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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Medionidus Simpson, 1900: cladomics
     6 Recent valid species (7 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (7 total)
analyzed in 19 cladograms in 12 publications
maximum number of terminals in an analysis = 2 (mean = 1.21, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 4
maximum number of included terminals = 2, (mean = 2 median = 2)
proportion of cladograms that support basic monophyly = 0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Zanatta & Murphy, 2006 [Publ 2672], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
26025446220
                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus **  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus ** 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
23025546223
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus **  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus ** 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
290210041224
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus **        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus **         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
260210041224
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus **         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus **        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
11012001311--
  +-------------- Potamilus_leptodon   
  |                                    
  |  +----------- Potamilus_alatus     
--+  |                                 
  |  |  +-------- Venustaconcha        
  |  |  |                              
  +--+  +-------- Leaunio              
     |  |                              
     |  |     +-- Ellipsaria           
     |  +-----+                        
     +--+     +-- Truncilla            
        |                              
        |     +-- Medionidus_conradicus **
        +-----+                        
        |     +-- Ptychobranchus       
        |                              
        |  +----- Cambarunio           
        +--+                           
           |  +-- Ortmanniana          
           +--+                        
              +-- Lampsilis            

Roe, 2013 [Publ 3571], fig. 4

iTtTbiriSoTroTroSiD
110110105--
  +----------- Obliquaria           
  |                                 
--+----------- Lampsilis            
  |                                 
  |  +-------- Medionidus_conradicus **
  +--+                              
     |  +----- Ptychobranchus       
     +--+                           
        |  +-- Cyprogenia           
        +--+                        
           +-- Dromus               

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
11012001311--
  +----- Ellipsaria           
  |                           
  +----- Potamilus_leptodon   
  |                           
  +----- Medionidus_conradicus **
  |                           
  +----- Potamilus_alatus     
  |                           
  +----- Ptychobranchus       
--+                           
  +----- Truncilla            
  |                           
  +----- Venustaconcha        
  |                           
  +----- Cambarunio           
  |                           
  +----- Leaunio              
  |                           
  |  +-- Ortmanniana          
  +--+                        
     +-- Lampsilis            

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
1101403519--
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus_conradicus **  
  |                 +--+                          
  |                 |  +-- Lemiox                 
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria             
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa                
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria               
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis              

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
11011981311--
  +-------------------- Potamilus_leptodon   
  |                                          
--+  +----------------- Potamilus_alatus     
  |  |                                       
  |  |              +-- Medionidus_conradicus **
  +--+  +-----------+                        
     |  |           +-- Ptychobranchus       
     |  |                                    
     +--+           +-- Ellipsaria           
        |  +--------+                        
        |  |        +-- Truncilla            
        +--+                                 
           |  +-------- Leaunio              
           |  |                              
           +--+     +-- Venustaconcha        
              |  +--+                        
              |  |  +-- Cambarunio           
              +--+                           
                 |  +-- Ortmanniana          
                 +--+                        
                    +-- Lampsilis            

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
11012865--
  +----- Medionidus_conradicus **
--+                           
  |  +-- Dromus               
  +--+                        
     +-- Cyprogenia           

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
110132107--
           +----- Ptychobranchus       
  +--------+                           
  |        |  +-- Potamilus_fragilis   
  |        +--+                        
  |           +-- Medionidus_conradicus **
--+                                    
  |           +-- Villosa              
  |  +--------+                        
  |  |        +-- Ellipsaria           
  +--+                                 
     |  +-------- Lampsilis_teres      
     +--+                              
        |  +----- Lampsilis_siliquoidea
        +--+                           
           |  +-- Potamilus_purpuratus 
           +--+                        
              +-- Potamilus_alatus     

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
11012686--
  +-------------- Potamilus_fragilis   
  |                                    
--+  +----------- Ptychobranchus       
  |  |                                 
  +--+  +-------- Medionidus_conradicus **
     |  |                              
     +--+  +----- Potamilus_purpuratus 
        |  |                           
        +--+----- Lampsilis            
           |                           
           |  +-- Ellipsaria           
           +--+                        
              +-- Villosa              

Burlakova et al., 2019 [Publ 3899], fig. 1

iTtTbiriSoTroTroSiD
1101322517--
  +----------------- Toxolasma             
  |                                        
  |  +-------------- Potamilus             
  |  |                                     
  |  |           +-- Ellipsaria            
  |  +-----------+                         
--+  |           +-- Truncilla             
  |  |                                     
  |  |        +----- Medionidus_acutissimus **
  |  +--------+                            
  |  |        |  +-- Lemiox                
  |  |        +--+                         
  +--+           +-- Dromus                
     |                                     
     |  +----------- Cambarunio            
     |  |                                  
     |  +----------- Leaunio               
     |  |                                  
     |  +----------- Villosa               
     |  |                                  
     |  |        +-- Sagittunio            
     |  +--------+                         
     +--+        +-- Lampsilis_ovata       
        |                                  
        |     +----- Venustaconcha         
        +-----+                            
        |     |  +-- Epioblasma            
        |     +--+                         
        |        +-- Obovaria              
        |                                  
        |  +-------- Lampsilis_teres       
        +--+                               
           |  +----- Lampsilis_siliquoidea 
           +--+                            
              |  +-- Lampsilis_hydiana     
              +--+                         
                 +-- Lampsilis_straminea   

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
11011382--
  +-------------------- Medionidus_conradicus **   
  |                                             
--+  +----------------- Lampsilis_teres         
  |  |                                          
  |  +----------------- Villosa                 
  +--+                                          
     |              +-- Lampsilis_satura        
     |  +-----------+                           
     |  |           +-- Lampsilis_cardium       
     +--+                                       
        |  +----------- Lampsilis_reeveiana     
        |  |                                    
        +--+  +-------- Lampsilis_hydiana__1    
           |  |                                 
           +--+-------- Lampsilis_hydiana__2    
              |                                 
              |  +----- Lampsilis_siliquoidea__2
              +--+                              
                 |  +-- Lampsilis_siliquoidea__1
                 +--+                           
                    +-- Lampsilis_powellii      

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013376--
  +----------------- Medionidus_conradicus **
--+                                       
  |  +-------------- Potamilus_fragilis   
  +--+                                    
     |  +----------- Ptychobranchus       
     +--+                                 
        |  +-------- Potamilus_purpuratus 
        +--+                              
           |  +----- Lampsilis            
           +--+                           
              |  +-- Villosa              
              +--+                        
                 +-- Ellipsaria           

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012786--
  +-------------------- Obliquaria           
--+                                          
  |  +----------------- Medionidus_conradicus **
  +--+                                       
     |  +-------------- Ptychobranchus       
     +--+                                    
        |  +----------- Potamilus_fragilis   
        +--+                                 
           |  +-------- Potamilus_purpuratus 
           +--+                              
              |  +----- Ellipsaria           
              +--+                           
                 |  +-- Villosa              
                 +--+                        
                    +-- Lampsilis            

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013376--
-- Medionidus_conradicus **

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
11012865--
  +----- Medionidus_conradicus **
--+                           
  |  +-- Dromus               
  +--+                        
     +-- Cyprogenia           

Grobler et al., 2011 [Publ 3359], fig. 2

iTtTbiriSoTroTroSiD
110116159--
        +-------- Lemiox               
  +-----+                              
  |     |  +----- Medionidus_conradicus **
  |     +--+                           
  |        |  +-- Dromus               
  |        +--+                        
--+           +-- Cyprogenia           
  |                                    
  |  +----------- Potamilus            
  |  |                                 
  |  |     +----- Lampsilis_siliquoidea
  |  |  +--+                           
  +--+  |  |  +-- Lampsilis_higginsii  
     |  |  +--+                        
     |  |     +-- Ortmanniana          
     |  |                              
     |  |  +----- Lampsilis_fasciola   
     +--+--+                           
        |  |  +-- Lampsilis_cardium    
        |  +--+                        
        |     +-- Lampsilis_ovata      
        |                              
        |     +-- Cambarunio           
        |     |                        
        +-----+-- Ligumia              
              |                        
              +-- Epioblasma           
 
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