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Mya margaritifera Linnaeus, 1758

Lister, 1678. Hist. Anim. Anglic.: [app.] 15, pl. 1, fig. 1. [pre-linnaean]
Lister, 1685. Conch.: pl. 149, fig. 4. [pre-linnaean]
Linnaeus, 1758. Syst. Nat. 1: 671, sp. 20.
Linnaeus, 1767. Syst. Nat. 1(2): 1112, sp. 29.
Type(s): LECTOTYPE LS_29, “Habitat in totius orbis arctici cataractis.”.

Margaritana margaritifera — Simpson (1900), Kobelt (1904), Ortmann (1909), Ortmann (1910), Ortmann (1912), ... Haas (1924), Frierson (1927), Germain (1931), Zhadin (1938), Zhadin (1965)
Margaritana (s.s.) margaritifera — Thiele (1934)
Margaritifera margaritifera — Mandahl-Barth (1938), Modell (1942), Kira (1959), Heard & Guckert (1970), Burch (1975), ... Dai et al. (2025), Chen et al. (2025), FMCS (2025), Chen et al. (2025), Wang et al. (2025)
Margaritifera margaritifera margaritifera — Haas (1940), Bogatov (2022)
Unio margaritifera — Kira (1954)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969), Falkner et al. (2001), Falkner et al. (2002)
Margaritifera (s.s.) margaritifera — Haas (1969), Bolotov et al. (2016)
Margaritiferidae — Krebs et al. (2003)

Margaritana fluviatilis Schumacher, 1817

Schumacher, 1816. Overs. Dansk. Vid. Selsk. Forhandl. Kjobehhaven 1816: 7. [nomen nudum]
Schumacher, 1817. Essai. Nouv. Syst.: 124, pl. 10, fig. 4.

Margaritifera margaritifera — Habe (1977), Ziuganov et al. (1994), Watters et al. (2009), Vinarski & Kantor (2016)

Unio elongata Lamarck, 1819

Pennant, 1777. British Zool. 4: pl. 43, fig. 18.
Lamarck, 1819. Hist. Natur. Anim. sans Vert. 6: 70, sp. 2.
Type(s): POSSIBLE SYNTYPE MNHN_MPL_0028, Habite dans les rivières de l’Angleterre, et probablement du nord de l’Europe [Lives in the rivers of England and Northern Europe probably].

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Graf (2007), Araujo et al. (2009), Watters et al. (2009), Kantor et al. (2010), Fauna Europaea (2012), Bolotov et al. (2013), Vinarski & Kantor (2016)
Margaritifera elongata — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritifera margaritifera elongata — Bogatov (2022)

Alasmodonta arcuata Barnes, 1823

Barnes, 1823. Amer. J. Sci. 6 (1-2): 277, pl. 12, figs. 20a-d.

Margaritana margaritifera — Simpson (1900), Hannibal (1912), Preston (1912), Simpson (1914), Frierson (1927)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Strayer & Jirka (1997), Araujo et al. (2009), Watters et al. (2009), Vinarski & Kantor (2016)

Unio sinuata C. Pfeiffer, 1825

C. Pfeiffer, 1825. Naturgesch. Dtsch. Land-Süßw. Moll. 2: ??.
(misidentified reference to Unio sinuata Lamarck, 1819)

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914)

Unio roissyi Michaud, 1831

Michaud, 1831. Compl. Hist. Nat. Moll., Draparnaud: 112, pl. 16, figs. 27-28.

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Margaritana roissyi — Rochebrune (1903)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Watters et al. (2009), Vinarski & Kantor (2016)

Unio margaritifer var. minor Rossmässler, 1835

Rossmässler, 1835. Icon. Land- und Süssw.-Moll. 1 (2): 21, pl. 9, fig. 129.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera margaritifera — Watters et al. (2009), Vinarski & Kantor (2016)

Unio brunneus Bonhomme, 1840

Bonhomme, 1840. Mém. Aveyron 2: 460.

Margaritana margaritifera — Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera (s.s.) margaritifera brunnea — Falkner et al. (2001), Falkner et al. (2002)
Margaritifera margaritifera — Araujo et al. (2009), Vinarski & Kantor (2016), Lopes-Lima et al. (2017)

Alasmodon margaritiferus var. arcuata Brown, 1844

Brown, 1844. Ill. Rec. Conch. Gr. Brit. & Ireland: 84, pl. 30*, fig. 3.
Brown, 1845. Ill. Land & Fr. Water Conch. Gr. Brit. & Ireland: 115, pl. 23, fig. 3.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758; homonym of Alasmodonta arcuata Barnes, 1823)

Margaritana margaritifera — Simpson (1900), Simpson (1914)

Alasmodon margaritiferus var. olivacea Brown, 1844

Brown, 1844. Ill. Rec. Conch. Gr. Brit. & Ireland: 84, pl. 30*, fig. 1.
Brown, 1845. Ill. Land & Fr. Water Conch. Gr. Brit. & Ireland: 115, pl. 23, fig. 1.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera — Simpson (1900), Simpson (1914)

Unio tristis Morelet, 1845

Morelet, 1845. Moll. Terr. Fluv. Portugal: 107, pl. 31, fig. 2.
Type(s): HOLOTYPE BMNH_93-2-4-1961, la Tamega, pres a Amarante, Portugal.

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Watters et al. (2009), Vinarski & Kantor (2016)

Unio raveneliana Chenu, 1859

‘Lea’ Chenu, 1859. Manuel Conchyl. 2: 144, fig. 714.
(misidentified reference to Unio ravenelianus Lea, 1834)

Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914)

Margaritana borealis Westerlund, 1871

Westerlund, 1871. Nova Reg. Soc. Scient. 8: 168.

Margaritifera borealis — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritifera margaritifera — Graf (2007), Kantor et al. (2010), Bolotov et al. (2013), Vinarski & Kantor (2016)
Margaritifera margaritifera borealis — Bogatov (2022)

Margaritana freytagi Kobelt, 1886

Kobelt, 1886. Jb. Nassau. Ver. Naturk. 39: 98, pl. 8, figs. 1-4.
Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: 45, pl. 55, figs. 291-294.
Type(s): LECTOTYPE SMF_4984, Nister b. Hachenburg.

Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana michaudi Locard, 1889

Michaud, 1831. Compl. Hist. Nat. Moll., Draparnaud: 113, pl. 16, fig. 29 [as ‘Unio elongata’].
Locard, 1889. Contr. Faune Malacol. Franc. 13: 123 [17], 180 [74].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 150.
Type(s): PARALECTOTYPE MNHN_22368, Vire [Calvados].

Margaritana margaritifera — Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana pyrenaica Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 123 [17], 181 [75].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 150.
Type(s): SYNTYPE MNHN_22478, Carcenac (Aveyron).

Margaritana margaritifera — Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana margaritifera forma jetschini Westerlund, 1890

Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 185. [infra-subsp.]
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana alleni Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 212.

Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana margaritifera parvula Haas, 1908

Haas, 1908. Beitr. Kenntn. Mittleleurop. Najadeen, Beilage zu... 1908 (1): 15.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)
Type(s): LECTOTYPE SMF_4982, Ulfenbach bei Affolterbach.

Margaritana margaritifera parvula — Haas (1924)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera (s.s.) margaritifera parvula — Falkner et al. (2001)
Margaritifera margaritifera — Vinarski & Kantor (2016), Lopes-Lima et al. (2017)

Margaritana margaritifera var. siluriana Bloomer, 1927

Bloomer, 1927. Proc. Malac. Soc. London 17: 211.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana durrovensis Phillips, 1928

Phillips, 1928. Proc. Malac. Soc. London 18: 71, pls. 3-5.
Bloomer, 1928. Proc. Malac. Soc. London 18: 74, text fig.
Type(s): SYNTYPES BMNH_1928-10-23-1-2, River Nore, Durrow, Queen’s Co., Ireland.

Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016), Lopes-Lima et al. (2017)
Margaritifera margaritifera durrovensis — IUCN (1996), Machordom et al. (2003), Walker et al. (2006)
Margaritifera (s.s.) margaritifera durrovensis — Falkner et al. (2001)

Mya margaritifera Da Costa, 1778

Da Costa, 1778. British Conch.: 225, pl. 15, fig. 3.
(reference to Mya margaritifera Linnaeus, 1758)

Unio elongata Michaud, 1831

Michaud, 1831. Compl. Hist. Nat. Moll., Draparnaud: 113, pl. 16, fig. 29.
(reference to Unio elongata Lamarck, 1819)

Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Margaritana margaritifera — Germain (1931)

Unio margaritifera Rossmässler, 1835

Rossmässler, 1835. Icon. Land- und Süssw.-Moll. 1 (1): pl. 4, fig. 74. [in part]
(reference to Mya margaritifera Linnaeus, 1758)

Unio rivalis Lea, 1836

‘Zeig.’ Lea, 1836. Synopsis Fam. Naiades: 46.
‘Zeig.’ Lea, 1838. Synopsis Fam. Naiades: 28.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Unio roisii Lea, 1852

‘Mich.’ Lea, 1852. Synopsis Fam. Naiades: 40.
(error for Unio roissyi Michaud, 1831)

Unio reyniesii Lea, 1870

‘Mich.’ Lea, 1870. Synopsis Fam. Naiades: 64.
(error for Unio roissyi Michaud, 1831)

Margaritana margaritifera var. compressa Westerlund, 1873

Westerlund, 1873. Fauna Moll. Terr. Fluv. Suec. Norv. Dan. (2): 578.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana margaritifera var. ventricosa Westerlund, 1873

Westerlund, 1873. Fauna Moll. Terr. Fluv. Suec. Norv. Dan. (2): 578.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana durrovensis Bloomer, 1928

Bloomer, 1928. Proc. Malac. Soc. London 18: 74, text fig.
(reference to Margaritana durrovensis Phillips, 1928)

Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Fauna Europaea (2012)

Margaritana durrowensis Haas, 1940

‘Phillips’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 120.
(error for Margaritana durrovensis Phillips, 1928)

Margaritana tenerus Ziuganov et al., 1994

Ziuganov et al., 1994. Freshw. Pearl Muss. & Fish: 59.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Margaritifera margaritifera — Ziuganov et al. (1994)

Margaritana vulgaris Ziuganov et al., 1994

Ziuganov et al., 1994. Freshw. Pearl Muss. & Fish: 59.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Margaritifera margaritifera — Ziuganov et al. (1994)

Margaritana pirenaica Vinarski & Kantor, 2016

‘Boruguignat in Locard’ Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 59.
(error for Margaritana pyrenaica Locard, 1889)