Bivalvia | Unionida | Unionoidea | Margaritiferidae | Gibbosulinae

genus Gibbosula Simpson, 1900: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 81 cladograms in 46 publications
maximum number of terminals in an analysis = 4 (mean = 1.35, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 4, (mean = 2.33 median = 2)
proportion of cladograms that support basic monophyly = 0.81

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Gibbosula cladogram distances
PCoA Ordination

NJ Tree of Gibbosula cladogram distances
NJ Tree

Cladograms

Chen et al., 2025 [Publ 4455], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 4 22 0 0 0 
     +-- Gibbosula_laosensis **     
  +--+                           
  |  +-- Gibbosula_sinensis **      
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3 3 1 3 138 0 0 3 
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
3 3 1 3 628 0 0 1 
  +----- Gibbosula_laosensis **     
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 25 0 0 3 
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Lyubas et al., 2019 [Publ 3963], fig. 10

iTtTbiriSoTroTroSiD
3 3 1 3 13 0 0 3 
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 25 0 0 3 
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 0 0 -- 
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2 13 0 2 198 0 0 13 
  +-------- Gibbosula_rochechouartii **
  |                                 
--+  +----- Gibbosula_laosensis **     
  |  |                              
  +--+  +-- Cumberlandia            
     |  |                           
     +--+-- Pseudunio               
        |                           
        +-- Margaritifera

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 12 0 2 32 0 0 12 
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2 13 0 2 196 0 0 13 
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 0 0 -- 
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2 13 0 2 198 0 0 13 
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Cumberlandia            
     +--+                              
        |  +-- Pseudunio               
        +--+                           
           +-- Margaritifera

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 27 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 8 0 0 -- 
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 0 0 -- 
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Gibbosula_laosensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_rochechouartii **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Gibbosula_laosensis **

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Gibbosula_laosensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Gibbosula_laosensis **

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_rochechouartii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Gibbosula_rochechouartii **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 94 0 0 -- 
-- Gibbosula_rochechouartii **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Gibbosula_laosensis **

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Gibbosula_laosensis **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Gibbosula_crassa **

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Gibbosula_crassa **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Gibbosula_rochechouartii **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 47 0 0 -- 
-- Gibbosula_laosensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 0 0 -- 
-- Gibbosula_crassa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_crassa **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 0 0 -- 
-- Gibbosula_laosensis **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Gibbosula_laosensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Gibbosula_rochechouartii **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_rochechouartii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Gibbosula_laosensis **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Gibbosula_crassa **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Gibbosula_rochechouartii **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 53 0 0 -- 
-- Gibbosula_crassa **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Gibbosula_crassa **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 50 0 0 -- 
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Gibbosula_rochechouartii **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 116 0 0 -- 
-- Gibbosula_laosensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Gibbosula_rochechouartii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Gibbosula_crassa **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Gibbosula_laosensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Gibbosula_laosensis **

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Gibbosula_rochechouartii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Gibbosula_rochechouartii **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 64 0 0 -- 
-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 0 0 -- 
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Gibbosula_crassa **