species Unio tumidus Philipsson in Retzius, 1788: synonymy

Unio tumidus Philipsson in Retzius, 1788

Philipsson in Retzius, 1788. Diss. Nat.-Hist. Test. Gen.: 17.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (Groupe Tumidusiana) tumidus — Caziot (1907)
Unio tumidus — Germain (1908), Kobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), ... Nagel (2025), Campbell & Lydeard (2026), Wu et al. (2026), Wu et al. (2026), Li et al. (2026)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio (s.s.) tumidus — Zhadin (1938), Vinarski & Kantor (2016)
Unio tumidus tumidus — Haas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017), Bogatov (2022)
Unio (Tumidusiana) tumidus — Zatravkin (1983), Korniushin (1998), Gozhik (2006)
Unio (s.s.) tumidus tumidus — Falkner et al. (2001), Falkner et al. (2002)
Tumidiana tumida — Starobogatov et al. (2004)
Tumidiana tumida tumida — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio tumidu [sic] — Li et al. (2010)

Unio conus Spengler, 1793

Lister, 1685. Conch.: pl. 146, fig. 1. [pre-linnaean]
Spengler, 1793. Skr. Naturhist. Selsk. 3: 60, no. 26.

Unio (sec. Lymnium) pictorum — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) pictorum — Simpson (1914)
Unio tumidus — Germain (1931), Graf (2007), IUCN (2011), Klishko et al. (2017), Bogatov et al. (2018), Anisimova et al. (2018)
Unio (s.s.) tumidus var. conus — Zhadin (1938)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (Tumidusiana) conus — Zatravkin (1983), Gozhik (2006)
Tumidiana conus — Starobogatov et al. (2004)
Tumidiana conus conus — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio (s.s.) conus — Vinarski & Kantor (2016)
Unio tumidus conus — Bogatov (2022)

Mya depressa Donovan, 1801

Donovan, 1801. Brit. Shells 3: pl. 101.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio tumidus tumidus — Haas (1940), Lopes-Lima et al. (2017)
Unio (s.s.) tumidus depressus — Falkner et al. (2001), Falkner et al. (2002)
Unio tumidus — Prié & Puillandre (2014)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Mya ovata Donovan, 1802

Donovan, 1802. Brit. Shells 4: pl. 122, figs. 1-3.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio tumidus — Germain (1908), Germain (1931)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio tumidus tumidus — Haas (1940), Haas (1969), Fauna Europaea (2012)

Mya ovalis Montagu, 1803

(misidentified reference to Mya ovalis Lightfoot, 1786)
Philipsson in Retzius, 1788. Diss. Nat.-Hist. Test. Gen.: 17.
Pulteney, 1799. Cat. Birds. Shells. Dorsetshire: 27. [nomen nudum]
Montagu, 1803. Test. Brit. 1-2: 34.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Tumidiana conus conus — Kantor & Sysoev (2005)
Unio tumidus — Klishko et al. (2017)

Mysca solida Turton, 1822

Montagu, 1803. Test. Brit. 1-2: 36 [as ‘Mya solida’].
Turton, 1822. Conch. Ins. Brit.: 246, pl. 16, fig. 2.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio inflata Hecart, 1833

(homonym of Unio inflatus Studer, 1820)
Hecart, 1833. Mém. Soc. Agricult. Valanciennes 1: 245.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio muelleri Rossmässler, 1838

Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): 41, pl. 40, fig. 541.
Rossmässler, 1842. Icon. Land- und Süssw.-Moll. 2 (5): 13, pl. 54, fig. 739.
LECTOTYPE SMF 10815 (by Zilch, 1967: 82), Schulensee, near Kiel, Germany.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) muelleri — Zatravkin (1983), Stadnichenko (1984), Vinarski & Kantor (2016)
Tumidiana muelleri — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Unio tumidus — Graf (2007), IUCN (2011), Klishko et al. (2017), Nagel (2025)

Unio tumidus var. lacustris Rossmässler, 1838

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): pl. 40, fig. 542 [as ‘Unio tumidus’].
Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): pl. 60, fig. 775 [as ‘Unio tumidus’].
Küster, 1853. Conchyl. Cab. 9(2): 75, pl. 18, fig. [as ‘Unio tumidus’].

Unio tumidus var. radiatus Colbeau, 1839

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Colbeau, 1839. Matér. Faune Malacol. Belg.: ??.

Unio pictorum Brown, 1844

(misidentified reference to Mya pictorum Linnaeus, 1758)
Brown, 1844. Ill. Rec. Conch. Gr. Brit. & Ireland: 81, pl. 31, figs. 8-11.

Unio (sec. Lymnium) tumidus — Simpson (1900)
Unio (sec. Lymnium) (Group U. pictorum) tumidus — Simpson (1914)

Unio tumidus var. corrosus Zelebor, 1851

(homonym of Unio corrosus Villa, 1841; intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zelebor, 1851. Ber. Mit. Freunde Naturw. Wien 7: 230, sp. 7.
SYNTYPES SMF 10812, March bei Thebensee und Neudorf, Slovakia.

Unio tumidus tumidus — Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio tumidus var. solidus Zelebor, 1851

(homonym of Unio solidus Lea, 1838; intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zelebor, 1851. Ber. Mit. Freunde Naturw. Wien 7: 230, sp. 6.

Unio tumidus tumidus — Haas (1969)

Unio limaniae Dupuy, 1852

Bouill., 1836. Cat. Moll. Auv.: 81. [nomen nudum]
‘Bouillet’ Dupuy, 1852. Hist. Natur. Moll. France (6): 653 [as ‘limaniæ’].

Unio tumidus var. saccatus Rossmässler, 1859

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Rossmässler, 1859. Icon. Land- und Süssw.-Moll. 3 (5-6): 138, pl. 89, fig. 969.
LECTOTYPE SMF 10802 (by Zilch, 1967: 82), Köflefors-Å”, Sweden.

Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (Tumidusiana) tumidus var. saccatus — Stadnichenko (1984)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio pictorum var. niger de Joannis, 1859

(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; homonym of Unio (Elliptio) nigra Rafinesque, 1820)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 294, pl. 12, fig. 2.

Unio (Rostratiana) niger — Germain (1904)
Unio tumidus — Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio pictorum var. vinceleus de Joannis, 1859

(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; renamed Unio vinceloti Germain, 1904)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 294, pl. 12, figs. 4-5.

Unio (Vincelotiana) vinceloti — Germain (1904)
Unio (Groupe des Vincelensiana) [sic] vincelens [sic] — Caziot (1907)
Unio tumidus — Germain (1931), Prié & Puillandre (2014)
Unio pictorum rostratus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus vinceleus — Falkner et al. (2001), Falkner et al. (2002)
Unio (s.s.) rostrata — Vinarski & Kantor (2016)
Unio tumidus tumidus — Lopes-Lima et al. (2017)

Unio tumidus picta Beck in Mørch, 1863

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788; homonym of Unio pictus Lea, 1834)
Beck in Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 339.
Mørch, 1864. Syn. Moll. Daniae: 77.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio pictorum maxima Mørch, 1863

(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 340.
Mørch, 1864. Syn. Moll. Daniae: 78.

Unio tumidus var. maximus — Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus limicola Mørch, 1863

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 339.
Mørch, 1864. Syn. Moll. Daniae: 77.

Unio (s.s.) tumidus limicola — Zhadin (1938)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (Tumidusiana) tumidus var. limicola — Stadnichenko (1984)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio conus compressa Mørch, 1863

(homonym of Unio compressus Rafinesque, 1831; intro'd as intra-sp. of Unio conus Spengler, 1793)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 340.

Unio conus compressissima Mørch, 1863

(homonym of Unio compressissimus Lea, 1845; intro'd as intra-sp. of Unio conus Spengler, 1793)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 340.

Unio robianoi Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 33, pl. 1, fig. 1-2.

Unio robianoi var. inflatus Malzine, 1867

(intro'd as intra-sp. of Unio robianoi Malzine, 1867; homonym of Unio inflatus Studer, 1820)
Malzine, 1867. Essai faun. malac. Belg.: 33, pl. 1, fig. 3-4.

Unio batavus var. incurvatus Colbeau, 1868

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 2.

Unio (Alpecanusiana) incurvatus — Germain (1904)
Unio tumidus — Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. heckingi Colbeau, 1868

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 1.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. rohrmanni Kobelt, 1879

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Kobelt, 1879. Icon. Land- und Süssw.-Moll. 7: 32, pl. 192, fig. 1951.
LECTOTYPE SMF 10826 (by Zilch, 1967: 82), Świerzyna, Poland.

Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio tumidus var. borysthenensis Kobelt, 1879

(renamed Unio tumidus var. borysthenicus Servain, 1888; intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Kobelt, 1879. Icon. Land- und Süssw.-Moll. 7: 32, pl. 192, fig. 1950.
LECTOTYPE SMF 10820 (by Zilch, 1967: 83), Dnieper River, Dnipro, Ukraine.

Unio tumidus borysthenensis — Haas (1940), Haas (1969), Fauna Europaea (2012)
Tumidiana conus borysthenensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio (s.s.) conus — Vinarski & Kantor (2016)
Unio tumidus borysthensis [sic] — Lopes-Lima et al. (2017)
Unio tumidus — Araujo et al. (2018), Nagel (2025)

Unio gerstfeldtianus Clessin, 1880

Clessin, 1880. Nachr. Bl. Dtsch. Malak. Ges. 12: 81.

Unio (s.s.) tumidus var. gerstfeldtianus — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969), Fauna Europaea (2012)
Unio tumidus var. gerstfeldtianus — Zhadin (1965)
Tumidiana tumida tumida — Kantor & Sysoev (2005)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio bardus Bourguignat, in sched. Servain, 1881

Bourguignat, in sched. Servain, 1881. Malac. Lac Balaton: 98.
Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 323.

Unio (Rostratiana) bardus — Germain (1904)
Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio tumidus var. falcatulus Drouët, 1881

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Drouët, 1881. Unionidae Russie: 9.

Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Unio (Tumidusiana) tumidus falcatulus — Stadnichenko (1984)
Tumidiana tumida falcatulus — Kantor & Sysoev (2005)
Tumidiana tumida falcatula — Kantor et al. (2010)
Unio tumidus — IUCN (2011)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. minor Drouët, 1881

(homonym of Unio margaritifer var. minor Rossmässler, 1835)
Drouët, 1881. Unionidae Russie: 9.

Unio edyus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 299, 367.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 177 [71].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 215.

Unio tumidus — Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio alpecanus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 177 [71].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 215.
LECTOTYPE MNHN_22094 (by Graf, 2011: 457), l’Oise à Creil.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio fourneli Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 300, 367.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 177 [71].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 214.
IDIOTYPE MNHN_22225, La Marne à Lagny (Seine-et-Marne).

Unio tumidus — Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio schroederi Bourguignat in Schröder, 1885

Bourguignat in Schröder, 1885. Bull. Soc. Malac. France 2: 224.

Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio rhenanus Kobelt, 1886

Kobelt, 1886. Jb. Nassau. Ver. Naturk. 39: 96, pl. 5, fig. 3.
Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: 48, pl. 56, fig. 297.
HOLOTYPE SMF 10821, Rheingau Region, Germany.

Unio tumidus rhenanus — Kobelt & Haas (1910)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio tumidus var. godetiana Clessin, 1887

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Clessin, 1887. Moll.-Fauna Mitteleuropas 2: 738, text fig. 496.

Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus godetianus — Falkner et al. (2001)
Unio tumidus godetianus — Lopes-Lima et al. (2017)

Unio mulierum Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 321.

Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio tumidus var. borysthenicus Servain, 1888

(new name for Unio tumidus var. borysthenensis Kobelt, 1879)
Kobelt, 1879. Icon. Land- und Süssw.-Moll. 7: 32, pl. 192, fig. 1950 [as ‘borysthenensis’].
‘Kobelt’ Servain, 1888. Bull. Soc. Malac. France 5: 323.

Unio (s.s.) tumidus var. borysthenicus — Zhadin (1938)
Unio tumidus var. borysthenicus — Zhadin (1965)
Unio (Tumidusiana) conus borysthenicus — Stadnichenko (1984)

Unio spengeli Bourguignat in Servain, 1888

Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 323.

Unio anabaenus Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 324 [as ‘anabænus’].

Unio macrorhynchus Borcherding, 1889

(homonym of Unio macrorhynchus Bourguignat in Locard, 1882; renamed Unio borcherdingi Bourguignat in Locard, 1889)
Borcherding, 1889. Abh. Naturw. Ver. Bremen 10: 342, pl. 4, figs. 3-3b.
LECTOTYPE SMF_XXXX (by Zilch, 1967: 81), Dümmer See, Germany.

Unio tumidus tumidus — Haas (1969)
Unio tumidus — Nagel (2025)

Unio borcherdingi Bourguignat in Locard, 1889

(new name for Unio macrorhynchus Borcherding, 1889)
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 153 [47].

Unio tumidus tumidus — Haas (1969)
Unio tumidus — Nagel (2025)

Unio tumidulus Locard, 1889

(homonym of Unio tumidulus Lea, 1856)
Rossmässler, 1836. Icon. Land- und Süssw.-Moll. 1 (3): pl. 14, fig. 203 [as ‘Unio tumidus’].
Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): pl. 40, fig. 542 [as ‘Unio tumidus’].
Locard, 1889. Contr. Faune Malacol. Franc. 13: 176 [70], 261 [155].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 213.
LECTOTYPE MNHN_22586 (by Graf, 2011: 510), La Seine à Rouen (Seine-Inférieure).

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio aldemaricus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 176 [70], 262 [156].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 213.
LECTOTYPE MNHN_22091 (by Graf, 2011: 456), l’Aisne, à Guignicourt (Ardennes).

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio giberti Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 131 [25], 191 [85].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 215, fig. 227.
LECTOTYPE MNHN_22237 (by Graf, 2011: 477), La Marne à Lagny (Seine-et-Marne).

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio rothomagensis Locard, 1893

Locard, 1893. Coqu. Eau Douce Saumâtre France: 213.
LECTOTYPE MNHN_22503 (by Graf, 2011: 501), la Seine près Rouen.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio elboviensis Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 4.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 215.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio lacrymiformis Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 12.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 216, fig. 228.
LECTOTYPE MNHN_22300 (by Graf, 2011: 483), La Seine au nord de Rouen.

Unio tumidus — Germain (1908), Germain (1931)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio levoiturieri Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 7.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 216.

Unio tumidus — Germain (1908)
Unio tumidus tumidus — Haas (1940), Haas (1969)

Unio catinulus Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 10.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 217.
SYNTYPE MNHN_22172, La Seine à Elbeuf (Seine-Inférieure).

Unio tumidus — Germain (1908), Germain (1931)

Unio vinceloti Germain, 1904

(new name for Unio pictorum var. vinceleus de Joannis, 1859)
‘de Joannis’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 276 [38].

Unio (Vincelotiana) vinceloti — Germain (1904)

Unio vinceloti var. cuneiformis Germain, 1904

(intro'd as intra-sp. of Unio vinceloti Germain, 1904)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 276 [38], pl. 2, fig. 37.

Unio (Vincelotiana) vinceloti cuneiformis — Germain (1904)

Unio lauterborni Haas, 1909

Haas, 1909. Beitr. Kenntn. Mittleleurop. Najadeen, Beilage zu... 41: 46.
LECTOTYPE SMF 10819 (by Zilch, 1967: 82), Altrein, at Neuhofen, near Ludwigshafen, Germany.

Unio tumidus lauterborni — Kobelt & Haas (1910)
Unio tumidus tumidus — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio tumidus var. okae Kobelt & Haas, 1910

(renamed Unio tumidus var. kobeltianus Zhadin, 1938; homonym of Unio crassus var. okae Kobelt & Haas, 1910; intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Kobelt & Haas, 1910. Icon. Land- und Süssw.-Moll. 17: 58, index, pl. 480, figs. 2565-2566.

Unio (s.s.) tumidus var. kobeltianus — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1969)
Unio tumidus — Nagel (2025)

Unio tumidus var. succinctus Hilbert, 1912

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Hilbert, 1912. Schr. Königsberg. Phys. Ges. 52: 271.

Unio tumidus tumidus — Haas (1969)

Unio tumidus ilekensis Kobelt, 1912

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 28, pl. 492, fig. 2629.
HOLOTYPE SMF 11055, Ilek River, Bestamak, Kazakhstan.

Unio (s.s.) tumidus var. ilikensis [sic] — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Tumidiana tumida tumida — Kantor & Sysoev (2005)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)
Unio tumidus — Nagel (2025)

Unio tumidus var. natermanni Honigmann, 1914

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Honigmann, 1914. Abh. Ber. Mus. Magdeburg 2: 7.
SYNTYPE SMF 654, Neajlov River, Comana, Romania.

Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Unio tumidus — Nagel (2025)

Unio tumidus var. bashkiricus Zhadin, 1938

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zhadin, 1938. Faune USSR Mollusques 4(1): 81, text fig. 19.

Unio (s.s.) tumidus var. bashkiricus — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Unio tumidus var. bashkiricus — Zhadin (1965)
Tumidiana tumida tumida — Kantor & Sysoev (2005)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. moltshanovi Zhadin, 1938

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zhadin, 1938. Faune USSR Mollusques 4(1): 85, text fig. 20.

Unio (s.s.) tumidus var. moltshanovi — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. fridmani Zhadin, 1938

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zhadin, 1938. Faune USSR Mollusques 4(1): 86, text fig. 21.

Unio (s.s.) tumidus var. fridmani — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Unio tumidus var. fridmani — Zhadin (1965)
Tumidiana tumida tumida — Kantor & Sysoev (2005)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. kobeltianus Zhadin, 1938

(homonym of Unio batavus kobeltianus Haas, 1913; new name for Unio tumidus var. okae Kobelt & Haas, 1910; intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Zhadin, 1938. Faune USSR Mollusques 4(1): 82.

Unio (s.s.) tumidus var. kobeltianus — Zhadin (1938)
Unio tumidus borysthenensis — Haas (1940), Haas (1969)
Tumidiana tumida tumida — Kantor & Sysoev (2005)
Unio (Tumidusiana) kobeltianus — Gozhik (2006)
Unio (s.s.) tumidus — Vinarski & Kantor (2016)

Unio tumidus var. ilikensis ‘Kobelt’ of Authors

(error for Unio tumidus ilekensis Kobelt, 1912)
‘Kobelt’ of Authors.
Zhadin, 1938. Faune USSR Mollusques 4(1): 83.
Kantor & Sysoev, 2005. Cat. Moll. Russia: 327.

Unio limagnae ‘Bouillet’ of Authors

(error for Unio limaniae Dupuy, 1852)
‘Bouillet’ of Authors.
Jay, 1835. Cat. Rec. Shells: 16 [as ‘limagnæ’].
Lea, 1836. Synopsis Fam. Naiades: 36 [as ‘limagnæ’].
Lea, 1838. Synopsis Fam. Naiades: 24 [as ‘limagnæ’].

Unio concavus ‘Zeleb.’ of Authors

‘Zeleb.’ of Authors.

Unio elongatus ‘Rm.’ of Authors

‘Rm.’ of Authors.

Unio muelleri var. compressus Westerl. Faun. Moll. Suec. of Authors

Westerl. Faun. Moll. Suec. of Authors.

Mytilus angustior Lister, 1685

Lister, 1685. Conch.: ??. [pre-linnaean]

Unio rostrata Waardenburg, 1827

(misidentified reference to Unio rostrata Lamarck, 1819)
Waardenburg, 1827. Moll. Belgico: 36.

Unio tumidus — Germain (1908), Germain (1931)

Unio tumidus Rossmässler, 1836

(reference to Unio tumidus Philipsson in Retzius, 1788)
Rossmässler, 1836. Icon. Land- und Süssw.-Moll. 1 (3): pl. 14, fig. 203.
Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): 41, pl. 40, fig. 542.
Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): pl. 60, fig. 773.
Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 28, fig. 20.
Turton, 1857. Man. Land Fresh-Water Shells Brit. Is.: pl. 2, fig. 13.
Forbes & Hanley, 1859. Brit. Moll.: pl. 40, fig. 1.

Unio tumidus — Germain (1908), Germain (1931)

Unio batavus Rossmässler, 1836

(misidentified reference to Mya batava Maton & Rackett, 1807)
Rossmässler, 1836. Icon. Land- und Süssw.-Moll. 1 (3): pl. 15, fig. 214.

Unio tumidus var. radiolata Krynicki, 1837

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Krynicki, 1837. Bull. Soc. imp. Nat. Moscou 10(2): 59. [nomen nudum]

Unio tumidus Mousson, 1849

(reference to Unio tumidus Philipsson in Retzius, 1788)
Mousson, 1849. Land-Süßw.-Moll. Java: 95, pl. 17 [16], fig. 1.

Rectidens sumatrensis — Haas (1914)
Unio tumidus — van Benthem Jutting (1953)
Rectidens sumatrensis sumatrensis — Haas (1969)

Unio zelebori Zelebor, 1851

‘Parreyss’ Zelebor, 1851. Ber. Mit. Freunde Naturw. Wien 7: 231, sp. 8. [nomen nudum]

Unio (s.s.) tumidus zelebori — Falkner et al. (2001)
Unio tumidus zelebori — Lopes-Lima et al. (2017)

Unio pictorum var. vinceleus de Joannis, 1859

(reference to Unio pictorum var. vinceleus de Joannis, 1859)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 12, fig. 5. [in part]

Unio (Batavusiana) potamius — Germain (1904)
Unio batavus — Germain (1931)

Unio vincelleus Locard, 1882

(error for Unio pictorum var. vinceleus de Joannis, 1859)
‘de Joannis’ Locard, 1882. Prodr. Malac. Franç. 1: 298.

Unio pictorum var. vincelus Locard, 1889

(error for Unio pictorum var. vinceleus de Joannis, 1859)
‘de Joannis’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 140 [34].

Unio alpacamus Locard, 1889

(error for Unio alpecanus Bourguignat in Locard, 1882)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 178 [72].

Unio vincelens Caziot, 1907

(error for Unio pictorum var. vinceleus de Joannis, 1859)
‘Bourg.’ Caziot, 1907. Bull. Soc. Sci. Hist. Nat. Yonne 60: 276.

Unio elbovensis Germain, 1931

(error for Unio elboviensis Locard, 1893)
‘Locard’ Germain, 1931. Faune de France 22: 741.

Unio albovicensis Haas, 1940

(error for Unio elboviensis Locard, 1893)
‘Locard’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 129.

Unio subtumidus ‘Mousson, in litt.’ van Benthem Jutting, 1953

(intro'd as syn of Unio tumidus Philipsson in Retzius, 1788)
‘Mousson, in litt.’ van Benthem Jutting, 1953. Treubia 22(1): 45.

Unio tumidus — van Benthem Jutting (1953)

Unio pictorum var. vincelius Haas, 1969

(error for Unio pictorum var. vinceleus de Joannis, 1859)
‘Joannis’ Haas, 1969. Das Tierreich 88: 26.

Unio elbovicensis Haas, 1969

(error for Unio elboviensis Locard, 1893)
‘Locard’ Haas, 1969. Das Tierreich 88: 21.

Unio (Unio) tumidus ssp. [Bassin de la Saône] Falkner et al., 2002

(intro'd as intra-sp. of Unio tumidus Philipsson in Retzius, 1788)
Falkner et al., 2002. Patrim. Natur. 52: 55. [nomen nudum]

Unio (s.s.) tumidus ssp. [Bassin de la Saône] — Falkner et al. (2002)

Unio tumidu Li et al., 2010

(error for Unio tumidus Philipsson in Retzius, 1788)
Li et al., 2010. Hereditas 32(4): 363.

Unio lateborni Vinarski & Kantor, 2016

(error for Unio lauterborni Haas, 1909)
‘Haas’ Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 23.

Unio tumidus borysthensis Lopes-Lima et al., 2017

(error for Unio tumidus var. borysthenensis Kobelt, 1879)
‘Kobelt’ Lopes-Lima et al., 2017. Biol. Rev. 92(1): ??.