Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Leoparreysiini

genus Leoparreysia Vikhrev, Bolotov & Aksenova in Bolotov et al., 2017: cladomics

9 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
analyzed in 45 cladograms in 28 publications
maximum number of terminals in an analysis = 5 (mean = 3.07, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 34
maximum number of included terminals = 5, (mean = 3.74 median = 4)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Leoparreysia cladogram distances
PCoA Ordination

NJ Tree of Leoparreysia cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2019 [Publ 3974], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 13 0 0 **
Cladomics cladogram Nitia Coelatura Indochinella Radiatula Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia canefrii

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 112 0 0 2.0
Cladomics cladogram Parreysia Leoparreysia burmana Leoparreysia whitteni Leoparreysia tavoyensis Leoparreysia olivacea Leoparreysia canefrii

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 106 0 0 0.0
Cladomics cladogram Parreysia Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia canefrii

Bolotov et al., 2022 [Publ 4239], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 53 0 0 0.0
Cladomics cladogram Parreysia Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Bolotov et al., 2022 [Publ 4239], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 53 0 0 0.0
Cladomics cladogram Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Nitia Coelatura aegyptiaca Coelatura kunenensis Nyassunio Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 53 0 0 2.0
Cladomics cladogram Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Nitia Coelatura aegyptiaca Coelatura kunenensis Nyassunio Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4250], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 44 0 0 0.0
Cladomics cladogram Indonaia Indochinella Radiatula Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Bolotov et al., 2022 [Publ 4250], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 0 0 0 2.0
Cladomics cladogram Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia canefrii

Bolotov et al., 2022 [Publ 4252], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 56 0 0 0.0
Cladomics cladogram Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Nitia Coelatura aegyptiaca Nyassunio Coelatura kunenensis Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4252], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 0 5 56 0 0 3.0
Cladomics cladogram Leoparreysia burmana Leoparreysia canefrii Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia whitteni Coelatura aegyptiaca Coelatura kunenensis Nitia Nyassunio Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2024 [Publ 4369], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 57 0 0 0.0
Cladomics cladogram Assamnaia Parreysia Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Bolotov et al., 2024 [Publ 4369], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 58 0 0 0
Cladomics cladogram Lamellidens Balwantia Trapezidens Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia canefrii

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 628 0 0 2.0
Cladomics cladogram Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Pseudomulleria Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies Assamnaia Parreysia Nitia monceti 2 Nitia teretiuscula Nitia monceti 1 Grandidieria Pseudospatha tanganyicensis 2 Moncetia Pseudospatha tanganyicensis 1 Coelatura leopoldvillensis 2 Coelatura leopoldvillensis 1 Coelatura mesafricana Coelatura aegyptiaca 3 Coelatura hauttecoeuri 2 Nitia acuminata Coelatura aegyptiaca 2 Coelatura alluaudi 2 Coelatura hauttecoeuri 1 Coelatura stuhlmanni 2 Coelatura aegyptiaca 1 Coelatura stuhlmanni 1 Coelatura alluaudi 1 Coelatura bakeri 1 Coelatura bakeri 2 Coelatura horei 3 Coelatura ratidota Coelatura 2 Coelatura horei 1 Coelatura 1 Coelatura horei 2 Coelatura gabonensis Coelatura stanleyvillensis Coelatura kunenensis Nyassunio Coelatura hypsiprymna 2 Coelatura mossambicensis 1 Coelatura hypsiprymna 3 Coelatura mossambicensis 2 Coelatura luapulaensis 2 Coelatura choziensis 2 Coelatura hypsiprymna 1 Coelatura choziensis 1 Coelatura luapulaensis 1 Coelatura briarti Prisodontopsis aviculaeformis 1 Prisodontopsis mweruensis 1 Prisodontopsis aviculaeformis 2 Prisodontopsis aviculaeformis 3 Prisodontopsis mweruensis 2

Konopleva et al., 2026 [Publ 4847], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 61 0 0 0
Cladomics cladogram Assamnaia Parreysia Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia tavoyensis Leoparreysia canefrii

Konopleva et al., 2026 [Publ 4847], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 61 0 0 0.0
Cladomics cladogram Assamnaia Parreysia Leoparreysia burmana Leoparreysia whitteni Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Konopleva et al., 2026 [Publ 4847], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 62 0 0 0.0
Cladomics cladogram Lamellidens Balwantia Trapezidens Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Konopleva et al., 2026 [Publ 4847], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 5 5 2 5 62 0 0 1.0
Cladomics cladogram Lamellidens Balwantia Trapezidens Leoparreysia whitteni Leoparreysia burmana Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 2 3 122 0 0 3.0
Cladomics cladogram Leoparreysia olivacea Leoparreysia tavoyensis 2 Leoparreysia canefrii Leoparreysia tavoyensis 1 Nitia Coelatura gabonensis Prisodontopsis mweruensis Coelatura kunenensis Coelatura aegyptiaca 1 Coelatura aegyptiaca 2 Prisodontopsis aviculaeformis Coelatura luapulaensis Coelatura choziensis Indochinella Scabies Radiatula Indonaia shurtleffiana Indonaia rugosa Indonaia andersoniana Indonaia caerulea Indonaia bonneaudii Lamellidens

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 79 0 0 2.0
Cladomics cladogram Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Coelatura Nitia Indonaia Indochinella Radiatula Scabies

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 0 3 198 0 0 3.0
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia canefrii Leoparreysia olivacea Indonaia Indochinella Scabies Radiatula Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Nitia Coelatura gabonensis Coelatura kunenensis Prisodontopsis

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 196 0 0 4.0
Cladomics cladogram Leoparreysia canefrii Leoparreysia tavoyensis Leoparreysia olivacea Indonaia Indochinella Scabies Radiatula Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Nitia Coelatura kunenensis Coelatura gabonensis Prisodontopsis

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 0 0 4.0
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia canefrii Leoparreysia olivacea Indonaia Indochinella Scabies Radiatula Coelatura aegyptiaca 2 Nitia Coelatura aegyptiaca 1 Coelatura gabonensis Coelatura kunenensis Prisodontopsis

Konopleva et al., 2020 [Publ 4072], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 0 3 9 0 0 3.0
Cladomics cladogram Lamellidens Trapezidens Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 2.0
Cladomics cladogram Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Coelatura Indonaia Scabies Radiatula

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 2.0
Cladomics cladogram Leoparreysia olivacea Leoparreysia canefrii Leoparreysia tavoyensis Coelatura Indonaia Radiatula myitkyinae Scabies Radiatula mouhoti Radiatula chaudhurii

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 38 0 0 --
Cladomics cladogram Leoparreysia olivacea Leoparreysia tavoyensis Coelatura Indochinella Radiatula Scabies

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Nitia Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Prisodontopsis Coelatura gabonensis Coelatura kunenensis Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 0 0 --
Cladomics cladogram Leoparreysia olivacea Leoparreysia tavoyensis Nitia Coelatura aegyptiaca 1 Coelatura aegyptiaca 2 Coelatura gabonensis Coelatura kunenensis Prisodontopsis Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Coelatura Indochinella Radiatula Scabies

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 26 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Nitia Coelatura Prisodontopsis Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Coelatura Prisodontopsis Indonaia Indochinella Scabies Harmandia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Coelatura Prisodontopsis Indonaia Indochinella Scabies Harmandia

Bolotov et al., 2022 [Publ 4250], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 13 0 0 --
Cladomics cladogram Parreysia Leoparreysia tavoyensis Leoparreysia olivacea

Chen et al., 2026 [Publ 4833], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 25 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Leoparreysia olivacea Nitia Coelatura Prisodontopsis Indochinella Indonaia Radiatula Songkhlanaia Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Radiatula Indochinella Nitia Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Coelatura gabonensis Prisodontopsis

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Radiatula Indochinella Nitia Coelatura aegyptiaca 1 Coelatura aegyptiaca 2 Coelatura gabonensis Prisodontopsis

Graf et al., 2014 [Publ 3591], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Nitia Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Coelatura gabonensis Coelatura kunenensis Prisodontopsis Coelatura choziensis 1 Coelatura choziensis 2 Coelatura luapulaensis 1 Coelatura luapulaensis 2 Coelatura luapulaensis 3 Coelatura luapulaensis 4 Coelatura luapulaensis 5

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Coelatura Indochinella

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Leoparreysia canefrii Parreysia Coelatura Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 0 0 --
Cladomics cladogram Coelatura Leoparreysia tavoyensis Radiatula

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Lamellidens Leoparreysia canefrii

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Scabies Grandidieria Pseudospatha Moncetia Nitia Coelatura aegyptiaca Coelatura alluaudi Coelatura bakeri Coelatura hauttecoeuri Coelatura stuhlmanni Coelatura leopoldvillensis Coelatura mesafricana Coelatura horei Coelatura ratidota Coelatura Coelatura gabonensis Coelatura stanleyvillensis Coelatura kunenensis Prisodontopsis aviculaeformis Coelatura briarti Prisodontopsis mweruensis Coelatura choziensis Coelatura hypsiprymna Coelatura luapulaensis Coelatura mossambicensis Nyassunio

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Indochinella Indonaia Scabies Grandidieria Pseudospatha Moncetia Nitia Coelatura aegyptiaca Coelatura alluaudi Coelatura bakeri Coelatura hauttecoeuri Coelatura stuhlmanni Coelatura leopoldvillensis Coelatura mesafricana Coelatura horei Coelatura ratidota Coelatura Coelatura gabonensis Coelatura stanleyvillensis Coelatura kunenensis Coelatura briarti Prisodontopsis mweruensis Prisodontopsis aviculaeformis Coelatura choziensis Coelatura hypsiprymna Coelatura luapulaensis Coelatura mossambicensis Nyassunio

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 0 0 --
Cladomics cladogram Leoparreysia tavoyensis Indonaia Indochinella Radiatula

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 0 0 --
Cladomics cladogram Leoparreysia burmana Unionetta Coelatura Nitia