species Unio delphinus Spengler, 1793: synonymy

Unio delphinus Spengler, 1793

Spengler, 1793. Skr. Naturhist. Selsk. 3: 63, no. 28.

Unio delphinus — Simpson (1914), Araujo et al. (2009), Khalloufi et al. (2011), IUCN (2011), Graf & Cummings (2011), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Campbell & Lydeard (2026)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Hyriopsis bialata — Klymus et al. (2025)

Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844

‘Moq.-Tand. in litt.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 26, pl. 56, fig. 747.

Unio (sec. Lymnium) turtoni — Simpson (1900)
Unio (sec. Lymnium) (Group Unio pictorum) turtonii — Simpson (1914)
Unio requieni — Germain (1931)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009), Fauna Europaea (2012)
Unio mancus requienii — Prie (2012)

Unio dactylus Morelet, 1845

(homonym of Unio dactylus Lea, 1840)
Morelet, 1845. Moll. Terr. Fluv. Portugal: 110, pl. 14, fig. 2.
Type(s): HOLOTYPE BMNH_93-2-4-1587, Guadiana, near Castro, Portugal.

Unio (sec. Lymnium) pictorum — Simpson (1900)
Unio (sec. Lymnium) (Group Unio pictorum) pictorum — Simpson (1914)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009), Fauna Europaea (2012)

Unio mucidus Morelet, 1845

Morelet, 1845. Moll. Terr. Fluv. Portugal: 111, pl. 14, fig. 3.
Type(s): HOLOTYPE BMNH_92-2-4-1974, le Cavado, Portugal.

Unio (sec. Lymnium) pictorum — Simpson (1900)
Unio (sec. Lymnium) (Group Unio pictorum) pictorum — Simpson (1914)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Fauna Europaea (2012), Lopes-Lima et al. (2017), Araujo et al. (2018)
Unio pictorum — He & Zhuang (2013)

Unio hyperephanus da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 289.
Castro in Locard, 1899. Conchyliol. Portug.: 244.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio nevesi da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 290.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio simoesi da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 292.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio (hispanus var.?) sevillensis Kobelt, 1887

(intro'd as intra-sp. of Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)
Kobelt, 1887. An. Soc. Españ. Hist. Nat. 16: 437.
Kobelt, 1888. Icon. Land- und Süssw.-Moll. 3: 56, pl. 89, fig. 496.

Unio pictorum delphinus — Haas (1940), Haas (1969)

Unio sousanus Locard, 1889

Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3 [as ‘Unio requienii var. acruata’].
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2 [as ‘Unio requienii var. acruatus’].
‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 197, fig. 211 [as ‘souzanus’].
‘Castro’ Locard, 1899. Conchyliol. Portug.: 258.
Type(s): LECTOTYPE MNHN_22545 (by Graf, 2011: 505), le Rhône, à Valence (Drôme).

Unio (Cyrtusiana) souzanus [sic] — Germain (1904)
Unio requieni — Germain (1931)
Unio elongatulus requienii — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)
Unio mancus requienii — Prie (2012)

Unio subhispanus Locard, 1889

‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 171 [65], 253 [147].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 200.
‘Castro’ Locard, 1899. Conchyliol. Portug.: 244.
Type(s): LECTOTYPE MNHN_22562 (by Graf, 2011: 507), La Loire, à Saint-Gemmes, près Angers.

Unio (Hispaniana) subhispanus — Germain (1904)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)
Unio mancus requienii — Prie (2012)

Unio hispanus var. sphenoides Westerlund, 1892

(intro'd as intra-sp. of Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)
Westerlund, 1892. An. Soc. Españ. Hist. Nat. 21: 390.

Unio pictorum delphinus — Haas (1940), Haas (1969)

Unio cameratus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 183, pl. 2, fig. 8.

Unio cameratus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio decurtatus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 170.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 185, pl. 1, fig. 9.

Unio decurtatus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio gravatus Drouët, 1893

Drouët, 1893. Unionidae de l’Espagne: 49.

Unio gravatus — Kobelt (1903)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio limosellus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 184, pl. 12, fig. 4.

Unio limosellus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio souzanus ‘Castro’ of Authors

(error for Unio sousanus Locard, 1889)
‘Castro’ of Authors.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 197.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 275 [37].
Germain, 1931. Faune de France 22: 737.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 126.

Unio abrantesianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 253.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio allenianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 255.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio barbozanus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 233.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio castroi Bourguignat in Locard, 1899

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Bourguignat in Locard, 1899. Conchyliol. Portug.: 234.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chasmirhynchus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 246.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chorellinus Locard, 1899

Locard, 1899. Conchyliol. Portug.: 228.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chorellus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52] [as ‘chlorellus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 227.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio cyrtus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 257.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio euchasmus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 249.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio hypoxanthus Casto in Locard, 1899

Casto in Locard, 1899. Conchyliol. Portug.: 226.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio mundanus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 247.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio neothaumus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 259.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio novus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 258.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio ocresanus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 236.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio oncomensis Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46] [as ‘oncomenus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 225.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio paulinoi Locard, 1899

Locard, 1899. Conchyliol. Portug.: 242.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio scalabisianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 254.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio schousboei Bourguignat in Locard, 1899

Bourguignat in Locard, 1899. Conchyliol. Portug.: 250.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio silvai Bourguignat in Locard, 1899

Bourguignat in Locard, 1899. Conchyliol. Portug.: 235.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio submucidus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 230.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio taganus Servain in Locard, 1899

Servain in Locard, 1899. Conchyliol. Portug.: 252.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio tameganus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 221.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio requienii taginus Kobelt, 1903

(homonym of Unio littoralis taginus Kobelt, 1903; intro'd as intra-sp. of Unio requienii Michaud, 1831)
Kobelt, 1903. Icon. Land- und Süssw.-Moll. 10: 28, pl. 280, fig. 1796.

Unio requienii taginus — Kobelt (1903)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio subhispanicus ‘Castro in Locard’ of Authors

(error for Unio subhispanus Locard, 1889)
‘Castro in Locard’ of Authors.
Germain, 1931. Faune de France 22: 737.
Prie, 2012. MalaCo 8: 432.

Unio requieni — Germain (1931)

Hyriopsis bialata Klymus et al., 2025

(misidentified reference to Hyriopsis bialatus Simpson, 1900)
Klymus et al., 2025. mtDNA B 10(6): Table 1.

Unio hispanicus Graëls, 1846

(error for Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)
Graëls, 1846. Catal. Moll. Esp.: 22.
‘Moquin. Rossm.’ Lea, 1852. Synopsis Fam. Naiades: 30.
‘Moquin. Rossm.’ Lea, 1870. Synopsis Fam. Naiades: 48.

Unio requienii var. arcuata Drouët, 1857

(misidentified reference to Unio arcuata Jacquemin, 1835)
Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3.
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2.

Unio (Cyrtusiana) souzanus [sic] — Germain (1904)
Unio requieni — Germain (1931)

Unio hispanus Lowe, 1860

(reference to Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)
Lowe, 1860. unknown publication: 176.

Unio elongatulus durieui — Daget (1998)

Unio chlorellus Castro in Locard, 1889

(error for Unio chorellus Castro in Locard, 1899)
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52].

Unio oncomenus Castro in Locard, 1889

(error for Unio oncomensis Castro in Locard, 1899)
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46].