Pilsbry, H.A. 1920. Costa Rican land and freshwater mollusks. Proceedings of the Academy of Natural Sciences of Philadelphia 72: 2-10. Available electronically.
‘Lea’ Pilsbry, 1920. Proc. Acad. Nat. Sci. 72: 10.
(error for Unio macnielii Lea, 1869)

Pilsbry, 1920. Proc. Acad. Nat. Sci. 72: 9, fig. 6.
Type(s): LECTOTYPE [AS HOLOTYPE] ANSP_105225 (by R.I. Johnson, 1980: 278), Rio Tempisque, Filadelphia, Costa Rica.