cyrtonaias_berlandierii **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- |
+----------- Obliquaria --+ | +-------- Cyrtonaias_berlandierii ** +--+ | +----- Leaunio +--+ | +-- Lampsilis +--+ +-- Pachynaias
Smith et al., 2020 [Publ 4129], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- |
+-------------- Obliquaria | --+ +----------- Cyrtonaias_berlandierii ** | | | | +-- Pachynaias +--+ +-----+ | | +-- Lampsilis | | +--+ +-- Ellipsaria | +--+ | | +-- Truncilla +--+ | +-- Atlanticoncha +--+ +-- Potamilus
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- |
+-- Glebula
+--------+
| +-- Cyrtonaias_berlandierii **
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias_berlandierii **
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Freeman et al., 2025 [Publ 4443], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 10 | 2 | 2 | -- |
+----- Obliquaria --+ | +-- Glebula +--+ +-- Cyrtonaias_berlandierii **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- |
-- Cyrtonaias_berlandierii **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- |
+----------- Obliquaria --+ | +-------- Cyrtonaias_berlandierii ** +--+ | +----- Leaunio +--+ | +-- Lampsilis +--+ +-- Pachynaias
Hoeh et al., 2002 [Publ 2653], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+-- Glebula --+ +-- Cyrtonaias_berlandierii **
Campbell et al., 2008 [Publ 2675], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- |
-- Cyrtonaias_berlandierii **
Ouyang et al., 2015 [Publ 4493], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- |
+----- Cyrtonaias_berlandierii ** --+ | +-- Lampsilis +--+ +-- Epioblasma
Burlakova et al., 2012 [Publ 3486], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
-- Cyrtonaias_berlandierii **
Chapman et al., 2008 [Publ 2667], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- |
+-------------------------- Toxolasma | --+ +----------------------- Obliquaria | | | | +-- Cyrtonaias_berlandierii ** +--+ +-----------------+ | | +-- Glebula | | +--+ +-- Ptychobranchus | +--------------+ | | +-- Lemiox | | +--+ +-- Venustaconcha | +-----------+ | | +-- Obovaria +--+ | +----------- Hamiota +--+ | +-------- Lampsilis_ovata +--+ | +----- Ortmanniana +--+ | +-- Lampsilis_straminea +--+ +-- Lampsilis_hydiana
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 42 | 4 | 3 | -- |
-- Cyrtonaias_berlandierii **
Campbell et al., 2008 [Publ 2675], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- |
-- Cyrtonaias_berlandierii **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- |
-- Cyrtonaias_berlandierii **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+-- Cyrtonaias_berlandierii **
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Ortmanniana
+--+
| +-- Ligumia
+--+
+-- Potamilus
