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Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio tumidus Philipsson in Retzius, 1788: specimens
     61 available nominal species synonyms (89 total synonyms)

Recent Taxonomic Opinions

Unio tumidus — Graf (2007), Graf & Cummings (2007), Coffinet et al. (2008), Manendo et al. (2008), Chong et al. (2008), ... Dai et al. (2024), Wu et al. (2024), Hou et al. (2024), Dai et al. (2025), Chen et al. (2025)
Tumidiana tumida tumida — Kantor et al. (2010)
Unio tumidu [sic] — Li et al. (2010)
Tumidiana conus conus — Kantor et al. (2010)
Tumidiana muelleri — Kantor et al. (2010)
Tumidiana conus borysthenensis — Kantor et al. (2010)
Tumidiana tumida falcatula — Kantor et al. (2010)
Unio tumidus tumidus — Fauna Europaea (2012), Lopes-Lima et al. (2017), Bogatov (2022)
Unio tumidus borysthenensis — Fauna Europaea (2012)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio (s.s.) conus — Vinarski & Kantor (2016)
Unio (s.s.) muelleri — Vinarski & Kantor (2016)
Unio (s.s.) rostrata — Vinarski & Kantor (2016)
Unio tumidus zelebori — Lopes-Lima et al. (2017)
Unio tumidus borysthensis [sic] — Lopes-Lima et al. (2017)
Unio tumidus godetianus — Lopes-Lima et al. (2017)
Unio tumidus conus — Bogatov (2022)

Range

Widespread in Europe. Introduced to the Amur Basin.

Map of Georeferenced Specimens

INHS_3821.

INHS_3822.

INHS_21663. Lake Druesen, Schleswig- Holstein, Germany

INHS_21556. Pest, Budapest, Central Hungary, Hungary

INHS_22462. Tetezev River, Zhilomyz, Ukraine

INHS_21555. Uleser, Hungary

INHS_46937. Vantaanjoki, 2.5 km ESE Nurmijärvi, Myllykoski recreation area, off Siippoontie Rd. just E jct. Rt. E12/13/3, , Nyland, Finland

INHS_21431. Zijl River, near Leiden, South Holland, Netherlands

England

INHS_23642. New Bedford River, Sutton Gault, 11 km from Ely, Cambridgeshire, East of England, England

France

MNHN_22507. Elbeuf (Seine-Inf.)

MNHN_22301. La Charente, à Saintes

MNHN_22238. La Marne à Lagny (Seine-et-Marne)

MNHN_22092. La Rille, à Pont-Audemer (Eure)

MNHN_22506. la Seine à la Bouville [?]

MNHN_22504. la Seine près Rouen

MNHN_22239. la Seine, à Corbeil

MNHN_22505. la Seine, au nord du Rouen

MNHN_22240. le Grand-Morin, à Coulommiers

Netherlands

INHS_21432. De Diem, near Diemen (Amsterdam), North Holland, Netherlands

unknown

Georeferenced Genbank Vouchers

France

MNHN_IM-2009-12607. (16S: KC703377, 28S: KC703640, COI: KC703874, go to Genbank) France: Aisne River.

MNHN_IM-2009-12608. (16S: KC703378, COI: KC703875, go to Genbank) ditto.

MNHN_IM-2009-12609. (16S: KC703379, COI: KC703876, go to Genbank) ditto.

MNHN_IM-2009-12610. (16S: KC703380, COI: KC703877, go to Genbank) ditto.

MNHN_IM-2009-12658. (16S: KC703387, 28S: KC703646, COI: KC703881, go to Genbank) ditto.

MNHN_IM-2009-12659. (16S: KC703388, COI: KC703882, go to Genbank) ditto.

MNHN_IM-2009-12660. (16S: KC703389, COI: KC703883, go to Genbank) ditto.

MNHN_IM-2009-12661. (16S: KC703390, COI: KC703884, go to Genbank) ditto.

MNHN_IM-2009-12664. (16S: KC703391, COI: KC703885, go to Genbank) ditto.

MNHN_IM-2009-12665. (16S: KC703392, COI: KC703886, go to Genbank) ditto.

MNHN_IM-2009-12669. (16S: KC703396, COI: KC703887, go to Genbank) ditto.

MNHN_IM-2009-12670. (16S: KC703397, COI: KC703888, go to Genbank) ditto.

MNHN_IM-2009-12671. (16S: KC703398, COI: KC703889, go to Genbank) ditto.

MNHN_IM-2009-17800. (16S: KC703480, 28S: KC703725, COI: KC703907, go to Genbank) ditto.

MNHN_IM-2009-21209. (16S: KC703519, 28S: KC703767, COI: KC703924, go to Genbank) France: Canal Bruche Eckbolsheim.

MNHN_IM-2009-21210. (16S: KC703520, 28S: KC703768, COI: KC703925, go to Genbank) ditto.

MNHN_IM-2009-21211. (16S: KC703521, 28S: KC703769, go to Genbank) ditto.

MNHN_IM-2009-12575. (16S: KC703359, 28S: KC703627, COI: JX046560, go to Genbank) France: Meuse River.

MNHN_IM-2009-12578. (16S: KC703362, 28S: KC703629, COI: KC703871, go to Genbank) ditto.

MNHN_IM-2009-12620. (16S: KC703383, 28S: KC703642, go to Genbank) ditto.

MNHN_IM-2009-12683. (16S: KC703408, 28S: KC703653, COI: KC703893, go to Genbank) France: Oise River.

MNHN_IM-2009-12698. (16S: KC703415, 28S: KC703660, COI: KC703894, go to Genbank) France: Saône River.

MNHN_IM-2009-22393. (16S: KC703609, 28S: KC703858, COI: KC703957, go to Genbank) France: Saône River.

MNHN_IM-2009-12606. (16S: KC703376, 28S: KC703639, COI: KC703873, go to Genbank) France: Seine River.

Germany

SMF_354467. (16S: OK055689, 28S: OK077672, COI: OK050297, go to Genbank) Horloff, Oberflorstadt (50.3291, 8.871).

Kazakhstan

RMBH_biv_623_1. (COI: MK603937, go to Genbank) Kazakhstan: Cherny Irtysh River (48.0511°N, 85.0675° E), Ob’ Basin, Kara Sea drainage.

RMBH_biv_623_2. (COI: MK603938, go to Genbank) ditto.

RMBH_biv_623_3. (COI: MK603939, go to Genbank) ditto.

RMBH_biv_594_1. (COI: MK603931, go to Genbank) Kazakhstan: Ural River (50.7131°N, 51.3808 °E), Caspian Sea drainage.

RMBH_biv_594_2. (COI: MK603932, go to Genbank) ditto.

Russia

RMBH_biv_566_1. (COI: ON648758, go to Genbank) Russia: Aksai, Don Basin (47.3582, 40.0632).

RMBH_biv_584_3. (COI: MZ165238, go to Genbank) Russia: Bashkortostan Region, Belaya River.

RMBH_biv_609. (COI: MK603940, go to Genbank) Russia: Eastern Siberia, Transbaikalia, Ingoda River (52.0072°N, 113.5212°E), Amur Basin, Okhotsk Sea drainage.

RMBH_biv_568_1. (16S: MK574399, 28S: MK574540, COI: MK574263, go to Genbank) Russia: European part, Arkhangelsk Region, Lacha Lake (61.3079°N, 38.8429°E), Onega Basin, White Sea drainage.

RMBH_biv_568_2. (16S: MK574400, 28S: MK574541, COI: MK574264, go to Genbank) ditto.

RMBH_biv_278_1. (16S: MK574392, 28S: MK574533, COI: MK574253, go to Genbank) Russia: European part, Kursk Region, Seym River (51.7699°N, 36.2186°E), Dnieper Basin, Black Sea drainage.

RMBH_biv_278_2. (16S: MK574393, 28S: MK574534, COI: MK574254, go to Genbank) ditto.

RMBH_biv_278_3. (COI: MK574255, go to Genbank) ditto.

RMBH_biv_304_1. (16S: MK574394, 28S: MK574535, COI: MK574257, go to Genbank) Russia: European part, Perm Region, Iren River (57.4167°N, 56.9496°E), Volga Basin, Caspian Sea drainage.

RMBH_biv_304_6. (COI: MK574258, go to Genbank) ditto.

RMBH_biv_281_1. (16S: MK574395, 28S: MK574536, COI: MK574259, go to Genbank) Russia: European part, Saratov Region, Khopyor River (51.6330°N, 43.2296°E), Don Basin, Azov Sea drainage.

RMBH_biv_281_2. (16S: MK574396, 28S: MK574537, COI: MK574260, go to Genbank) ditto.

RMBH_biv_281_3. (16S: MK574397, COI: MK574261, go to Genbank) ditto.

RMBH_biv_281_4. (COI: MK574262, go to Genbank) ditto.

RMBH_biv_280_7. (COI: MK574256, go to Genbank) Russia: European part, Voronezh Region, Khopyor River (51.4878°N, 42.5931°E), Don Basin, Azov Sea drainage.

RMBH_biv_718_6. (COI: MZ165243, go to Genbank) Russia: Kaluga Region, Oka River.

RMBH_biv_556_1. (COI: ON648756, go to Genbank) Russia: Kirpili (45.3540, 39.8053).

RMBH_biv_725_1. (COI: MZ165244, go to Genbank) Russia: Krasnodar Region, Beysuzhek Levyi River.

RMBH_biv_725_4. (COI: MZ165247, go to Genbank) ditto.

RMBH_biv_556_3. (COI: MZ165232, go to Genbank) Russia: Krasnodar Region, Kirpili River.

Mp_3904_U10. (16S: MH544953, 5.8S: MH544849, ITS1: MH544849, go to Genbank) Russia: River Ivitza, Volga basin, Rameshkovsky District, Tver Oblast.

Mp_3904_U12. (16S: MH544960, ITS1: MH544858, go to Genbank) ditto.

Mp_3904_U42. (16S: MH544956, COI: MH539803, go to Genbank) ditto.

Mp_3904_U44. (16S: MH544955, 5.8S: MH544850, COI: MH539802, ITS1: MH544850, go to Genbank) ditto.

Mp_3904_U52. (16S: MH544954, go to Genbank) ditto.

Mp_3904_U67. (16S: MH544963, ITS1: MH544855, go to Genbank) ditto.

Mp_3904_U78. (16S: MH544964, 5.8S: MH544854, ITS1: MH544854, go to Genbank) ditto.

Mp_3904_U83. (16S: MH544965, 5.8S: MH544853, ITS1: MH544853, go to Genbank) ditto.

RMBH_biv_566_2. (COI: MZ165235, go to Genbank) Russia: Rostov Region, Aksai River.

RMBH_biv_566_3. (COI: MZ165236, go to Genbank) ditto.

RMBH_biv_766_1. (COI: MZ165256, go to Genbank) Russia: Voronezh Region, Don River.

RMBH_biv_766_2. (COI: MZ165257, go to Genbank) ditto.

RMBH_biv_766_3. (COI: MZ165258, go to Genbank) ditto.

RMBH_biv_774_1. (COI: MZ165263, go to Genbank) Russia: Voronezh Region, Khoper River.

RMBH_biv_774_2. (COI: MZ165264, go to Genbank) ditto.

RMBH_biv_774_4. (COI: MZ165266, go to Genbank) ditto.

Sweden

GNM-MOLOV000009576. (COI: PP204156, go to Genbank) Sweden (58.572278 N 13.942361 E).

GNM-MOLOV000009577. (COI: PP204157, go to Genbank) ditto.

Switzerland

NMBE_565032. (16S: OK055609, 28S: OK077595, COI: OK050213, go to Genbank) Aare, Häftli (47.14706, 7.37087).

NMBE_565033. (16S: OK055610, 28S: OK077596, COI: OK050214, go to Genbank) ditto.

NMBE_566652. (16S: OK055681, 28S: OK077665, COI: OK050290, go to Genbank) Alpnachersee, kleine Schlieren (46.9554, 8.2848).

NMBE_566653. (16S: OK055683, 28S: OK077666, COI: OK050291, go to Genbank) ditto.

NMBE_566654. (16S: OK055684, 28S: OK077667, COI: OK050292, go to Genbank) ditto.

NMBE_559886. (16S: OK055602, 28S: OK077587, COI: OK050205, go to Genbank) Greifensee, Bad (47.3669, 8.6708).

NMBE_566655. (16S: OK055685, 28S: OK077668, COI: OK050293, go to Genbank) Klingnauer Stausee (47.5829, 8.2343).

NMBE_566656. (16S: OK055686, 28S: OK077669, COI: OK050294, go to Genbank) ditto.

NMBE_566657. (16S: OK055687, 28S: OK077670, COI: OK050295, go to Genbank) ditto.

NMBE_559866. (16S: OK055586, 28S: OK077570, COI: OK050188, go to Genbank) Lac de Morat, Muntelier (46.93769, 7.12439).

NMBE_559867. (16S: OK055587, 28S: OK077571, COI: OK050189, go to Genbank) ditto.

NMBE_559872. (16S: OK055590, 28S: OK077575, COI: OK050193, go to Genbank) Lac Morat, Plage d'Avenches (46.9053, 7.0559).

NMBE_559873. (16S: OK055591, 28S: OK077576, COI: OK050194, go to Genbank) ditto.

NMBE_559880. (16S: OK055596, 28S: OK077581, COI: OK050199, go to Genbank) Lac Neuchâtel, St-Blaise (47.0109, 6.9806).

NMBE_565048. (16S: OK055623, 28S: OK077609, COI: OK050224, go to Genbank) Léman, Vieux Rhône (46.39471, 6.8733).

NMBE_565049. (16S: OK055624, 28S: OK077610, COI: OK050225, go to Genbank) Léman, Villeneuve port (46.39677, 6.92102).

NMBE_565050. (16S: OK055625, 28S: OK077611, COI: OK050226, go to Genbank) Léman, Villeneuve port (46.39686, 6.92097).

NMBE_565007. (16S: OK055607, 28S: OK077592, COI: OK050210, go to Genbank) Vierwaldstättersee, Horw (47.00701, 8.31999).

NMBE_565076. (16S: OK055648, 28S: OK077633, COI: OK050247, go to Genbank) ditto.

Turkey

GCMAE_M34_02. (16S: MZ518959, 28S: MZ519196, COI: MZ509536, go to Genbank) Turkey: Çamlıca: Maritsa Basin (40.768715, 26.514664).

GCMAE_M34_03. (16S: MZ518960, 28S: MZ519197, COI: MZ509537, go to Genbank) ditto.

GCMAE_M34_04. (COI: MZ509538, go to Genbank) ditto.

NCSM_107595_1. (16S: MZ518958, 28S: MZ519195, COI: MZ509535, go to Genbank) ditto.

NCSM_107595_2. (COI: MZ509539, go to Genbank) ditto.

Ukraine

MNCN_N1724. (16S: KX400553, go to Genbank) Gnilopyat River, Ukraine.

MNCN_N1735. (16S: KX400556, COI: KX400164, go to Genbank) ditto.

ZSU-U9. (COI: KY930357, go to Genbank) South Bug, Ukraine.

ZSU-U7. (COI: KY930358, go to Genbank) Vorsakla River, Ukraine.