MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Publication Haas (1924): summary
Full Citation
Haas, F. 1924. Beiträge zu einer Monographie der asiatischen Unioniden. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 38: 129-203. Available electronically.
Attributed Nominal Genera
Chrysopseudodon Haas, 1920
Haas, 1920. Conch. Cab. 9 (pt. 2, h. 53): 317.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 129.
(intro'd as sub-gen. of Pseudodon Gould, 1844)
Type species: Psudodon [sic] aureus Heude, 1885 (orig. desig.) = Pseudobaphia aurea
Cosmopseudodon Haas, 1920
Haas, 1920. Conch. Cab. 9 (pt. 2, h. 53): 310.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 129.
(intro'd as sub-gen. of Pseudodon Gould, 1844)
Type species: Pseudodon resupinatus Martens, 1902 (orig. desig.) = Cosmopseudodon resupinatus
Diplopseudodon Haas, 1920
Haas, 1920. Conch. Cab. 9 (pt. 2, h. 53): 313.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 129.
(intro'd as sub-gen. of Pseudodon Gould, 1844)
Type species: Pseudodon crassus Drouët in Drouët & Chaper, 1892 (orig. desig.) = Pseudodon walpolei
Prohyriopsis Haas, 1914
Haas, 1914. Nachr. Bl. Dtsch. Malak. Ges. 46: 76.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 163.
Type species: Unio stolatus Martens, 1900 (orig. desig.) = Prohyriopsis stolata
Rhombuniopsis Haas, 1920
Haas, 1920. Senckenbergiana 2: 149.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 185.
Type species: Unio (Cuneopsis) tauriformis Fulton, 1906 (orig. desig.) = Rhombuniopsis tauriformis
Attributed Nominal Species
Unio sacellus ‘Drouet & Chaper’ of Authors = Lens lugens
‘Drouet & Chaper’ of Authors.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 818.
Simpson, 1914. Descr. Cat. Naiades: 1011.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 198.
(error for Unio saccellus Drouët & Chaper, 1892)
Parreysia simpularis ‘Heude’ of Authors = Ptychorhynchus murinum
‘Heude’ of Authors.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 845.
Simpson, 1914. Descr. Cat. Naiades: 1122.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 159.
Haas, 1969. Das Tierreich 88: 106.
He & Zhuang, 2013. Freshwater Bivalves China: 95.
(error for Unio simpularius Heude, 1884)
Pseudodon solidus Haas, 1911 = Ptychorhynchus murinum
Haas, 1911. Nachr. Bl. Dtsch. Malak. Ges. 43: 46.
Haas, 1913. Conch. Cab. 9 (pt. 2, h. 47): pl. 41, fig. 4.
Haas, 1924. Abh. Senckenb. Naturf. Ges. 38: 136.
Taxonomic Opinions
Arcidopsis footei (Theobald, 1876)
Arconaia lanceolata (Lea, 1856) = Lanceolaria lanceolata
+ Triquetra contorta Lea, 1857
+ Unio contortus var. muticus Heude, 1877
+ Unio contortus var. conjungens Heude, 1877
Chamberlainia hainesiana (Lea, 1856) = Chamberlainia hainesiana + Chamberlainia duclerci + Sinohyriopsis goliath
+ Unio imperialis Morelet, 1862
+ Dysnomia pavonina Rochebrune, 1882
+ Unio duclerci Rochebrune, 1882
+ Simpsonia demangei Rochebrune, 1904
Chamberlainia paviei (Morelet, 1891) = Chamberlainia hainesiana
+ Chamberlainia pavici Rochebrune, 1904
Contradens ascia ascia (Hanley, 1856) = Lens contradens
There are 113 species taxonomic opinions attributed to this publication.
Contradens ascia dimotus (Lea, 1870) = Lens contradens
Contradens ascia fischerianus (Morlet, 1883) = Lens micropterus
Contradens ascia lugens (Drouet & Chaper, 1892) = Lens lugens
Contradens ascia sacellus [sic] (Drouet & Chaper, 1892) = Lens lugens
Ctenodesma (Nesonaia) beauforti (Bavay, 1908) = Microdontia anodontaeformis
Ctenodesma (Nesonaia) lorentzi (Schepman, 1919) = Hyridella misoolensis
Ctenodesma (s.s.) borneensis (Issel, 1874) = Ctenodesma borneensis
Cuneopsis capitata (Heude, 1874) = Pseudocuneopsis capitata
Cuneopsis celtiformis (Heude, 1874)
Cuneopsis heudei (Heude, 1874) = Cuneopsis heudei + Cuneopsis szechenyii
+ Unio corderii Heude, 1875
+ Unio szechenyii Neumayr, 1899
Cuneopsis pisciculus (Heude, 1874) = Tchangsinaia piscicula
+ Unio retortus Martens, 1875
+ Unio flexuosus Neumayr, 1899
+ Unio flexicosus Simpson, 1914
Cuneopsis rufescens (Heude, 1874)
Diaurora aurorea (Heude, 1883)
Discomya rhomboidea (Simpson, 1900) = Discomya radulosa
Hyriopsis bialata Simpson, 1900
+ Unio delphinus Gruner, 1841
+ Unio megapterus Morelet, 1864
+ Unio delphinopterus Dautzenberg & Fischer, 1905
+ Hyriopsis gracilis Haas, 1910
Hyriopsis cumingii (Lea, 1852) = Sinohyriopsis cumingii + Quadrula verrucosa + Sinohyriopsis goliath
+ Unio gigas Sowerby, 1867
+ Hyriopsis goliath Rolle, 1904
Hyriopsis delaportei (Crosse & Fischer, 1876)
Hyriopsis myersiana (Lea, 1856)
+ Unio housei Lea, 1856
Hyriopsis schlegelii (von Martens, 1861) = Sinohyriopsis schlegelii
Hyriopsis sutrangensis (Morlet, 1889) = Hyriopsis myersiana
+ Unio stungtrangensis Fischer & Dautzenberg, 1904
+ Unio patrangensis Morlet, 1904
Hyriopsis velthuizeni (Schepman, 1895)
+ Hyriopsis velthuzeni Simpson, 1900
Lanceolaria gladiolus (Heude, 1877) = Lanceolaria gladiola
Lanceolaria grayana (Lea, 1834) = Lanceolaria grayii
Margaritanopsis woodthorpi (Gowin-Austen, 1919) = Gibbosula laosensis
Nodularia (s.s.) douglasiae (Griffith & Pidgeon, 1834) = Nodularia douglasiae
+ Unio kobelti Neumayr, 1899
+ Unio boettgeri Neumayr, 1899
Nodularia (s.s.) douglasiae taiwanica Pilsbry, 1905 = Nodularia douglasiae
Nodularia (s.s.) pantoensis (Neumayr, 1899) = Inversidens pantoensis
+ Nodularia continentalis Haas, 1910
Nodularia (Scabies) fluctigera (Lea, 1859)
Nodularia (Scabies) scobinata (Lea, 1856) = Scabies scobinatus + Nodularia persculpta
+ Nodularia persculpta Haas, 1910
+ Unio pellislacerti Morelet, 1865
+ Unio pellis-lacertae Haas, 1911
Oxynaia micheloti (Morlet, 1886) = Nodularia micheloti + Nodularia gladiator
+ Unio gladiator Ancey, 1881
Oxynaia pugio (Benson, 1862) = Indochinella pugio + Diplodon parallelopipedon
+ Unio digitiformis Sowerby, 1868
Physunio (s.s.) ferrugineus Annandale, 1918 = Lamellidens ferrugineus
Physunio (s.s.) micropteroides Annandale, 1918 = Lamellidens ferrugineus
Physunio (s.s.) micropterus (Morelet, 1866) = Lens micropterus
Physunio (Velunio) friersoni Simpson, 1914 = Lamellidens friersoni
+ Unio velaris ‘Benson, in Hanley’ Sowerby, 1868
Pressidens exanthematicus (Küster, 1861)
Prohyriopsis stolata (von Martens, 1900)
Pseudodon (Bineurus) thomsoni Morlet, 1884 = Songkhlanaia moreleti
Pseudodon (Monodontina) cambodjensis (Petit, 1865) = Pseudodon cambodjensis + Pseudodon vagulus
+ Unio subtrigonus Sowerby, 1867
Pseudodon (Monodontina) chaperi (Morgan, 1885) = Sundadontina cumingii
Pseudodon (s.s.) avae (Theobald, 1873) = Indopseudodon salwenianus
Pseudodon (s.s.) callifer (von Martens, 1860) = Thaiconcha callifera + Sundadontina harmandi
+ Pseudodon harmandi Crosse & Fischer, 1876
Pseudodon (s.s.) cumingii (Lea, 1850) = Sundadontina cumingii
Pseudodon (s.s.) inoscularis (Gould, 1844) = Pseudodon inoscularis
Pseudodon (s.s.) moreleti Crosse & Fischer, 1876 = Songkhlanaia moreleti
+ Monocondylea [sic] tumida Deshayes & Jullien, 1876
Pseudodon (s.s.) salweniana (Gould, 1844) = Indopseudodon salwenianus
Pseudodon (s.s.) solidus Haas, 1911 = Ptychorhynchus murinum
Pseudodon (s.s.) sulcatus Rochebrune, 1882 = Sundadontina sulcata
Pseudodon (s.s.) tumidus (Morelet, 1866) = Sundadontina tumida
Pseudodon (s.s.) walpolei (Hanley, 1871) = Pseudodon walpolei
Pseudodon (Trigonodon) crebristriata (Anthony, J.G., 1865) = Indopseudodon crebristriatus + Pseudodon inoscularis
+ Pseudodon (Trigonodon) crebristriatus var. curvata Preston, 1912
+ Unio vondembuschii Sowerby, 1868
Pseudodon (Trigonodon) peguensis (Anthony, J.G., 1865) = Indopseudodon salwenianus
Ptychorhynchus apicellatum (Heude, 1883) = Ptychorhynchus murinum
Ptychorhynchus murinum (Heude, 1883)
+ Unio compressus Heude, 1877
+ Unio morinus Paetel, 1890
+ Ptychorhynchus incertus Simpson, 1900
Ptychorhynchus pfisteri (Heude, 1874)
+ Unio pfisteri var. insperatus Heude, 1877
+ Ptychorhynchus pfisteri var. inspiratus Simpson, 1900
+ Unio mediastinus Heude, 1883
Ptychorhynchus schomburgianum (Heude, 1885)
Ptychorhynchus simpulare [sic] (Heude, 1884) = Ptychorhynchus murinum
+ Unio modestus Heude, 1877
Rhombuniopsis superstes (Neumayr, 1899) = Leoparreysia superstes + Rhombuniopsis heres
+ Unio heres Neumayr, 1899
Rhombuniopsis tauriformis (Fulton, 1906)
Simpsonella serradelli (Drouet, 1892) = Simpsonella gracilis
Trapezoideus prashadi Haas, 1922 = Arcidopsis footei
+ Arcidopsis footei ‘Theobald’ of Authors
Virgus (Leiovirgus) misoolensis (Schepman, 1897) = Hyridella misoolensis
