MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Cambarunio Watters, 2018: cladomics
7 Recent valid species (7 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 7 (7 total)
analyzed in 36 cladograms in 24 publications
maximum number of terminals in an analysis = 4 (mean = 1.08, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 1
maximum number of included terminals = 4, (mean = 4 median = 4)
proportion of cladograms that support basic monophyly = 0
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 68 | 76 | 4 | 628 | 89 | 26 | 0 |
+-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus **
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus **
+--+ | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Cambarunio_simus **
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
+--+ | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- |
+----------- Ligumia | +----------- Truncilla | | +-- Ptychobranchus +--------+ --+ +-- Sagittunio | | +-- Lampsilis_siliquoidea | +-----+ | | +-- Epioblasma | | +--+ +----- Ortmanniana | | | +----- Cambarunio_iris ** +--+ +----- Leaunio | | +-- Lampsilis_cardium +--+ +-- Lampsilis_fasciola
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- |
+-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio_iris **
|
+-- Pachynaias
|
+-- Mesonaias
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio_iris **
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Bolotov et al., 2018 [Publ 3826], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 81 | 2 | 2 | -- |
-- Cambarunio_iris **
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- |
+----- Truncilla | +----- Ptychobranchus --+ | +-- Ortmanniana | | | +-- Cambarunio_iris ** +--+ +-- Ligumia | +-- Epioblasma
Bolotov et al., 2017 [Publ 3761], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 137 | 2 | 2 | -- |
+----- Cambarunio_iris ** --+ | +-- Ortmanniana +--+ +-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- |
+----------------------- Cyrtonaias | --+ +-------------------- Ptychobranchus | | +--+ +----------------- Ligumia | | +--+ +-------------- Potamilus | | +--+ +----------- Pachynaias | | | | +-- Epioblasma +--+ +-----+ | | +-- Truncilla | | +--+ +-- Mesonaias | +--+ | | +-- Cambarunio_iris ** +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 17 | 16 | 5 | -- |
+----------------------- Obliquaria | | +-- Cambarunio_iris ** | +-----------------+ | | +-- Leaunio --+ | | | +----------- Lampsilis_teres | | | | +--------+ +-------- Ortmanniana | | | | | | +--+ +-- Lampsilis_hydiana +--+ | +--+ | | | +-- Lampsilis_virescens | +--+ | | +-- Lampsilis_siliquoidea | +--+ | +-- Lampsilis_radiata | | +----------------- Obovaria | | +--+ +-------------- Lampsilis_bracteata | | +--+ +----------- Lampsilis_ornata | | +--+ +-------- Lampsilis_ovata | | +--+ +----- Lampsilis_cardium__3 | | +--+ +-- Lampsilis_satura | | +--+-- Lampsilis_cardium__1 | +-- Lampsilis_cardium__2
Bolotov et al., 2017 [Publ 3761], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 137 | 2 | 2 | -- |
+----- Ortmanniana --+ | +-- Lampsilis +--+ +-- Cambarunio_iris **
Buhay et al., 2002 [Publ 2661], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 3 | -- |
+----- Lampsilis | --+----- Epioblasma | | +-- Cambarunio_taeniatus ** +--+ +-- Villosa
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- |
+----------------------- Ellipsaria | --+ +-------------------- Truncilla | | +--+ +----------------- Obliquaria | | | | +-------------- Potamilus +--+ | | | +----- Lampsilis_ornata | | +-----+ | | | | +-- Lampsilis_cardium +--+ | +--+ | | +-- Lampsilis_ovata | | | | +-------- Lampsilis_teres | +--+ +--+ | +----- Lampsilis_siliquoidea | +--+ | | +-- Ortmanniana | +--+ | +-- Lampsilis_higginsii | | +----- Hamiota | | +-----+ +-- Epioblasma | | +--+-- Cambarunio_iris ** | +-- Ligumia
Grobler et al., 2011 [Publ 3359], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 16 | 15 | 9 | -- |
+-------- Lemiox
+-----+
| | +----- Medionidus
| +--+
| | +-- Dromus
| +--+
--+ +-- Cyprogenia
|
| +----------- Potamilus
| |
| | +----- Lampsilis_siliquoidea
| | +--+
+--+ | | +-- Lampsilis_higginsii
| | +--+
| | +-- Ortmanniana
| |
| | +----- Lampsilis_fasciola
+--+--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-- Cambarunio_iris **
| |
+-----+-- Ligumia
|
+-- Epioblasma
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- |
+-------------- Obliquaria | +-------------- Truncilla --+ +-------------- Ptychobranchus | | +----------- Ligumia +--+ | +-------- Cambarunio_iris ** +--+ | +----- Epioblasma +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- |
+-------------- Potamilus_leptodon | | +----------- Potamilus_alatus --+ | | | +-------- Venustaconcha | | | +--+ +-------- Leaunio | | | | +-- Ellipsaria | +-----+ +--+ +-- Truncilla | | +-- Medionidus +-----+ | +-- Ptychobranchus | | +----- Cambarunio_iris ** +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- |
+-- Cambarunio_iris ** --+ +-- Ptychobranchus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- |
+-------------------- Truncilla | --+ +----------------- Ligumia | | | | +-- Ptychobranchus +--+ +-----------+ | | +-- Sagittunio | | +--+ +----------- Lampsilis_siliquoidea | | | | +----- Ortmanniana +--+ +--+ | | | +-- Lampsilis_cardium | | +--+ +--+ +-- Lampsilis_fasciola | | +----- Epioblasma +--+ | +-- Cambarunio_iris ** +--+ +-- Leaunio
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- |
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio_iris **
+--+
+-- Ligumia
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 17 | 16 | 5 | -- |
+----------------- Lampsilis_teres | +----------------- Obovaria | | +-- Leaunio +--------------+ | +-- Cambarunio_iris ** | | +-------- Ortmanniana | | --+--------+ +-- Lampsilis_hydiana | | +--+ | | | +-- Lampsilis_virescens | +--+ | | +-- Lampsilis_siliquoidea | +--+ | +-- Lampsilis_radiata | | +-------------- Lampsilis_bracteata | | +--+ +----------- Lampsilis_ornata | | +--+ +-------- Lampsilis_ovata | | +--+ +----- Lampsilis_cardium__3 | | +--+ +-- Lampsilis_satura | | +--+-- Lampsilis_cardium__1 | +-- Lampsilis_cardium__2
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- |
+----- Ortmanniana --+ | +-- Cambarunio_iris ** +--+ +-- Lampsilis
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio_iris **
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 79 | 2 | 2 | -- |
+----- Cambarunio_iris ** --+ | +-- Lampsilis +--+ +-- Ortmanniana
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 197 | 13 | 11 | -- |
+-------------------- Potamilus_leptodon | --+ +----------------- Potamilus_alatus | | | | +-- Medionidus +--+ +-----------+ | | +-- Ptychobranchus | | +--+ +-- Ellipsaria | +--------+ | | +-- Truncilla +--+ | +-------- Leaunio | | +--+ +-- Venustaconcha | +--+ | | +-- Cambarunio_iris ** +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- |
+----- Ortmanniana --+ | +-- Lampsilis +--+ +-- Cambarunio_iris **
Bolotov et al., 2017 [Publ 3764], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 123 | 2 | 2 | -- |
-- Cambarunio_iris **
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- |
+----------------- Ptychobranchus --+ | +-------------- Truncilla +--+ | +----------- Ortmanniana +--+ | +-------- Lampsilis +--+ | +----- Cambarunio_iris ** +--+ | +-- Ligumia +--+ +-- Epioblasma
Inoue et al., 2013 [Publ 3528], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 16 | 9 | 5 | -- |
+-------- Lampsilis | --+ +----- Leaunio | | +--+----- Obovaria | | +-- Villosa +--+ +-- Cambarunio_iris **
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- |
+----------------- Ptychobranchus --+ | +-------------- Truncilla +--+ | +----------- Ortmanniana +--+ | +-------- Lampsilis +--+ | +----- Cambarunio_iris ** +--+ | +-- Ligumia +--+ +-- Epioblasma
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- |
+----- Ellipsaria | +----- Potamilus_leptodon | +----- Medionidus | +----- Potamilus_alatus | +----- Ptychobranchus --+ +----- Truncilla | +----- Venustaconcha | +----- Cambarunio_iris ** | +----- Leaunio | | +-- Ortmanniana +--+ +-- Lampsilis
Chase et al., 2018 [Publ 3847], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 38 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Venustaconcha | | +--+ +-- Lemiox | +--------+ | | +-- Lampsilis +--+ | +-------- Potamilus +--+ | +----- Cambarunio_iris ** +--+ | +-- Ellipsaria +--+ +-- Truncilla
Breton et al., 2011 [Publ 3376], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Ellipsaria | | +--+ +----------- Truncilla | | +--+ +-- Lemiox | +-----+ | | +-- Potamilus +--+ | +----- Lampsilis +--+ | +-- Cambarunio_iris ** +--+ +-- Venustaconcha
Bolotov et al., 2020 [Publ 4084], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 2 | 2 | -- |
-- Cambarunio_iris **
Buhay et al., 2002 [Publ 2661], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 3 | -- |
+----- Lampsilis | --+----- Epioblasma | | +-- Cambarunio_taeniatus ** +--+ +-- Villosa
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio_iris **
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- |
+-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio_iris **
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- |
+----------------- Toxolasma | | +-------------- Potamilus | | | | +-- Ellipsaria | +-----------+ --+ | +-- Truncilla | | | | +----- Medionidus | +--------+ | | | +-- Lemiox | | +--+ +--+ +-- Dromus | | +----------- Cambarunio_simus ** | | | +----------- Leaunio | | | +----------- Villosa | | | | +-- Sagittunio | +--------+ +--+ +-- Lampsilis_ovata | | +----- Venustaconcha +-----+ | | +-- Epioblasma | +--+ | +-- Obovaria | | +-------- Lampsilis_teres +--+ | +----- Lampsilis_siliquoidea +--+ | +-- Lampsilis_hydiana +--+ +-- Lampsilis_straminea
