MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina
genus Songkhlanaia Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023: cladomics
3 Recent valid species (3 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 3 (3 total)
analyzed in 8 cladograms in 6 publications
maximum number of terminals in an analysis = 3 (mean = 1.75, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 3
maximum number of included terminals = 3, (mean = 3 median = 3)
proportion of cladograms that support basic monophyly = 0.67
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 582 | 651 | 4 | 629 | 37 | 9 | 0 |
+----- Amuranodonta
+-----------------------------------+
| | +-- Anemina
| +--+
| +-- Buldowskia
|
| +-- Beringiana
| +-----+
| | +-- Pletholophus
| +-----------------------------+
+-----------------------+ | | +----- Mimo
| | | +--+
| | | | +-- Acudonta
| | | +--+
| | | +-- Sinanodonta
| | |
| | | +----------------------------------- Cristaria
| | | |
| | | | +----------------------------- Lepidodesma
| +--+ | |
| | | | +----------- Lanceolaria
| | | | +--------------+
| | | +--+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | +--+
| | | | | | | +----- Acuticosta_chinensis__2
| | | | | | +--+
| | | | +--+ | +-- Acuticosta_chinensis__1
| | | | | +--+
| | | | | +-- Pseudoacuticosta_retiaria__1
| +--+ | |
| | | | +----------------------- Globunio
| | | | |
| | | +--+ +-------------------- Scriptolamprotula
| | | | |
| | | +--+ +----------------- Middendorffinaia
| | | | |
| | | +--+ +-------------- Unio
| | | | |
| | | | | +-- Schistodesmus
| | | +--+ +--------+
| | | | | +-- Nodularia
| | | | |
| | | +--+ +-- Tchangsinaia
| +--+ | +-----+
| | | | +-- Inversiunio
| | +--+
| | | +----- Pseudocuneopsis
| | +--+
| | | +-- Diaurora
| | +--+
| | +-- Cuneopsis
| |
| | +-------- Pseudodontoideus
| | +--------------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| | | +--+
| | | +-- Arcidens
| | |
| | | +----------------------- Anodonta_anatina
| | | |
+-----------------------+ | | +--+ +-------------------- Pseudanodonta
| | | | | | |
| | +--+ | +--+ +----------------- Brachyanodon_oregonensis__2
| | | | | |
| | | | | | +----- Anodonta_cygnea
| | | | +--+ +--------+
| | | | | | | +-- Anodonta_seddonae
| | | | | | +--+
| | | | +--+ +-- Anodonta_exulcerata
| | | | |
| | | | | +----------- Brachyanodon_impura
| | | | | |
| | | | +--+ +-------- Brachyanodon
| | | | | |
| | +--+ +--+ +----- Brachyanodon_nuttalliana__1
| | | | |
| | | +--+----- Brachyanodon_nuttalliana__2
| | | |
| | | | +-- Brachyanodon_californiensis
| | | +--+
| | | +-- Brachyanodon_oregonensis__1
| | |
| | | +-------------- Utterbackia
| | | +--+
| | | | | +----------- Pyganodon_grandis
| | | | +--+
| | | | | +-------- Pyganodon_lacustris
| | | | +--+
| | | | | +----- Pyganodon_cataracta__2
| | +--------+ +--+
| | | | +-- Pyganodon_cataracta__1
| | | +--+
| | | +-- Potamilus_fragilis__3
| | |
| | | +-------------- Utterbackiana
| | | |
| | +--+ +----------- Simpsonaias
| | | |
| | | | +-- Alasminota
| | +--+ +-----+
| | | | +-- Alasmidonta
| | | |
| | +--+ +-- Prolasmidonta
| | | +--+
| | | | +-- Platynaias
| | +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
| |
| | +-------------------------------------------------------------- Lamellidens
| | |
| | | +-- Balwantia
| +--+ +--------------------------------------------------------+
| | | +-- Trapezidens
| | |
| +--+ +-------------------------------------------------------- Leoparreysia
| | |
| | | +----------------------------------------------------- Pseudomulleria
| | | |
| +--+ | +-- Pseudoscabies
| | | +--------------+
| | | | +-- Indonaia
| | | +--------------------------------+
| | | | | +-------------- Indochinella
| +--+ | | |
| | | +--+ +----------- Radiatula
| | | | |
| | | +--+ +-- Songkhlanaia_songkhramensis__2 **
| | | | +-----+
| | | | | +-- Unionetta_fabagina__2
| | | +--+
| | | | +----- Harmandia
| +--+ +--+
| | | +-- Unionetta_fabagina__1
| | +--+
| | +-- Scabies
| |
| | +-- Assamnaia
| | +--------------------------------------------+
| | | +-- Parreysia
| | |
| | | +----- Nitia_monceti__2
| +--+ +--------------------------------------+
| | | | +-- Nitia_teretiuscula
| | | +--+
| | | +-- Nitia_monceti__1
| +--+
| | +----------------------------------------- Grandidieria
| | |
| | | +----- Pseudospatha_tanganyicensis__2
| | | +--------------------------------+
| +--+ | | +-- Moncetia
| | | +--+
| | | +-- Pseudospatha_tanganyicensis__1
| | |
| | | +----- Coelatura_leopoldvillensis__2
| +--+ +-----------------------------+
| | | | +-- Coelatura_leopoldvillensis__1
| | | +--+
| | | +-- Coelatura_mesafricana
| | |
| | | +-------------------------------- Coelatura_aegyptiaca__3
| | | |
| +--+ | +-------- Coelatura_hauttecoeuri__2
| | | |
| | | +-------- Nitia_acuminata
| | | |
| | | | +----- Coelatura_aegyptiaca__2
| | | +--------------------+ |
| | | | | +----- Coelatura_alluaudi__2
| +--+ | | |
| | | | +----- Coelatura_hauttecoeuri__1
| | | | |
| | | +--+----- Coelatura_stuhlmanni__2
--+ | | |
| | | | +-- Coelatura_aegyptiaca__1
| | | | |
| | | | +-- Coelatura_stuhlmanni__1
| | | | |
| | | +--+-- Coelatura_alluaudi__1
| +--+ |
| | +-- Coelatura_bakeri__1
| | |
| | +-- Coelatura_bakeri__2
| |
| | +----------- Coelatura_horei__3
| | |
| | +--------------+ +-------- Coelatura_ratidota
| | | | |
| | | +--+ +----- Coelatura__2
| | | | |
| | | +--+ +-- Coelatura_horei__1
| | | | |
| +--+ +--+-- Coelatura__1
| | |
| | +-- Coelatura_horei__2
| |
| | +----------------------- Coelatura_gabonensis
| | |
| +--+ +-------------------- Coelatura_stanleyvillensis
| | |
| | | +----------------- Coelatura_kunenensis
| +--+ |
| | | +-------------- Nyassunio
| | | |
| +--+ | +----------- Coelatura_hypsiprymna__2
| | | |
| | | +----------- Coelatura_mossambicensis__1
| | | |
| +--+ | +-- Coelatura_hypsiprymna__3
| | +--------+
| | | +-- Coelatura_mossambicensis__2
| | |
| | | +----- Coelatura_luapulaensis__2
| | | |
| | | +----- Coelatura_choziensis__2
| +--+-----+
| | +----- Coelatura_hypsiprymna__1
| | |
| | | +-- Coelatura_choziensis__1
| | +--+
| | +-- Coelatura_luapulaensis__1
| |
| | +----- Coelatura_briarti
| | +--+
| | | | +-- Prisodontopsis_aviculaeformis__1
| | | +--+
| +--+ +-- Prisodontopsis_mweruensis__1
| |
| | +-- Prisodontopsis_aviculaeformis__2
| | |
| +-----+-- Prisodontopsis_aviculaeformis__3
| |
| +-- Prisodontopsis_mweruensis__2
|
| +-------------------------- Khairuloconcha
| |
| | +----------------------- Ctenodesma
| | |
| +-----------------------------------------------------------+ | +-- Physunio_modelli
| | | | +-----------+
| | | | | +-- Physunio_pellucidus
| | | | |
| | +--+ +-----+ +-- Physunio_massini
| | | | | +--------+
| | | | | | +-- Physunio_superbus
| | | | +--+
| | | | | +-------- Physunio
| | | | +--+
| | | | | +----- Trapezoideus
| | +--+ +--+
| | | | +-- Yaukthwa
| | | +--+
| | | +-- Lens
| | |
| | | +----------------- Ensidens
| | | |
| | | | +----- Rectidens
| | +--+ +--------+
| | | | | +-- Hyriopsis_delaportei
| | | | +--+
| | | | +-- Hyriopsis_desowitzi
| | +--+
| | | +----------- Hyriopsis_panhai
| | | |
| | | | +-- Hyriopsis_bialata
| | +--+ +-----+
| | | | +-- Hyriopsis_khoratensis
| | | |
| | +--+ +-- Hyriopsis_kratiensis
| | | +--+
| | | | +-- Hyriopsis_phuphaniensis
| | +--+
| | | +-- Hyriopsis_myersiana
| | +--+
| | +-- Hyriopsis_sakhonensis
| |
| | +-------- Pseudunio
| | +--------------------------------------------------------------------------+
| | | | +----- Gibbosula
| | | +--+
| | | | +-- Cumberlandia
| | | +--+
| | | +-- Margaritifera
| | |
| | | +----------------------------- Indopseudodon
| | | |
| | | | +-- Lannanaia
| | | +--+ +-----------------------+
+--+ | | | | +-- Pilsbryoconcha
| | | | |
| | | | | +----- Songkhlanaia_tamodienica **
| | | +--+ +-----------------+
| | | | | | +-- Songkhlanaia_moreleti **
| | | | | +--+
| | | | | +-- Songkhlanaia_songkhramensis__1 **
| | | | |
| | | +--+ +-- Nyeinchanconcha
| | | | +-----------------+
| | | | | +-- Sundadontina
| | | | |
| | | | | +-- Isannaia
| | | +--+ +--+
| | | | | +-- Namkongnaia
| | | | +-----------+
| | | | | | +-- Thaiconcha
| | | | | +--+
| | | +--+ +-- Bineurus
| | | |
| | | | +-------------- Pseudodon_walpolei
| | | | |
| | | +--+ +----------- Pseudodon_vagulus
| | | | |
| | +-----------------------------------------------+ | | +----- Pseudodon_vondembuschianus
| | | | +--+ +--+
| | | | | | | +-- Pseudodon_mekongi
| | | | | | +--+
| | | | +--+ +-- Schepmania
| | | | |
| | | | | +----- Pseudodon_lenyanensis__2
| | | | +--+
| | | | | +-- Pseudodon_inoscularis
| | | | +--+
| | | | +-- Pseudodon_lenyanensis__1
| | | |
| | | | +-- Chamberlainia
| | | | +-----------------+
| | | | | +-- Sinohyriopsis
| | | | +--+
| | | | | | +----------------- Parvasolenaia
| | | | | | |
| | | | | +--+ +-------------- Pseudopostolata
| | | | | | |
| | | | | +--+ +----------- Koreosolenaia
| | | | | | |
| | | | | +--+ +-- Cosmopseudodon
| | | | | | +-----+
| | | | | | | +-- Guiunio
| | | | | +--+
+--+ | +--------+ | +----- Ptychorhynchus
| | | +--+
| | | | +-- Obovalis
| | | +--+
| | | +-- Postolata
| | |
| | | +-------- Caudiculatus
| | | +--------+
| | | | | +----- Lamprotula
| | | | +--+
| | | | | +-- Pronodularia
| | | | +--+
| | +-----+ +-- Potomida
| | |
| | | +-------------- Sinosolenaia_carinata
| | | |
| | | | +----- Gonidea
| | +--+ +-----+
| | | | | +-- Microcondylaea
| | | | +--+
| | +--+ +-- Leguminaia
| | |
| | | +-------- Inversidens
| | +--+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Amblema
| | +--------------------------------------------------------------------+
| | | | +-- Nephronaias
| | | +--+
| | | +-- Psoronaias
| | |
| | +--+ +----------------------------------------------------------------------- Reginaia
| | | | |
| | | | | +----- Popenaias
| | | | | +--------------------------------------------------------------+
| | | | | | | +-- Cyrtonaias
| | | +--+ | +--+
| | | | | +-- Glebula
| | | | |
| | | | | +----------------------------------------------------------------- Paetulunio
| | | | | |
| | | +--+ | +----------------- Toxolasma_parvum
| | | | | |
| | | | | | +----- Toxolasma_lividum__2
| | | | | | +--------+
| | | | | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus
| | | | | | | | +--+
| | | | | | | | +-- Toxolasma_lividum__1
| | | | | | | |
| | | +--+ | +--+ +-------- Toxolasma_texasiense__2
| | | | | | +--+
| | | | | | | | +----- Toxolasma_glans
| | | | | | | +--+
| | | | | | | | +-- Toxolasma_paulum
| | | | | +--+ +--+
| | | | | | +-- Toxolasma_texasiense__1
| | | | | |
+--+ | | | | +-------- Toxolasma__4
| | | | +--+
| | +--+ | +----- Toxolasma_granulatum
| | | +--+
| | | | +-- Toxolasma__2
| | | +--+
| | | +-- Toxolasma__3
| | |
| | | +----- Toxolasma_pullus
| | | +-----------------------------------------------------+
| | | | | +-- Toxolasma_germanum
| | | | +--+
| | | | +-- Toxolasma__1
| | | |
| | | | +-- Obliquaria
| | | | +-----------------------------------------------------+
| | | | | +-- Plectomerus
| | +--+ |
| | | | +-------- Potamilus_inflatus
| | | | +--------+
| | | | | | +----- Potamilus_ohiensis
| | | | | +--+
| | | | | | +-- Potamilus_amphichaenus
| | | | | +--+
| | | | +-----------------------------------+ +-- Potamilus_streckersoni
| | | | | |
| | +--+ | | +----- Potamilus_fragilis__2
| | | | | +--------+
| | | | | | | +-- Potamilus_leptodon
| | | | | | +--+
| | | | +--+ +-- Potamilus_fragilis__1
| | | | |
| | | | | +----------- Potamilus_metnecktayi
| | | | | |
| | | | +--+ +-------- Potamilus_alatus__3
| | | | | |
| | | | +--+ +----- Potamilus_alatus__2
| | +--+ | |
| | | +--+----- Potamilus_purpuratus__2
| | | |
| | | | +-- Potamilus_alatus__1
| | | +--+
| | | +-- Potamilus_purpuratus__1
| | |
| | | +-- Villosa_villosa
| | | +-----------------------------------------------+
| | | | +-- Atlanticoncha
| | | |
| | | | +-- Cambarunio_nebulosus
| | | | +--------------------------------------------+
| | +--+ | +-- Ligumia
| | | |
| | | | +-------- Leaunio_umbrans
| | | | +-----------------------------------+
| | | | | | +----- Cambarunio_taeniatus
| | +--+ | +--+
| | | | | +-- Cambarunio_iris
| | | | +--+
| | | | +-- Cambarunio_simus
| | | |
| | | | +----------------------------------------- Hamiota
| | | | |
| | +--+ | +----------------------- Pachynaias
| | | | |
| | | | +--------------+ +-------------------- Sagittunio
| | | | | | |
| | | | | | | +----- Ellipsaria
| | | | | +--+ +-----------+
| | | | | | | | +-- Potamilus_capax
| | | | | | | +--+
| | | | | | | +-- Truncilla
| | +--+ | +--+
| | | | | +----- Medionidus_acutissimus
| | | | | +--------+
| | | | | | | +-- Lemiox
| | | | | | +--+
| | | | +--+ +-- Medionidus_penicillatus
| | | | |
| | | | | +----------- Ptychobranchus
| | | | | |
| | | | +--+ +-- Cyprogenia_aberti__2
| | | | | +-----+
| | +--+ | | +-- Dromus
| | | +--+
| | | | +----- Cyprogenia_stegaria__2
| | | +--+
| | | | +-- Cyprogenia_aberti__1
| | | +--+
| | | +-- Cyprogenia_stegaria__1
+--+ |
| | +-- Villosa_delumbis
| | +--------------------------------+
| | | +-- Villosa_vibex
| | |
| | | +-------------- Obovaria_olivaria
| | | |
| | | +-----------------+ +-- Venustaconcha
| | | | | +--------+
| | | | | | +-- Epioblasma
| | | | +--+
| | | | | +-------- Obovaria_retusa
| +--+ | +--+
| | | | +----- Obovaria_choctawensis
| | | +--+
| | | | +-- Obovaria_unicolor
| | | +--+
| | | +-- Obovaria_subrotunda
| | |
| | | +-- Mesonaias
| | | +-----------------------+
| | | | +-- Obovaria_arkansasensis
| | | |
| +--+ +--+ +-- Lampsilis_splendida
| | | | +--------------------+
| | | | | +-- Lampsilis_bracteata
| | | | |
| | | +--+ +-- Lampsilis_fasciola
| | | | +-----------------+
| | | | | +-- Ortmanniana_pectorosa
| | | +--+
| | | | +----------------- Lampsilis_ornata
| | | | |
| | | +--+ +-------------- Lampsilis_ovata__2
| | | | |
| | | +--+ +----------- Lampsilis_dolabraeformis
| | | | |
| | | | | +----- Lampsilis_cardium__2
| | | +--+ +--+
| | | | | | +-- Lampsilis_cardium__1
| +--+ | | +--+
| | +--+ +-- Lampsilis_satura
| | |
| | | +----- Lampsilis_cariosa__2
| | +--+
| | | +-- Lampsilis_ovata__1
| | +--+
| | +-- Lampsilis_cariosa__1
| |
| | +----------- Ortmanniana_ligamentina__2
| | +--------------+
| | | | +-------- Lampsilis_abrupta
| | | +--+
| | | | +----- Lampsilis_higginsii__2
| | | +--+
| | | | +-- Ortmanniana_ligamentina__1
| | | +--+
| +--+ +-- Lampsilis_higginsii__1
| |
| | +----- Medionidus_conradicus
| | +-----------------+
| | | | +-- Leaunio_lienosus
| | | +--+
| | | +-- Leaunio_vanuxemensis
| | |
| +--+ +-- Lampsilis_floridensis
| | +-----------------+
| | | +-- Lampsilis_teres
| | |
| | | +-- Lampsilis_siliquoidea__1
| +--+ |
| | +-- Lampsilis_siliquoidea__2
| | +--------------+
| | | +-- Lampsilis_powellii__1
| | | |
| +--+ +-- Lampsilis_powellii__2
| |
| | +-------------- Lampsilis_sietmani
| | |
| +--+ +-- Lampsilis_straminea__2
| | +--------+
| | | +-- Lampsilis_virescens
| +--+
| | +-------- Lampsilis_hydiana__2
| +--+
| | +----- Lampsilis_hydiana__1
| +--+
| | +-- Lampsilis_bergmanni
| +--+
| +-- Lampsilis_straminea__1
|
| +-- Megalonaias
| +-----------------+
| | +-- Uniomerus
| |
| | +-- Theliderma_johnsoni
| | +--------+
| | | +-- Theliderma_metanevra
| | |
| +--------------------------------------------+ +-----+ +-- Quadrula_nobilis
| | | | | +-----+
| | | | | | +-- Quadrula_verrucosa
| | | | +--+
| | | | | +----- Theliderma_cylindrica
| | | | +--+
| | | | | +-- Quadrula_fragosa
| | +--+ +--+
| | | +-- Quadrula_quadrula
| | |
| | | +----- Pustulosa_kieneriana
| | | +--------+
| | | | | +-- Pustulosa_infucata
| | | | +--+
| | +--+ +-- Pustulosa_kleiniana
| | |
+-----------+ | +----------- Cyclonaias
| | |
| +--+ +-- Pustulosa_pustulosa
| | +-----+
| | | +-- Pustulosa_succissa
| +--+
| | +----- Pustulosa_nodulata
| +--+
| | +-- Pustulosa_necki
| +--+
| +-- Pustulosa_petrina
|
| +-------------------------------------------------------------- Elliptoideus
| |
| | +----------------------------------------------------------- Parvaspina
| | |
| | | +-- Elliptio_fisheriana
+--+ | +-----------------------------------------------------+
| | | +-- Elliptio_nasutidus
| | |
| | | +----------------------------------------------------- Elliptio_spinosa
| | | |
+--+ | | +----------------------------- Elliptio_dariensis__5
| | | |
| | | | +----- Elliptio_arctata__3
| | | | |
| | | | +----- Elliptio_crassidens__4
| | | | +--------------------+
| | | | | +----- Elliptio_mcmichaeli__2
| | | | | |
| | | | | | +-- Elliptio_crassidens__5
+--+ | | | +--+
| | +--------------------+ | +-- Elliptio_roanokensis
| | | | |
| | | | | +----- Elliptio_complanata__2
| | | | | |
| | | | | +----- Elliptio_pullata__2
| | | | | |
| | | | | +----- Elliptio_icterina__3
| | | | | |
| | | | +--------------------+ +-- Elliptio_dariensis__4
| | | | | +--+
| | | | | | +-- Elliptio_jayensis__2
| | | +--+ |
| | | | | +-- Elliptio_congaraea__2
+--+ | | | |
| | | | +-- Elliptio_crassidens__3
| | | +--+
| | | +-- Elliptio_fumata__2
| | | |
| | | +-- Elliptio_mcmichaeli__1
| | |
| | | +----------------------- Elliptio_crassidens__2
| | | |
| | | | +-- Elliptio_arca__2
| | | +--------------------+
| | | | +-- Elliptio_crassidens__1
| | | |
| | | | +-------- Elliptio_icterina__2
| | | | +-----------+
| | +--+ | | +----- Elliptio_dariensis__3
| | | | +--+
| | | | | +-- Elliptio_dariensis__2
| | | | +--+
| | | | +-- Elliptio_mediocris
| | | |
| | | | +-- Elliptio_folliculata__2
| | | | +-----------+
| | | | | +-- Elliptio_purpurella__2
| | +--+ |
| | | | +-- Elliptio_jayensis__1
+--+ | +--+-----------+
| | | | +-- Elliptio_pullata__1
| | | |
| | | | +----------- Elliptio_arca__1
| | | +--+
| | | | +-------- Elliptio_arctata__2
| | | +--+
| | | | +----- Elliptio_fumata__1
| | | +--+
| +--+ | +-- Elliptio_arctata__1
| | +--+
| | +-- Elliptio_purpurella__1
| |
| | +-- Elliptio_congaraea__1
| | |
| | +--------+-- Elliptio__1
| | | |
| | | +-- Elliptio__2
| | |
| +-----+ +-- Elliptio_complanata__1
| | +-----+
| | | +-- Elliptio_hopetonensis__1
| | |
| +--+ +----- Elliptio_dariensis__1
| | |
| | +----- Elliptio_producta
| +--+
| | +-- Elliptio_hopetonensis__2
| | |
| +--+-- Elliptio_hopetonensis__3
| |
| +-- Elliptio_icterina__1
|
| +----- Elliptio_shepardiana
| +-----------------------------------------+
| | | +-- Elliptio_angustata
| | +--+
| | +-- Elliptio_folliculata__1
| |
+--+ +-------------------------------------------- Eurynia
| |
| | +-- Hemistena
| | +--------------------------------------+
| | | +-- Plethobasus_cyphyus
+--+ |
| | +-------------------------------------- Elliptio_fraterna
| | |
| | | +-------------------------------- Pleurobema_plenum
| | | |
+--+ | +--+ +----------------------------- Pleurobema_stabile
| | | | |
| | | | | +-------------------------- Pleuronaia
| | | +--+ |
| | | | | +-- Pleurobema_decisum
| | | | | +-----+
| | | | | | +-- Pleurobema_hanleyianum
| | | +--+ +--------------+
+--+ | | | | +----- Pleurobema_pyriforme
| | | | +--+
| | | | | +-- Pleurobema_athearni
| | | | +--+
| | | | +-- Pleurobema_georgianum
| | +--+
| | | +-- Pleurobema_clava__1
| | | +--+
| | | | +-- Pleurobema_oviforme__1
| | | +--------------+
| | | | | +-- Pleurobema_clava__2
| | | | +--+
| | +--+ +-- Pleurobema_oviforme__2
+--+ |
| | +----------------- Pleurobema_beadleianum__2
| +--+
| | +-------------- Pleurobema_rubellum
| +--+
| | +----------- Pleurobema_strodeanum__2
| +--+
| | +-------- Pleurobema_beadleianum__1
| +--+
| | +----- Pleurobema_taitianum
| +--+
| | +-- Pleurobema_strodeanum__1
| +--+
| +-- Pleurobema_perovatum
|
| +----- Pleurobema_riddellii
| +-----------------+
| | | +-- Plethobasus_cooperianus
| | +--+
| | +-- Pleurobema_sintoxia
| |
+-----------+ +-------- Pleurobema_cordatum
| +-----------+
| | | +----- Fusconaia_subrotunda
| | +--+
| | | +-- Fusconaia_cor
| | +--+
+--+ +-- Fusconaia_cuneolus
|
| +----------------- Fusconaia_ozarkensis
| |
| | +----- Fusconaia_escambia__2
| | +--------+
+--+ | | +-- Fusconaia_burkei
| | +--+
| | +-- Fusconaia_escambia__1
| |
+--+ +----- Fusconaia_masoni
| |
| +-----+ +-- Fusconaia_cerina__1
| | | |
| | +--+-- Fusconaia_cerina__2
| | |
+--+ +-- Fusconaia_flava__2
|
| +-- Fusconaia_askewi
| +-----+
| | +-- Fusconaia_chunii
+--+
| +----- Fusconaia_flava__1
+--+
| +-- Fusconaia_iheringi
+--+
+-- Fusconaia_mitchelli
Konopleva et al., 2025 [Publ 4446], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 41 | 36 | 9 | 688 |
+-------------------- Lannanaia | --+ +----------------- Pilsbryoconcha | | | | +----- Songkhlanaia_tamodienica ** +--+ +--------+ | | | +-- Songkhlanaia_moreleti ** | | +--+ | | +-- Songkhlanaia_songkhramensis ** +--+ | +----- Nyeinchanconcha | +-----+ | | | +-- Thaiconcha | | +--+ +--+ +-- Bineurus | | +-------- Sundadontina +--+ | +----- Pseudodon +--+ | +-- Namkongnaia +--+ +-- Isannaia
Jeratthitikul et al., 2025 [Publ 4423], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 47 | 35 | 9 | 688 |
+-- Lannanaia
+--------------+
| +-- Pilsbryoconcha
|
| +----- Songkhlanaia_tamodienica **
--+ +--------+
| | | +-- Songkhlanaia_moreleti **
| | +--+
| | +-- Songkhlanaia_songkhramensis **
+--+
| +----- Nyeinchanconcha
| +-----+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +-------- Sundadontina
+--+
| +----- Isannaia
+--+
| +-- Namkongnaia
+--+
+-- Pseudodon
Bolotov et al., 2023 [Publ 4312], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 39 | 34 | 7 | -- |
+-------------- Songkhlanaia_tamodienica ** | | +-- Nyeinchanconcha --+ +--------+ | | +-- Sundadontina | | +--+ +-- Pseudodon | +-----+ | | +-- Pilsbryoconcha +--+ | +----- Namkongnaia +--+ | +-- Thaiconcha +--+ +-- Bineurus
Bolotov et al., 2023 [Publ 4312], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 32 | 7 | -- |
+-------------- Pilsbryoconcha | | +----- Nyeinchanconcha --+ +-----+ | | | +-- Thaiconcha | | +--+ +--+ +-- Bineurus | | +-------- Songkhlanaia_tamodienica ** +--+ | +----- Sundadontina +--+ | +-- Namkongnaia +--+ +-- Pseudodon
Bolotov et al., 2023 [Publ 4312], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 32 | 7 | -- |
+----------------- Pilsbryoconcha | --+ +-------------- Songkhlanaia_tamodienica ** | | +--+ +-- Namkongnaia | +--------+ | | +-- Pseudodon +--+ | +-------- Sundadontina +--+ | +----- Nyeinchanconcha +--+ | +-- Thaiconcha +--+ +-- Bineurus
Konopleva et al., 2023 [Publ 4262], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 81 | 24 | 7 | -- |
+-------------------- Pilsbryoconcha --+ | +----------------- Songkhlanaia_tamodienica ** +--+ | +-------------- Namkongnaia +--+ | +----------- Pseudodon +--+ | +-------- Sundadontina +--+ | +----- Nyeinchanconcha +--+ | +-- Thaiconcha +--+ +-- Bineurus
Jeratthitikul et al., 2024 [Publ 4387], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 58 | 35 | 9 | -- |
+-------------- Pseudodon | | +-------- Sundadontina | +--+ --+ | | +----- Nyeinchanconcha | | +--+ | | | +-- Thaiconcha | | +--+ +--+ +-- Bineurus | | +-- Isannaia | +-----+ | | +-- Pilsbryoconcha +--+ | +----- Namkongnaia +--+ | +-- Lannanaia +--+ +-- Songkhlanaia_tamodienica **
