MUSSELpdb
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freshwater mussels
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Data last updated
Tue 02 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Leoparreysiini
genus Leoparreysia Vikhrev, Bolotov & Aksenova in Bolotov et al., 2017: cladomics
9 Recent valid species (9 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 9 (9 total)
analyzed in 41 cladograms in 27 publications
maximum number of terminals in an analysis = 5 (mean = 2.88, median = 3)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 5, (mean = 3.57 median = 3)
proportion of cladograms that support basic monophyly = 1
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Bolotov et al., 2019 [Publ 3974], fig. 1
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 13 | 0 | 0 | ** |
+-- Nitia
+--+
| +-- Coelatura
+-----+
| | +-- Indochinella
| +--+
| +-- Radiatula
--+
| +-- Leoparreysia_whitteni **
| +-----+
| | +-- Leoparreysia_burmana **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_tavoyensis **
+--+
+-- Leoparreysia_canefrii **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 112 | 0 | 0 | 2.0 |
+----------- Parreysia | --+ +-- Leoparreysia_burmana ** | +-----+ | | +-- Leoparreysia_whitteni ** +--+ | +----- Leoparreysia_tavoyensis ** +--+ | +-- Leoparreysia_olivacea ** +--+ +-- Leoparreysia_canefrii **
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 106 | 0 | 0 | 0.0 |
+----------- Parreysia | --+ +-- Leoparreysia_burmana ** | +-----+ | | +-- Leoparreysia_whitteni ** +--+ | +----- Leoparreysia_olivacea ** +--+ | +-- Leoparreysia_tavoyensis ** +--+ +-- Leoparreysia_canefrii **
Bolotov et al., 2022 [Publ 4239], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 53 | 0 | 0 | 0.0 |
+----------- Parreysia | --+ +-- Leoparreysia_burmana ** | +-----+ | | +-- Leoparreysia_whitteni ** +--+ | +----- Leoparreysia_olivacea ** +--+ | +-- Leoparreysia_canefrii ** +--+ +-- Leoparreysia_tavoyensis **
Bolotov et al., 2022 [Publ 4239], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 53 | 0 | 0 | 0.0 |
+-- Leoparreysia_burmana **
+-----+
| +-- Leoparreysia_whitteni **
+--------------+
| | +----- Leoparreysia_olivacea **
| +--+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_tavoyensis **
--+
| +-------- Nitia
| +-----------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
+--+ +--+
| +-- Nyassunio
|
| +----------------- Indonaia
+--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4239], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 53 | 0 | 0 | 2.0 |
+----------- Leoparreysia_burmana **
+-----------+
| | +-------- Leoparreysia_whitteni **
| +--+
| | +----- Leoparreysia_olivacea **
| +--+
| | +-- Leoparreysia_canefrii **
| +--+
--+ +-- Leoparreysia_tavoyensis **
|
| +-------- Nitia
| +-----------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
+--+ +--+
| +-- Nyassunio
|
| +----------------- Indonaia
+--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 44 | 0 | 0 | 0.0 |
+----- Indonaia
+-----+
| | +-- Indochinella
| +--+
| +-- Radiatula
--+
| +-- Leoparreysia_whitteni **
| +-----+
| | +-- Leoparreysia_burmana **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_canefrii **
+--+
+-- Leoparreysia_tavoyensis **
Bolotov et al., 2022 [Publ 4250], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 0 | 0 | 0 | 2.0 |
+----------- Leoparreysia_burmana ** --+ | +-------- Leoparreysia_whitteni ** +--+ | +----- Leoparreysia_olivacea ** +--+ | +-- Leoparreysia_tavoyensis ** +--+ +-- Leoparreysia_canefrii **
Bolotov et al., 2022 [Publ 4252], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 56 | 0 | 0 | 0.0 |
+-- Leoparreysia_burmana **
+-----+
| +-- Leoparreysia_whitteni **
+--------------+
| | +----- Leoparreysia_olivacea **
| +--+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_tavoyensis **
--+
| +-------- Nitia
| +-----------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Nyassunio
+--+ +--+
| +-- Coelatura_kunenensis
|
| +----------------- Indonaia
+--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4252], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 0 | 5 | 56 | 0 | 0 | 3.0 |
+-- Leoparreysia_burmana **
|
+-- Leoparreysia_canefrii **
|
+--------------------+-- Leoparreysia_olivacea **
| |
| +-- Leoparreysia_tavoyensis **
| |
| +-- Leoparreysia_whitteni **
|
--+ +-- Coelatura_aegyptiaca
| |
| +-- Coelatura_kunenensis
| +-----------------+
| | +-- Nitia
| | |
+--+ +-- Nyassunio
|
| +----------------- Indonaia
+--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2024 [Publ 4369], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 57 | 0 | 0 | 0.0 |
+-- Assamnaia
+--------+
| +-- Parreysia
|
--+ +-- Leoparreysia_burmana **
| +-----+
| | +-- Leoparreysia_whitteni **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_canefrii **
+--+
+-- Leoparreysia_tavoyensis **
Bolotov et al., 2024 [Publ 4369], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 58 | 0 | 0 | 0 |
+----- Lamellidens
+-----+
| | +-- Balwantia
| +--+
| +-- Trapezidens
--+
| +-- Leoparreysia_whitteni **
| +-----+
| | +-- Leoparreysia_burmana **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_tavoyensis **
+--+
+-- Leoparreysia_canefrii **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 627 | 0 | 0 | 2.0 |
+----------- Leoparreysia_burmana **
+--------------------------------------------+
| | +-------- Leoparreysia_whitteni **
| +--+
| | +----- Leoparreysia_olivacea **
| +--+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_tavoyensis **
--+
| +----------------------------------------------------- Pseudomulleria
| |
| | +-- Pseudoscabies
| | +--------------+
| | | +-- Indonaia
| | +--------------------------------+
| | | | +-------------- Indochinella
+--+ | | |
| | +--+ +----------- Radiatula
| | | |
| | +--+ +-- Scabiellus
| | | +-----+
| | | | +-- Unionetta_fabagina__2
| | +--+
| | | +----- Harmandia
+--+ +--+
| | +-- Unionetta_fabagina__1
| +--+
| +-- Scabies
|
| +-- Assamnaia
| +--------------------------------------------+
| | +-- Parreysia
| |
| | +----- Nitia_monceti__2
+--+ +--------------------------------------+
| | | +-- Nitia_teretiuscula
| | +--+
| | +-- Nitia_monceti__1
+--+
| +----------------------------------------- Grandidieria
| |
| | +----- Pseudospatha_tanganyicensis__2
| | +--------------------------------+
+--+ | | +-- Moncetia
| | +--+
| | +-- Pseudospatha_tanganyicensis__1
| |
| | +----- Coelatura_leopoldvillensis__2
+--+ +-----------------------------+
| | | +-- Coelatura_leopoldvillensis__1
| | +--+
| | +-- Coelatura_mesafricana
| |
| | +-------------------------------- Coelatura_aegyptiaca__3
| | |
+--+ | +-------- Coelatura_hauttecoeuri__2
| | |
| | +-------- Nitia_acuminata
| | |
| | | +----- Coelatura_aegyptiaca__2
| | +--------------------+ |
| | | | +----- Coelatura_alluaudi__2
+--+ | | |
| | | +----- Coelatura_hauttecoeuri__1
| | | |
| | +--+----- Coelatura_stuhlmanni__2
| | |
| | | +-- Coelatura_aegyptiaca__1
| | | |
| | | +-- Coelatura_stuhlmanni__1
| | | |
| | +--+-- Coelatura_alluaudi__1
+--+ |
| +-- Coelatura_bakeri__1
| |
| +-- Coelatura_bakeri__2
|
| +----------- Coelatura_horei__3
| |
| +--------------+ +-------- Coelatura_ratidota
| | | |
| | +--+ +----- Coelatura__2
| | | |
| | +--+ +-- Coelatura_horei__1
| | | |
+--+ +--+-- Coelatura__1
| |
| +-- Coelatura_horei__2
|
| +----------------------- Coelatura_gabonensis
| |
+--+ +-------------------- Coelatura_stanleyvillensis
| |
| | +----------------- Coelatura_kunenensis
+--+ |
| | +-------------- Nyassunio
| | |
+--+ | +----------- Coelatura_hypsiprymna__2
| | |
| | +----------- Coelatura_mossambicensis__1
| | |
+--+ | +-- Coelatura_hypsiprymna__3
| +--------+
| | +-- Coelatura_mossambicensis__2
| |
| | +----- Coelatura_luapulaensis__2
| | |
| | +----- Coelatura_choziensis__2
+--+-----+
| +----- Coelatura_hypsiprymna__1
| |
| | +-- Coelatura_choziensis__1
| +--+
| +-- Coelatura_luapulaensis__1
|
| +----- Coelatura_briarti
| +--+
| | | +-- Prisodontopsis_aviculaeformis__1
| | +--+
+--+ +-- Prisodontopsis_mweruensis__1
|
| +-- Prisodontopsis_aviculaeformis__2
| |
+-----+-- Prisodontopsis_aviculaeformis__3
|
+-- Prisodontopsis_mweruensis__2
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 2 | 3 | 122 | 0 | 0 | 3.0 |
+-------- Leoparreysia_olivacea **
+--------+
| | +----- Leoparreysia_tavoyensis__2 **
| +--+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_tavoyensis__1 **
|
| +-- Nitia
| |
| +-- Coelatura_gabonensis
| |
--+ +-- Prisodontopsis_mweruensis
| |
| +-- Coelatura_kunenensis
| |
| +-----------+-- Coelatura_aegyptiaca__1
| | |
| | +-- Coelatura_aegyptiaca__2
| | |
| | +-- Prisodontopsis_aviculaeformis
| | |
| | +-- Coelatura_luapulaensis
+--+ |
| +-- Coelatura_choziensis
|
| +----- Indochinella
| +-----+
| | | +-- Scabies
| | +--+
+--+ +-- Radiatula
|
| +-------- Indonaia_shurtleffiana
| |
+--+ +----- Indonaia_rugosa
| |
| | +-- Indonaia_andersoniana
+--+--+
| +-- Indonaia_caerulea
|
| +-- Indonaia_bonneaudii
+--+
+-- Lamellidens
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 79 | 0 | 0 | 2.0 |
+----- Leoparreysia_olivacea **
+--------+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_tavoyensis **
--+
| +-- Coelatura
| +--------+
| | +-- Nitia
+--+
| +-------- Indonaia
+--+
| +----- Indochinella
+--+
| +-- Radiatula
+--+
+-- Scabies
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 0 | 3 | 198 | 0 | 0 | 3.0 |
+-- Leoparreysia_tavoyensis **
|
+-----------+-- Leoparreysia_canefrii **
| |
| +-- Leoparreysia_olivacea **
|
--+ +-------- Indonaia
| +--+
| | | +----- Indochinella
| | +--+
| | | +-- Scabies
+--+ +--+
| +-- Radiatula
|
| +-------- Coelatura_aegyptiaca__2
| |
+--+ +----- Coelatura_aegyptiaca__1
| |
| +----- Nitia
+--+
| +-- Coelatura_gabonensis
| |
+--+-- Coelatura_kunenensis
|
+-- Prisodontopsis
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 196 | 0 | 0 | 4.0 |
+----- Leoparreysia_canefrii **
+-----------+
| | +-- Leoparreysia_tavoyensis **
| +--+
| +-- Leoparreysia_olivacea **
|
--+ +-------- Indonaia
| +-----+
| | | +----- Indochinella
| | +--+
| | | +-- Scabies
+--+ +--+
| +-- Radiatula
|
| +----------- Coelatura_aegyptiaca__2
| |
+--+ +-- Coelatura_aegyptiaca__1
| +-----+
| | +-- Nitia
+--+
| +----- Coelatura_kunenensis
+--+
| +-- Coelatura_gabonensis
+--+
+-- Prisodontopsis
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 198 | 0 | 0 | 4.0 |
+----- Leoparreysia_tavoyensis **
+--------+
| | +-- Leoparreysia_canefrii **
| +--+
| +-- Leoparreysia_olivacea **
|
--+ +-------- Indonaia
| +--+
| | | +----- Indochinella
| | +--+
| | | +-- Scabies
| | +--+
+--+ +-- Radiatula
|
| +-------- Coelatura_aegyptiaca__2
| |
| +-------- Nitia
+--+
| +----- Coelatura_aegyptiaca__1
| |
+--+ +-- Coelatura_gabonensis
| |
+--+-- Coelatura_kunenensis
|
+-- Prisodontopsis
Konopleva et al., 2020 [Publ 4072], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 0 | 3 | 9 | 0 | 0 | 3.0 |
+-- Lamellidens
+--+
| +-- Trapezidens
--+
| +-- Leoparreysia_olivacea **
| |
+--+-- Leoparreysia_canefrii **
|
+-- Leoparreysia_tavoyensis **
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 88 | 0 | 0 | 2.0 |
+----- Leoparreysia_olivacea **
+-----+
| | +-- Leoparreysia_canefrii **
| +--+
--+ +-- Leoparreysia_tavoyensis **
|
| +-------- Coelatura
+--+
| +----- Indonaia
+--+
| +-- Scabies
+--+
+-- Radiatula
Wu et al., 2021 [Publ 4366], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 88 | 0 | 0 | 2.0 |
+----- Leoparreysia_olivacea **
+-----------+
| | +-- Leoparreysia_canefrii **
| +--+
--+ +-- Leoparreysia_tavoyensis **
|
| +-------------- Coelatura
+--+
| +----------- Indonaia
+--+
| +-------- Radiatula_myitkyinae
+--+
| +----- Scabies
+--+
| +-- Radiatula_mouhoti
+--+
+-- Radiatula_chaudhurii
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 38 | 0 | 0 | -- |
+-- Leoparreysia_olivacea **
+--------+
| +-- Leoparreysia_tavoyensis **
--+
| +-------- Coelatura
+--+
| +----- Indochinella
+--+
| +-- Radiatula
+--+
+-- Scabies
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 137 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+-----------------+
| +-- Leoparreysia_olivacea **
|
| +-------------- Nitia
| +--+
--+ | | +----------- Coelatura_aegyptiaca__2
| | +--+
| | | +-------- Coelatura_aegyptiaca__1
| | +--+
| | | +----- Prisodontopsis
+--+ +--+
| | +-- Coelatura_gabonensis
| +--+
| +-- Coelatura_kunenensis
|
| +-------------- Indonaia
| |
+--+ +-- Indochinella
| +--------+
| | +-- Scabies_crispata
+--+
| +-------- Radiatula
+--+
| +----- Scabies_mandarinus
+--+
| +-- Scabies_humilis
+--+
+-- Scabies_phaselus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 137 | 0 | 0 | -- |
+-- Leoparreysia_olivacea **
+-----------------+
| +-- Leoparreysia_tavoyensis **
|
| +-------------- Nitia
| +--+
--+ | | +----------- Coelatura_aegyptiaca__1
| | +--+
| | | +-------- Coelatura_aegyptiaca__2
| | +--+
| | | +----- Coelatura_gabonensis
+--+ +--+
| | +-- Coelatura_kunenensis
| +--+
| +-- Prisodontopsis
|
| +-------------- Indonaia
| |
+--+ +-- Indochinella
| +--------+
| | +-- Scabies_crispata
+--+
| +-------- Radiatula
+--+
| +----- Scabies_mandarinus
+--+
| +-- Scabies_humilis
+--+
+-- Scabies_phaselus
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+--------+
| +-- Leoparreysia_olivacea **
--+
| +-------- Coelatura
+--+
| +----- Indochinella
+--+
| +-- Radiatula
+--+
+-- Scabies
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 26 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+-----------------------+
| +-- Leoparreysia_olivacea **
|
--+ +----- Nitia
| +-----------------+
| | | +-- Coelatura
| | +--+
+--+ +-- Prisodontopsis
|
| +-------------------- Indonaia
+--+
| +----------------- Indochinella
+--+
| +-------------- Radiatula
+--+
| +----------- Scabiellus
+--+
| +-------- Unionetta_fabagina__2
+--+
| +----- Harmandia
+--+
| +-- Unionetta_fabagina__1
+--+
+-- Scabies
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+-----------+
| +-- Leoparreysia_olivacea **
|
--+ +-- Coelatura
| +--------+
| | +-- Prisodontopsis
+--+
| +-------- Indonaia
+--+
| +----- Indochinella
+--+
| +-- Scabies
+--+
+-- Harmandia
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+-----------+
| +-- Leoparreysia_olivacea **
|
--+ +-- Coelatura
| +--------+
| | +-- Prisodontopsis
+--+
| +-------- Indonaia
+--+
| +----- Indochinella
+--+
| +-- Scabies
+--+
+-- Harmandia
Bolotov et al., 2022 [Publ 4250], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 13 | 0 | 0 | -- |
+----- Parreysia --+ | +-- Leoparreysia_tavoyensis ** +--+ +-- Leoparreysia_olivacea **
Chen et al., 2026 [Publ 4833], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 25 | 0 | 0 | -- |
+-- Leoparreysia_tavoyensis **
+--------------------+
| +-- Leoparreysia_olivacea **
|
| +----- Nitia
--+ +--------------+
| | | +-- Coelatura
| | +--+
| | +-- Prisodontopsis
+--+
| +-- Indochinella
| +--------------+
| | +-- Indonaia
+--+
| +-------------- Radiatula
| |
+--+ +----------- Songkhlanaia
| |
+--+ +-------- Harmandia
| |
+--+ +-- Unionetta_xishuangbannaensis
| +--+
| | +-- Unionetta_fabagina__2
+--+
| +-- Unionetta_fabagina__1
+--+
+-- Scabies
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+----------------- Leoparreysia_tavoyensis ** | --+ +-- Radiatula | +-----------+ | | +-- Indochinella +--+ | +----------- Nitia +--+ | +-------- Coelatura_aegyptiaca__2 +--+ | +----- Coelatura_aegyptiaca__1 +--+ | +-- Coelatura_gabonensis +--+ +-- Prisodontopsis
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+-------------- Leoparreysia_tavoyensis ** | --+ +-- Radiatula | +--------+ | | +-- Indochinella +--+ | +-------- Nitia | | +--+ +----- Coelatura_aegyptiaca__1 | | +--+----- Coelatura_aegyptiaca__2 | | +-- Coelatura_gabonensis +--+ +-- Prisodontopsis
Graf et al., 2014 [Publ 3591], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
+----------------------- Leoparreysia_tavoyensis ** | --+ +-------------------- Nitia | | +--+ +----------------- Coelatura_aegyptiaca__2 | | +--+ +-------------- Coelatura_aegyptiaca__1 | | +--+ +----------- Coelatura_gabonensis | | +--+ +-------- Coelatura_kunenensis | | | | +----- Prisodontopsis +--+ | | | +-- Coelatura_choziensis__1 | | | +--+ +-- Coelatura_choziensis__2 | | | +-- Coelatura_luapulaensis__1 | | +--+-- Coelatura_luapulaensis__2 | +-- Coelatura_luapulaensis__3 | +-- Coelatura_luapulaensis__4 | +-- Coelatura_luapulaensis__5
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 80 | 0 | 0 | -- |
+----- Leoparreysia_tavoyensis ** --+ | +-- Coelatura +--+ +-- Indochinella
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 0 | 0 | -- |
+-------------------------- Leoparreysia_canefrii ** --+ | +----------------------- Parreysia +--+ | +-------------------- Coelatura +--+ | +----------------- Indonaia +--+ | +-------------- Indochinella +--+ | +----------- Radiatula +--+ | +-------- Scabiellus +--+ | +----- Harmandia +--+ | +-- Unionetta +--+ +-- Scabies
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 121 | 0 | 0 | -- |
+-- Coelatura | --+-- Leoparreysia_tavoyensis ** | +-- Radiatula
Choi et al., 2020 [Publ 4169], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Lamellidens --+ +-- Leoparreysia_canefrii **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+-------------------------------------- Leoparreysia_tavoyensis ** | | +-------- Indonaia | | | +--------------------------+ +----- Indochinella --+ | | | | | +--+ +-- Radiatula_mouhoti | | | | | | | +-- Radiatula_myitkyinae | | +--+ +--+ +-- Radiatula_chaudhurii | | | +-- Scabies | | +-------------------------------- Grandidieria | | | | +-- Pseudospatha +--+ +--------------------------+ | | +-- Moncetia | | | | +-------------------------- Nitia +--+ | | | +-- Coelatura_aegyptiaca | | | | | +-- Coelatura_alluaudi +--+ | | +--------------------+-- Coelatura_bakeri | | | | | +-- Coelatura_hauttecoeuri | | | | | +-- Coelatura_stuhlmanni +--+ | +-- Coelatura_leopoldvillensis | +-----------------+ | | +-- Coelatura_mesafricana | | | | +-- Coelatura_horei +--+ | | +--------------+-- Coelatura_ratidota | | | | | +-- Coelatura +--+ | +-------------- Coelatura_gabonensis | | | +-------------- Coelatura_stanleyvillensis +--+ | +----------- Coelatura_kunenensis | | | | +----- Prisodontopsis_aviculaeformis +--+ +--+ | | | +-- Coelatura_briarti | | +--+ | | +-- Prisodontopsis_mweruensis +--+ | +-- Coelatura_choziensis | | | +-- Coelatura_hypsiprymna | | +-----+-- Coelatura_luapulaensis | +-- Coelatura_mossambicensis | +-- Nyassunio
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+-------------------------------- Leoparreysia_tavoyensis ** | | +-- Radiatula_mouhoti | | | +-- Radiatula_myitkyinae | | --+ +-- Radiatula_chaudhurii | +--------------------------+ | | +-- Indochinella | | | | | +-- Indonaia | | | | | +-- Scabies +--+ | +----- Grandidieria | +--------------------+ | | | +-- Pseudospatha | | +--+ | | +-- Moncetia | | +--+ +----------------------- Nitia | | | | +-- Coelatura_aegyptiaca | | | | | +-- Coelatura_alluaudi +--+ | | +-----------------+-- Coelatura_bakeri | | | | | +-- Coelatura_hauttecoeuri | | | | | +-- Coelatura_stuhlmanni +--+ | +-- Coelatura_leopoldvillensis | +--------------+ | | +-- Coelatura_mesafricana | | | | +-- Coelatura_horei +--+ | | +-----------+-- Coelatura_ratidota | | | | | +-- Coelatura | | +--+ +-- Coelatura_gabonensis | +--------+ | | +-- Coelatura_stanleyvillensis | | +--+ +-------- Coelatura_kunenensis | | | | +-- Coelatura_briarti | | | +--+ +--+-- Prisodontopsis_mweruensis | | | | | +-- Prisodontopsis_aviculaeformis | | +--+ +-- Coelatura_choziensis | | | +-- Coelatura_hypsiprymna | | +--+-- Coelatura_luapulaensis | +-- Coelatura_mossambicensis | +-- Nyassunio
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 55 | 0 | 0 | -- |
+-------- Leoparreysia_tavoyensis ** --+ | +----- Indonaia +--+ | +-- Indochinella +--+ +-- Radiatula
Zieritz et al., 2024 [Publ 4381], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 44 | 0 | 0 | -- |
+-------- Leoparreysia_burmana ** --+ | +----- Unionetta +--+ | +-- Coelatura +--+ +-- Nitia
