MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Fri 13 February 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Dromus Simpson, 1900: cladomics
1 Recent valid species (1 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 1 (1 total)
analyzed in 17 cladograms in 10 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)
Cladograms
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- |
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus_dromas **
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus_dromas **
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Grobler et al., 2011 [Publ 3359], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 16 | 15 | 9 | -- |
+-------- Lemiox
+-----+
| | +----- Medionidus
| +--+
| | +-- Dromus_dromas **
| +--+
--+ +-- Cyprogenia
|
| +----------- Potamilus
| |
| | +----- Lampsilis_siliquoidea
| | +--+
+--+ | | +-- Lampsilis_higginsii
| | +--+
| | +-- Ortmanniana
| |
| | +----- Lampsilis_fasciola
+--+--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-- Cambarunio
| |
+-----+-- Ligumia
|
+-- Epioblasma
Serb, 2006 [Publ 2678], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- |
+-------------- Obliquaria | | +-- Ptychobranchus +-----------+ --+ +-- Lampsilis | | +-- Cyprogenia_aberti__6 | +--------+ | | +-- Dromus_dromas ** +--+ | +-------- Cyprogenia_aberti__5 | | +--+ +----- Cyprogenia_aberti__4 | | | | +-- Cyprogenia_stegaria__1 +--+ | | +-- Cyprogenia_stegaria__2 | | +--+-- Cyprogenia_aberti__1 | +-- Cyprogenia_aberti__2 | +-- Cyprogenia_aberti__3
Serb, 2006 [Publ 2678], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- |
+-------------------- Obliquaria | +-------------------- Lampsilis --+ | +----------------- Ptychobranchus | | +--+ +-- Cyprogenia_aberti__5 | +-----------+ | | +-- Dromus_dromas ** +--+ | +----------- Cyprogenia_aberti__4 | | +--+ +-------- Cyprogenia_aberti__3 | | +--+ +----- Cyprogenia_stegaria__2 | | +--+----- Cyprogenia_aberti__2 | | +-- Cyprogenia_stegaria__1 +--+ +-- Cyprogenia_aberti__1
Serb, 2006 [Publ 2678], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- |
+-------------------- Obliquaria | +-------------------- Lampsilis --+ | +----------------- Ptychobranchus | | +--+ +-- Cyprogenia_aberti__5 | +-----------+ | | +-- Dromus_dromas ** +--+ | +----------- Cyprogenia_aberti__4 | | +--+ +-------- Cyprogenia_aberti__3 | | +--+ +----- Cyprogenia_stegaria__2 | | +--+----- Cyprogenia_aberti__2 | | +-- Cyprogenia_stegaria__1 +--+ +-- Cyprogenia_aberti__1
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- |
+-------------------------- Lampsilis | --+ +----------------------- Obliquaria | | +--+ +-------------------- Ptychobranchus | | +--+ +-- Cyprogenia_aberti__6 | +--------------+ | | +-- Dromus_dromas ** +--+ | +-------------- Cyprogenia_aberti__4 | | +--+ +----------- Cyprogenia_aberti__3 | | +--+ +-------- Cyprogenia_aberti__5 | | +--+ +----- Cyprogenia_stegaria__2 | | +--+----- Cyprogenia_aberti__2 | | +-- Cyprogenia_stegaria__1 +--+ +-- Cyprogenia_aberti__1
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- |
+----------------- Obliquaria | | +----- Potamilus --+ +--------+ | | | +-- Dromus_dromas ** | | +--+ +--+ +-- Lemiox | | +----------- Lampsilis_ovata | | +--+ +-------- Lampsilis_virescens | | +--+ +-- Epioblasma | +--+ | | +-- Obovaria_arkansasensis +--+ | +-- Obovaria_olivaria +--+ +-- Venustaconcha
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- |
+----------------------- Obliquaria | +----------------------- Lampsilis --+ | +-------------------- Ptychobranchus | | +--+ +-- Cyprogenia_aberti__6 | +--------------+ | | +-- Dromus_dromas ** +--+ | +-------------- Cyprogenia_aberti__5 | | +--+ +----------- Cyprogenia_aberti__4 | | +--+ +-------- Cyprogenia_aberti__3 | | +--+ +----- Cyprogenia_stegaria__2 | | +--+----- Cyprogenia_aberti__2 | | +-- Cyprogenia_stegaria__1 +--+ +-- Cyprogenia_aberti__1
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus_dromas **
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- |
+----------------- Toxolasma | | +-------------- Potamilus | | | | +-- Ellipsaria | +-----------+ --+ | +-- Truncilla | | | | +----- Medionidus | +--------+ | | | +-- Lemiox | | +--+ +--+ +-- Dromus_dromas ** | | +----------- Cambarunio | | | +----------- Leaunio | | | +----------- Villosa | | | | +-- Sagittunio | +--------+ +--+ +-- Lampsilis_ovata | | +----- Venustaconcha +-----+ | | +-- Epioblasma | +--+ | +-- Obovaria | | +-------- Lampsilis_teres +--+ | +----- Lampsilis_siliquoidea +--+ | +-- Lampsilis_hydiana +--+ +-- Lampsilis_straminea
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- |
+-------------- Potamilus_alatus | +-------------- Potamilus_capax | +-------------- Glebula | | +-- Toxolasma +-----------+ --+ +-- Paetulunio | | +-- Dromus_dromas ** +-----------+ | +-- Lemiox | | +-- Lampsilis_ovata | +--------+ | | +-- Obliquaria +--+ | +-------- Lampsilis_virescens | | +--+ +-- Epioblasma | +--+ | | +-- Obovaria_arkansasensis +--+ | +-- Obovaria_olivaria +--+ +-- Venustaconcha
Roe, 2013 [Publ 3571], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 10 | 10 | 5 | -- |
+----------- Obliquaria | --+----------- Lampsilis | | +-------- Medionidus +--+ | +----- Ptychobranchus +--+ | +-- Cyprogenia +--+ +-- Dromus_dromas **
Serb et al., 2003 [Publ 2659], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- |
+----- Medionidus --+ | +-- Dromus_dromas ** +--+ +-- Cyprogenia
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 103 | 10 | 7 | -- |
-- Dromus_dromas **
Serb et al., 2003 [Publ 2659], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- |
+----- Medionidus --+ | +-- Dromus_dromas ** +--+ +-- Cyprogenia
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus_dromas **
+--+
+-- Cyprogenia
