Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina

genus Pustulosa Frierson, 1927: cladomics

8 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 8 (9 total)
analyzed in 42 cladograms in 24 publications
maximum number of terminals in an analysis = 8 (mean = 3.14, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 25
maximum number of included terminals = 8, (mean = 4.6 median = 4)
proportion of cladograms that support basic monophyly = 0.24

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pustulosa cladogram distances
PCoA Ordination

NJ Tree of Pustulosa cladogram distances
NJ Tree

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 19 9 3 **
Cladomics cladogram Theliderma Cyclonaias Quadrula Pustulosa nodulata Pustulosa necki Pustulosa petrina Pustulosa kieneriana Pustulosa pustulosa Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Johnson et al., 2018 [Publ 3844], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 8 9 7 8 8 4 3 9.0
Cladomics cladogram Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa nodulata Pustulosa petrina

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.600 8 9 7 8 20 6 3 7.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa Quadrula nobilis Theliderma Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 8 9 7 8 21 9 3 9.0
Cladomics cladogram Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.600 8 9 7 8 18 6 3 9.0
Cladomics cladogram Theliderma Quadrula quadrula Quadrula verrucosa Quadrula nobilis Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.400 8 9 7 8 625 8 3 7.0
Cladomics cladogram Theliderma johnsoni Theliderma metanevra Quadrula nobilis Quadrula verrucosa Theliderma cylindrica Quadrula fragosa Quadrula quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Serb et al., 2003 [Publ 2659], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 6 23 7 2 7.0
Cladomics cladogram Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa petrina

Serb et al., 2003 [Publ 2659], fig. 3

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 23 7 2 8.0
Cladomics cladogram Quadrula verrucosa Quadrula nobilis Quadrula quadrula Theliderma Pustulosa kieneriana Pustulosa infucata Pustulosa succissa Pustulosa pustulosa Pustulosa nodulata Pustulosa petrina

Szumowski et al., 2012 [Publ 3569], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 8 7 2 8.0
Cladomics cladogram Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa petrina

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 4 5 2 4 98 5 3 6.0
Cladomics cladogram Theliderma Quadrula Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.600 4 5 2 4 98 5 3 6.0
Cladomics cladogram Theliderma Quadrula quadrula Quadrula nobilis Quadrula verrucosa Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.600 4 5 3 4 69 5 3 7.0
Cladomics cladogram Theliderma Quadrula verrucosa Quadrula quadrula Quadrula nobilis Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

cst iT tT bi riS oT roT roS iD
0.600 4 5 3 4 69 5 3 7.0
Cladomics cladogram Quadrula verrucosa Quadrula nobilis Quadrula quadrula Theliderma Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 4 5 3 4 38 2 2 7.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 4 5 3 4 38 2 2 7.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

cst iT tT bi riS oT roT roS iD
0.750 4 6 4 4 39 2 2 8.0
Cladomics cladogram Uniomerus Pustulosa infucata Pustulosa kleiniana Quadrula Cyclonaias Pustulosa succissa Pustulosa pustulosa

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 4 5 3 4 38 2 2 7.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 3 4 2 3 15 4 2 8.0
Cladomics cladogram Theliderma Quadrula quadrula Pustulosa infucata Quadrula verrucosa Pustulosa kieneriana Pustulosa succissa

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
0.600 2 3 0 2 33 4 3 7.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa Theliderma Pustulosa pustulosa Cyclonaias Pustulosa nodulata

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 2 3 0 2 36 4 3 7.0
Cladomics cladogram Theliderma Quadrula Pustulosa pustulosa Cyclonaias Pustulosa nodulata

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 2 3 0 2 36 3 2 7.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa Pustulosa pustulosa Pustulosa petrina Cyclonaias

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 1 1 --
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa succissa

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 12 1 1 7.0
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 1 1 --
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa succissa

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 12 1 1 7.0
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.600 1 5 0 1 104 4 3 12.0
Cladomics cladogram Uniomerus Cyclonaias Pustulosa pustulosa 1 Theliderma Pustulosa pustulosa 2 Quadrula quadrula Quadrula verrucosa

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 2 2 --
Cladomics cladogram Quadrula Cyclonaias Pustulosa pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Chapman et al., 2008 [Publ 2667], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Chapman et al., 2008 [Publ 2744], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 1 1 --
Cladomics cladogram Cyclonaias Pustulosa pustulosa

Pieri et al., 2018 [Publ 3867], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Quadrula Pustulosa petrina

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 1 1 --
Cladomics cladogram Pustulosa petrina Quadrula

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Pustulosa petrina Quadrula

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 10 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa Pleurobema Amblema Obovaria Ortmanniana Lampsilis

Freeman et al., 2025 [Publ 4443], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Freeman et al., 2025 [Publ 4443], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa

Freeman et al., 2025 [Publ 4443], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram Quadrula Pustulosa pustulosa