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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Scabiellus Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020: cladomics
     1 Recent valid species (1 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 1 (1 total)
analyzed in 12 cladograms in 7 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)


Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110161287--
                 +-- Pseudoscabies           
  +--------------+                           
  |              +-- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
110162287--
                 +-- Pseudoscabies           
  +--------------+                           
  |              +-- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
1101111196--
  +-------------------- Indonaia                
  |                                             
--+  +----------------- Indochinella            
  |  |                                          
  |  |           +----- Radiatula_mouhoti       
  +--+  +--------+                              
     |  |        |  +-- Radiatula_myitkyinae    
     |  |        +--+                           
     +--+           +-- Radiatula_chaudhurii    
        |                                       
        |  +----------- Radiatula_crispisulcata 
        +--+                                    
           |  +-------- Scabiellus_songkramensis **
           +--+                                 
              |  +----- Harmandia               
              +--+                              
                 |  +-- Unionetta               
                 +--+                           
                    +-- Scabies                 

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
110127166--
  +-------------------- Indonaia                
--+                                             
  |  +----------------- Indochinella            
  +--+                                          
     |  +-------------- Radiatula               
     +--+                                       
        |  +----------- Scabiellus_songkramensis **
        +--+                                    
           |  +-------- Unionetta_fabagina__2   
           +--+                                 
              |  +----- Harmandia               
              +--+                              
                 |  +-- Unionetta_fabagina__1   
                 +--+                           
                    +-- Scabies                 

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1101117206--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Scabies                 
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Harmandia               

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
110122176--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
110181136--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Scabies                 
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Harmandia               

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia                
--+                                          
  |  +-------------- Indochinella            
  +--+                                       
     |  +----------- Radiatula               
     +--+                                    
        |  +-------- Scabiellus_songkramensis **
        +--+                                 
           |  +----- Harmandia               
           +--+                              
              |  +-- Unionetta               
              +--+                           
                 +-- Scabies                 
 
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