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Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Unionoidea | Margaritiferidae | Gibbosulinae

genus Gibbosula Simpson, 1900: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 75 cladograms in 42 publications
maximum number of terminals in an analysis = 4 (mean = 1.32, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 4, (mean = 2.26 median = 2)
proportion of cladograms that support basic monophyly = 0.79

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Gibbosula cladogram distances
PCoA Ordination

NJ Tree of Gibbosula cladogram distances
NJ Tree

Cladograms

Chen et al., 2025 [Publ 4455], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	22	0	0	0

     +-- Gibbosula_laosensis **     
  +--+                           
  |  +-- Gibbosula_sinensis **      
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	25	0	0	3

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Lyubas et al., 2019 [Publ 3963], fig. 10

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	13	0	0	3

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	25	0	0	3

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	27	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
2	13	0	2	197	0	0	13

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
2	13	0	2	199	0	0	13

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Cumberlandia            
     +--+                              
        |  +-- Pseudunio               
        +--+                           
           +-- Margaritifera

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	0	0	--

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	66	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	13	0	2	199	0	0	13

  +-------- Gibbosula_rochechouartii **
  |                                 
--+  +----- Gibbosula_laosensis **     
  |  |                              
  +--+  +-- Cumberlandia            
     |  |                           
     +--+-- Pseudunio               
        |                           
        +-- Margaritifera

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	12	0	2	32	0	0	12

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	8	0	0	--

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	66	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	0	0	--

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	66	0	0	--

  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	67	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	0	0	--

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	51	0	0	--

-- Gibbosula_laosensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	0	0	--

-- Gibbosula_laosensis **

Bespalaya et al., 2018 [Publ 3811], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	0	0	--

-- Gibbosula_laosensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	117	0	0	--

-- Gibbosula_laosensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	0	0	--

-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Gibbosula_crassa **

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	0	0	--

-- Gibbosula_rochechouartii **

Wang et al., 2025 [Publ 4488], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	64	0	0	--

-- Gibbosula_rochechouartii **

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	0	0	--

-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4147], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Gibbosula_rochechouartii **

Vikhrev et al., 2017 [Publ 3810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Gibbosula_rochechouartii **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	0	0	--

-- Gibbosula_rochechouartii **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_crassa **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	0	0	--

-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Gibbosula_rochechouartii **

Chen et al., 2020 [Publ 4170], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Gibbosula_crassa **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	0	0	--

-- Gibbosula_laosensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	0	0	--

-- Gibbosula_crassa **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	0	0	--

-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	0	0	--

-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	0	0	--

-- Gibbosula_crassa **

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Gibbosula_rochechouartii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Gibbosula_laosensis **

Bolotov et al., 2016 [Publ 3743], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4385], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Gibbosula_laosensis **

Vikhrev et al., 2018 [Publ 3812], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	0	0	--

-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Gibbosula_crassa **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	0	0	--

-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	0	0	--

-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Gibbosula_crassa **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	0	0	--

-- Gibbosula_laosensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	0	0	--

-- Gibbosula_laosensis **

Huang et al., 2018 [Publ 3828], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	0	0	--

-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	0	0	--

-- Gibbosula_rochechouartii **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_rochechouartii **

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_rochechouartii **

Bolotov et al., 2016 [Publ 3743], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	0	0	--

-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	6	0	0	--

-- Gibbosula_crassa **

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Gibbosula_rochechouartii **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_crassa **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Gibbosula_rochechouartii **

Bolotov et al., 2016 [Publ 3732], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Gibbosula_laosensis **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Gibbosula_rochechouartii **

Araujo et al., 2016 [Publ 3746], fig. 2a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Gibbosula_laosensis **

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