species Unio delphinus Spengler, 1793: synonymy

Unio delphinus Spengler, 1793

Spengler, 1793. Skr. Naturhist. Selsk. 3: 63, no. 28.

Unio delphinus — Simpson (1900), Simpson (1914), Araujo et al. (2009), Khalloufi et al. (2011), IUCN (2011), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Campbell & Lydeard (2026)
Unio pictorum delphinus — Haas (1940), Haas (1969)

Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844

‘Moq.-Tand. in litt.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 26, pl. 56, fig. 747.

Unio (sec. Lymnium) turtoni — Simpson (1900), Simpson (1914)
Unio requieni — Germain (1931)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009), Fauna Europaea (2012)
Unio mancus requienii — Prie (2012)

Unio dactylus Morelet, 1845

Morelet, 1845. Moll. Terr. Fluv. Portugal: 110, pl. 14, fig. 2.
(homonym of Unio dactylus Lea, 1840)
Type(s): HOLOTYPE BMNH_93-2-4-1587, Guadiana, near Castro, Portugal.

Unio (sec. Lymnium) pictorum — Simpson (1900), Simpson (1914)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009), Fauna Europaea (2012)

Unio mucidus Morelet, 1845

Morelet, 1845. Moll. Terr. Fluv. Portugal: 111, pl. 14, fig. 3.
Type(s): HOLOTYPE BMNH_92-2-4-1974, le Cavado, Portugal.

Unio (sec. Lymnium) pictorum — Simpson (1900), Simpson (1914)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Fauna Europaea (2012), Lopes-Lima et al. (2017), Araujo et al. (2018)
Unio pictorum — He & Zhuang (2013)

Unio hyperephanus da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 289.
Castro in Locard, 1899. Conchyliol. Portug.: 244.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio nevesi da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 290.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio simoesi da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 292.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio (hispanus var.?) sevillensis Kobelt, 1887

Kobelt, 1887. An. Soc. Españ. Hist. Nat. 16: 437.
Kobelt, 1888. Icon. Land- und Süssw.-Moll. 3: 56, pl. 89, fig. 496.
(intro'd as intra-sp. of Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)

Unio pictorum delphinus — Haas (1940), Haas (1969)

Unio sousanus Locard, 1889

Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3 [as ‘Unio requienii var. acruata’].
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2 [as ‘Unio requienii var. acruatus’].
‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 197, fig. 211 [as ‘souzanus’].
‘Castro’ Locard, 1899. Conchyliol. Portug.: 258.
Type(s): LECTOTYPE MNHN_22545 (by Graf, 2011: 505), le Rhône, à Valence (Drôme).

Unio (Cyrtusiana) souzanus [sic] — Germain (1904)
Unio requieni — Germain (1931)
Unio elongatulus requienii — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)
Unio mancus requienii — Prie (2012)

Unio subhispanus Locard, 1889

‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 171 [65], 253 [147].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 200.
‘Castro’ Locard, 1899. Conchyliol. Portug.: 244.
Type(s): LECTOTYPE MNHN_22562 (by Graf, 2011: 507), La Loire, à Saint-Gemmes, près Angers.

Unio (Hispaniana) subhispanus — Germain (1904)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)
Unio mancus requienii — Prie (2012)

Unio hispanus var. sphenoides Westerlund, 1892

Westerlund, 1892. An. Soc. Españ. Hist. Nat. 21: 390.
(intro'd as intra-sp. of Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)

Unio pictorum delphinus — Haas (1940), Haas (1969)

Unio cameratus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 183, pl. 2, fig. 8.

Unio cameratus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio decurtatus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 170.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 185, pl. 1, fig. 9.

Unio decurtatus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio gravatus Drouët, 1893

Drouët, 1893. Unionidae de l’Espagne: 49.

Unio gravatus — Kobelt (1903)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio limosellus Drouët, 1893

Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 184, pl. 12, fig. 4.

Unio limosellus — Kobelt (1903)
Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio souzanus ‘Castro’ of Authors

‘Castro’ of Authors.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 197.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 275 [37].
Germain, 1931. Faune de France 22: 737.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 126.
(error for Unio sousanus Locard, 1889)

Unio abrantesianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 253.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio allenianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 255.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio barbozanus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 233.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio castroi Bourguignat in Locard, 1899

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Bourguignat in Locard, 1899. Conchyliol. Portug.: 234.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chasmirhynchus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 246.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chorellinus Locard, 1899

Locard, 1899. Conchyliol. Portug.: 228.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio chorellus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52] [as ‘chlorellus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 227.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio cyrtus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 257.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio euchasmus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 249.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio hypoxanthus Casto in Locard, 1899

Casto in Locard, 1899. Conchyliol. Portug.: 226.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio mundanus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 247.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio neothaumus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 259.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio novus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 258.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio ocresanus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 236.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio oncomensis Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46] [as ‘oncomenus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 225.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio paulinoi Locard, 1899

Locard, 1899. Conchyliol. Portug.: 242.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio scalabisianus Castro in Locard, 1899

Castro in Locard, 1899. Conchyliol. Portug.: 254.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio schousboei Bourguignat in Locard, 1899

Bourguignat in Locard, 1899. Conchyliol. Portug.: 250.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio silvai Bourguignat in Locard, 1899

Bourguignat in Locard, 1899. Conchyliol. Portug.: 235.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio submucidus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 230.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio taganus Servain in Locard, 1899

Servain in Locard, 1899. Conchyliol. Portug.: 252.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio tameganus Castro in Locard, 1899

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 221.

Unio pictorum mucidus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio requienii taginus Kobelt, 1903

Kobelt, 1903. Icon. Land- und Süssw.-Moll. 10: 28, pl. 280, fig. 1796.
(homonym of Unio littoralis taginus Kobelt, 1903; intro'd as intra-sp. of Unio requienii Michaud, 1831)

Unio requienii taginus — Kobelt (1903)
Unio pictorum delphinus — Haas (1940), Haas (1969)
Unio delphinus — Araujo et al. (2009)

Unio subhispanicus ‘Castro in Locard’ of Authors

‘Castro in Locard’ of Authors.
Germain, 1931. Faune de France 22: 737.
Prie, 2012. MalaCo 8: 432.
(error for Unio subhispanus Locard, 1889)

Unio requieni — Germain (1931)

Unio hispanicus Graëls, 1846

Graëls, 1846. Catal. Moll. Esp.: 22.
‘Moquin. Rossm.’ Lea, 1852. Synopsis Fam. Naiades: 30.
‘Moquin. Rossm.’ Lea, 1870. Synopsis Fam. Naiades: 48.
(error for Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)

Unio requienii var. arcuata Drouët, 1857

Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3.
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2.
(misidentified reference to Unio arcuata Jacquemin, 1835)

Unio (Cyrtusiana) souzanus [sic] — Germain (1904)
Unio requieni — Germain (1931)

Unio hispanus Lowe, 1860

Lowe, 1860. unknown publication: 176.
(reference to Unio hispanus ‘Moq.-Tand. in litt.’ Rossmässler, 1844)

Unio elongatulus durieui — Daget (1998)

Unio chlorellus Castro in Locard, 1889

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52].
(error for Unio chorellus Castro in Locard, 1899)

Unio oncomenus Castro in Locard, 1889

Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46].
(error for Unio oncomensis Castro in Locard, 1899)