MUSSELpdb

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Unionidae

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Species & Genus Richness by
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Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio crassus Philipsson in Retzius, 1788: specimens

113 available nominal species synonyms (191 total synonyms)

Recent Taxonomic Opinions

Unio crassus — Graf (2007), Graf & Cummings (2007), Manendo et al. (2008), Dhora (2009), Araujo et al. (2009), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Wang et al. (2025)
Unio crassus gontieri — Graf (2007)
Pleurobema sintoxia — Williams et al. (2008)
Crassiana crassa crassa — Kantor et al. (2010)
Crassiana cyprinorum — Kantor et al. (2010)
Crassiana irenjensis — Kantor et al. (2010)
Crassiana irgizlaica — Kantor et al. (2010)
Crassiana musiva musiva — Kantor et al. (2010)
Unio crassus crassus — Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017), Bogatov (2022)
Unio crassus batavus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio mancus mancus — Prie (2012)
Unio mancus moquinianus — Prie (2012)
Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016)
Unio (Crassiana) irenjensis — Vinarski & Kantor (2016)
Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016)
Unio (Crassiana) musivus — Vinarski & Kantor (2016)
Unio crassus cf. courtillieri — Klishko et al. (2017)
Unio crassus courtillierii [sic] — Araujo et al. (2018)
Unio c. crassus — Príe et al. (2021)
Unio c. courtillieri — Príe et al. (2021)
Unio cf. courtillieri — Lopes-Lima et al. (2021)
Unio crassus musivus — Bogatov (2022)

Range

Widespread in continental Europe through the Dardanelles to Transcaucasia.

Map of Georeferenced Specimens

MUSSELp Specimen Records

MNHN_22146. La Canne à Saint-Saulge

Bosnia and Herzegovina

INHS_8376. Lasva

France

MNHN_22128. a Grosne, à La Ferté

MNHN_22219. Domsure (Jura)

INHS_21545. France

INHS_21552. Hungary

MNHN_22218. L’Eure, près d’Évreux

MNHN_22353. l’Oise, à Compiègne

MNHN_22352. l’Oise, à Creil

MNHN_22354. l’Ornoise (Meuse)

MNHN_22184. La Drée (Saône-et-Loire)

MNHN_22183. La Drée, à Épinac (Saône-et-Loire)

MNHN_22244. La Grosne, à la Ferté (Saône-et-Loire)

MNHN_22245. La Grosne, à Marnay (Saône-et-Loire)

MNHN_22525. La Grosne, à Marnay (Saône-et-Loire)

MNHN_22361. La Loire à Balbigny (Loire)

MNHN_22242. La Loire à Nantes (Loire-Infereur)

MNHN_22524. La Loire à Nantes (Loire-Infereur)

MNHN_22126. La Loire à Villerest (Loire)

MNHN_22362. La Loire à Villerest (Loire)

MNHN_22123. La Loire, à Balbigny (Loire)

MNHN_MP_2453. La Loire, à Balbigny (Loire) / la Canne, Nievre

MNHN_22307. La Loire, à Nantes

MNHN_22356. La Marne à Lagny (Seine-et-Marne)

MNHN_22530. La Marne à Lagny (Seine-et-Marne)

MNHN_22560. La Marne à Lagny (Seine-et-Marne)

MNHN_22243. La Saône à Lyon [Rhône]

MNHN_22349. la Seine, à Charenton

MNHN_22348. la Seine, à Corbeil

MNHN_22561. le Grosne, à Marnay (Saône-et-Loire)

MNHN_22125. le Mandrezey, à Saulcy-sur-Meurthe (Vosges)

MNHN_22099. Le Mans (Sarthe)

MNHN_22351. le Meurthe à Lunéville

MNHN_22147. Marais de Dampierre

MNHN_22350. Pontoise

MNHN_22124. Saint-Germain-des Fossés

INHS_21549. Saône River, Lyon, Rhone, Rhône-Alpes, France

INHS_21550. Saône River, Saone, Rhône-Alpes, France

MNHN_22484. St-Amour (Jura)

INHS_21553. Trencsin, Hungary

Germany

INHS_21558. Germany

INHS_21560. Nister River, Nister, Rhineland-Palatinate, Germany

INHS_21561. Oberschlesien [Upper Silesia], Germany

INHS_21557. Oder River, Vizenot Oppehr. [?], Germany

INHS_21559. Ottmachari, Silesia, Germany

INHS_21665. Schleswig- Holstein, Germany

Iraq

Romania

INHS_21566. Transylvania, Romania

Switzerland

INHS_21563. Biwir Creek, Switzerland

INHS_21564. Geneva, Marais de Lionett [Lionett Marsh], Switzerland

INHS_21565. Lake Leman, Villeneuve, Vaud, Switzerland

Ukraine

INHS_22459. Pzisluch [Sluch] River, Rivne Region, Rivne Oblast, Ukraine

INHS_22460. Ubort River, near Olevsk, Zhytomyr Region, Zhytomyr Oblast, Ukraine

Genbank Vouchers

Georeferenced Genbank Vouchers

Austria

CIIMAR_BIV4658. (COI: OR841742, go to Genbank) Aschach River (Danube Basin), Austria (48.321463, 14.013764).

CIIMAR_BIV4659. (COI: OR841743, go to Genbank) ditto.

CIIMAR_BIV4660. (COI: OR841744, go to Genbank) ditto.

CIIMAR_BIV4661. (COI: OR841745, go to Genbank) ditto.

Belarus

RMBH_biv_1230_1. (COI: ON648805, go to Genbank) Belarus: Ross, Neman Basin (53.0442, 24.4229).

Belgium

TUM_GH02. (COI: OR841750, go to Genbank) Grande Honelles River (Scheldt Basin), Belgium (50.384939, 3.687902).

TUM_GH03. (COI: OR841751, go to Genbank) ditto.

TUM_GH04. (COI: OR841752, go to Genbank) ditto.

TUM_GH05. (COI: OR841753, go to Genbank) ditto.

TUM_ME01. (COI: OR841746, go to Genbank) Mehaigne River (Meuse Basin), Belgium (50.586811, 5.189187).

TUM_ME03. (COI: OR841747, go to Genbank) ditto.

TUM_ME04. (COI: OR841748, go to Genbank) ditto.

TUM_ME05. (COI: OR841749, go to Genbank) ditto.

Bulgaria

CIIMAR_BIV7928. (COI: OR841759, go to Genbank) Dyavolska River (Dyavolska Basin), Bulgaria (42.26879, 27.66478).

CIIMAR_BIV7929. (COI: OR841760, go to Genbank) ditto.

CIIMAR_BIV7930. (COI: OR841761, go to Genbank) ditto.

CIIMAR_BIV7931. (COI: OR841762, go to Genbank) ditto.

CIIMAR_BIV7768. (COI: OR841754, go to Genbank) Osam River (Danube Basin), Bulgaria (43.40044, 25.1551).

CIIMAR_BIV7769. (COI: OR841755, go to Genbank) ditto.

CIIMAR_BIV7770. (COI: OR841756, go to Genbank) ditto.

CIIMAR_BIV7771. (COI: OR841757, go to Genbank) ditto.

CIIMAR_BIV7772. (COI: OR841758, go to Genbank) ditto.

Croatia

CIIMAR_BIV6093. (COI: OR841763, go to Genbank) Kupa River (Danube Basin), Croatia (45.628231, 15.508737).

CIIMAR_BIV6094. (COI: OR841764, go to Genbank) ditto.

CIIMAR_BIV6095. (COI: OR841765, go to Genbank) ditto.

CIIMAR_BIV6096. (COI: OR841766, go to Genbank) ditto.

CIIMAR_BIV6097. (COI: OR841767, go to Genbank) ditto.

Czech Republic

Mp_3653_ACZ1. (COI: KJ526773, go to Genbank) Chumava River (Elbe Basin), Czechia (49.870833, 14.004722).

Mp_3653_ACZ11. (COI: KJ526782, go to Genbank) ditto.

Mp_3653_ACZ13. (COI: KJ526783, go to Genbank) ditto.

Mp_3653_ACZ2. (COI: KJ526774, go to Genbank) ditto.

Mp_3653_ACZ3. (COI: KJ526775, go to Genbank) ditto.

Mp_3653_ACZ4. (COI: KJ526776, go to Genbank) ditto.

Mp_3653_ACZ5. (COI: KJ526777, go to Genbank) ditto.

Mp_3653_ACZ6. (COI: KJ526778, go to Genbank) ditto.

Mp_3653_ACZ8. (COI: KJ526780, go to Genbank) ditto.

Mp_3653_ACZ9. (COI: KJ526781, go to Genbank) ditto.

CIIMAR_UC07. (COI: OR841768, go to Genbank) Luznice River (Elbe Basin), Czechia (49.311013, 14.502444).

CIIMAR_UC08. (COI: OR841769, go to Genbank) ditto.

CIIMAR_UC09. (COI: OR841770, go to Genbank) ditto.

Mp_3653_BCZ26. (COI: KJ526784, go to Genbank) Sazava River (Elbe Basin), Czechia (49.856111, 14.700278).

Mp_3653_BCZ27. (COI: KJ526785, go to Genbank) ditto.

Mp_3653_BCZ28. (COI: KJ526786, go to Genbank) ditto.

Mp_3653_BCZ29. (COI: KJ526787, go to Genbank) ditto.

Mp_3653_BCZ30. (COI: KJ526788, go to Genbank) ditto.

Mp_3653_BCZ32. (COI: KJ526789, go to Genbank) ditto.

Mp_3653_BCZ33. (COI: KJ526790, go to Genbank) ditto.

Mp_3653_BCZ34. (COI: KJ526791, go to Genbank) ditto.

Mp_3653_BCZ35. (COI: KJ526792, go to Genbank) ditto.

Mp_3653_BCZ37. (COI: KJ526793, go to Genbank) ditto.

Mp_3653_BCZ38. (COI: KJ526794, go to Genbank) ditto.

Mp_3653_BCZ40. (COI: KJ526795, go to Genbank) ditto.

Finland

CIIMAR_BIV3480. (COI: OR841771, go to Genbank) Kokemäenjoki River (Kokemäenjoki Basin), Finland (61.56193, 21.66649).

CIIMAR_BIV3481. (COI: OR841772, go to Genbank) ditto.

CIIMAR_BIV3482. (COI: OR841773, go to Genbank) ditto.

CIIMAR_BIV3483. (COI: OR841774, go to Genbank) ditto.

CIIMAR_BIV3484. (COI: OR841775, go to Genbank) ditto.

France

MNHN_IM-2009-17754. (16S: KC703451, 28S: KC703696, COI: KC703897, go to Genbank) Aisne River (Seine Basin), France (49.475847, 4.589161).

MNHN_IM-2009-17755. (16S: KC703452, 28S: KC703697, COI: KC703898, go to Genbank) ditto.

MNHN_IM-2009-17763. (16S: KC703458, 28S: KC703703, COI: KC703900, go to Genbank) ditto.

MNHN_IM-2009-17792. (16S: KC703472, 28S: KC703717, COI: KC703901, go to Genbank) ditto.

MNHN_IM-2009-17796. (16S: KC703476, 28S: KC703721, COI: KC703905, go to Genbank) ditto.

MNHN_IM-2009-22398. (16S: KC703613, 28S: KC703862, COI: KC703961, go to Genbank) ditto.

MNHN_IM-2009-17761. (16S: KC703457, 28S: KC703702, go to Genbank) France: Aisne River.

MNHN_IM-2009-17766. (16S: KC703461, 28S: KC703706, go to Genbank) ditto.

MNHN_IM-2009-17790. (16S: KC703470, 28S: KC703715, go to Genbank) ditto.

MNHN_IM-2009-17805. (16S: KC703485, 28S: KC703730, go to Genbank) ditto.

MNHN_IM-2009-12721. (16S: KC703435, 28S: KC703680, go to Genbank) France: Aubetin River.

MNHN_IM-2009-12617. (16S: KC703382, 28S: KC703641, go to Genbank) France: Boivre River.

MNHN_IM-2009-21230. (16S: KC703538, 28S: KC703787, go to Genbank) France: Ourcq River.

MNHN_IM-2009-21213. (16S: KC703523, 28S: KC703771, COI: KC703926, go to Genbank) La Bruche Canal (Rhine Basin), France (48.579961, 7.678739).

MNHN_IM-2009-12720. (16S: KC703434, 28S: KC703679, COI: KC703895, go to Genbank) Lunain River (Seine Basin), France (48.2812, 2.7954).

MNHN_IM-2009-21219. (16S: KC703529, 28S: KC703777, COI: KC703928, go to Genbank) Ourcq River (Seine Basin), France (49.186301, 3.360753).

MNHN_IM-2009-21221. (16S: KC703530, 28S: KC703779, COI: KC703929, go to Genbank) ditto.

MNHN_IM-2009-21227. (16S: KC703535, 28S: KC703784, COI: KC703930, go to Genbank) ditto.

MNHN_IM-2009-21232. (16S: KC703540, 28S: KC703789, COI: KC703931, go to Genbank) ditto.

MNHN_IM-2009-12677. (16S: KC703403, COI: KC703890, go to Genbank) Petit Gland River (Seine Basin), France (49.9059, 4.1585).

MNHN_IM-2009-12680. (16S: KC703405, 28S: KC703650, COI: KC703891, go to Genbank) ditto.

MNHN_IM-2009-17769. (16S: KC703516, 28S: KC703763, COI: KC703922, go to Genbank) ditto.

MNHN_IM-2009-17842. (16S: KC703508, 28S: KC703756, COI: KC703914, go to Genbank) ditto.

MNHN_IM-2009-17848. (16S: KC703514, 28S: KC703762, COI: KC703920, go to Genbank) ditto.

MNHN_IM-2009-17811. (16S: KC703491, 28S: KC703736, COI: KC703909, go to Genbank) Therain River (Seine Basin), France (49.50631, 1.971917).

MNHN_IM-2009-12564. (16S: KC703351, 28S: KC703622, COI: JX046553, go to Genbank) Vienne River (Loire Basin), France (47.017062, 0.563053).

MNHN_IM-2009-12566. (16S: KC703352, COI: KC703870, go to Genbank) ditto.

MNHN_IM-2009-12595. (16S: KC703369, 28S: KC703634, COI: KC703872, go to Genbank) ditto.

MNHN_IM-2009-12616. (16S: KC703381, COI: KC703878, go to Genbank) ditto.

MNHN_IM-2009-12629. (16S: KC703385, 28S: KC703644, COI: KC703879, go to Genbank) ditto.

MNHN_IM-2009-12630. (16S: KC703386, 28S: KC703645, COI: KC703880, go to Genbank) ditto.

Germany

TUM_AI02. (COI: OR841787, go to Genbank) Aisch River (Rhine Basin), Germany (49.70439, 10.854872).

TUM_AI03. (COI: OR841788, go to Genbank) ditto.

CIIMAR_BIV4202. (COI: OR841789, go to Genbank) Alte Schwentine River (Schlei/Trave Basin), Germany (54.185686, 10.241204).

CIIMAR_BIV4203. (COI: OR841790, go to Genbank) ditto.

CIIMAR_BIV4204. (COI: OR841791, go to Genbank) ditto.

CIIMAR_BIV4205. (COI: OR841792, go to Genbank) ditto.

CIIMAR_BIV4206. (COI: OR841793, go to Genbank) ditto.

CIIMAR_BIV4192. (COI: OR841777, go to Genbank) Bollingstedter Au River (Eider Basin), Germany (54.588631, 9.433135).

CIIMAR_BIV4193. (COI: OR841778, go to Genbank) ditto.

CIIMAR_BIV4194. (COI: OR841779, go to Genbank) ditto.

CIIMAR_BIV4195. (COI: OR841780, go to Genbank) ditto.

CIIMAR_BIV4196. (COI: OR841781, go to Genbank) ditto.

CIIMAR_BIV4197. (COI: OR841782, go to Genbank) Ilmenau River (Elbe Basin), Germany (53.148217, 10.467698).

CIIMAR_BIV4198. (COI: OR841783, go to Genbank) ditto.

CIIMAR_BIV4199. (COI: OR841784, go to Genbank) ditto.

CIIMAR_BIV4200. (COI: OR841785, go to Genbank) ditto.

CIIMAR_BIV4201. (COI: OR841786, go to Genbank) ditto.

TUM_RG03. (COI: OR841776, go to Genbank) Regen River (Danube Basin), Germany (49.211311, 12.218066).

Hungary

TUM_UC02. (COI: OR841794, go to Genbank) Ipoly River (Danube Basin), Hungary (47.930964, 18.771094).

TUM_UC03. (COI: OR841795, go to Genbank) ditto.

TUM_UC04. (COI: OR841796, go to Genbank) ditto.

TUM_UC06. (COI: OR841797, go to Genbank) ditto.

Kazakhstan

RMBH_biv_595_1. (COI: MK603933, go to Genbank) Kazakhstan: Ural River (50.7131°N, 51.3808 °E), Caspian Sea drainage.

RMBH_biv_595_2. (COI: MK603934, go to Genbank) ditto.

RMBH_biv_595_3. (COI: MK603935, go to Genbank) ditto.

RMBH_biv_595_4. (COI: MK603936, go to Genbank) ditto.

Latvia

TUM_DG04. (COI: OR841798, go to Genbank) Düna River (Western Dvina Basin), Latvia (55.89361, 26.875624).

TUM_DG05. (COI: OR841799, go to Genbank) ditto.

TUM_DG06. (COI: OR841800, go to Genbank) ditto.

TUM_DG07. (COI: OR841801, go to Genbank) ditto.

TUM_DG08. (COI: OR841802, go to Genbank) ditto.

TUM_IN01. (COI: OR841803, go to Genbank) Indrica River (Western Dvina Basin), Latvia (55.845895, 27.413246).

TUM_IN02. (COI: OR841804, go to Genbank) ditto.

TUM_IN05. (COI: OR841805, go to Genbank) ditto.

TUM_IN06. (COI: OR841806, go to Genbank) ditto.

TUM_IN07. (COI: OR841807, go to Genbank) ditto.

Lithuania

CIIMAR_BIV4401. (COI: OR841814, go to Genbank) Jūra River (Neman Basin), Lithuania (55.540367, 21.993161).

CIIMAR_BIV4402. (COI: OR841815, go to Genbank) ditto.

CIIMAR_BIV4403. (COI: OR841816, go to Genbank) ditto.

CIIMAR_BIV4404. (COI: OR841817, go to Genbank) ditto.

CIIMAR_BIV4405. (COI: OR841818, go to Genbank) ditto.

CIIMAR_BIV4406. (COI: OR841819, go to Genbank) ditto.

CIIMAR_BIV4461. (COI: OR841808, go to Genbank) Nemunėlis River/Lake (Lielupe Basin), Lithuania (56.151583, 25.139853).

CIIMAR_BIV4462. (COI: OR841809, go to Genbank) ditto.

CIIMAR_BIV4463. (COI: OR841810, go to Genbank) ditto.

CIIMAR_BIV4464. (COI: OR841811, go to Genbank) ditto.

CIIMAR_BIV4465. (COI: OR841812, go to Genbank) ditto.

CIIMAR_BIV4466. (COI: OR841813, go to Genbank) ditto.

CIIMAR_BIV4391. (COI: OR841820, go to Genbank) Ūla River (Neman Basin), Lithuania (54.046133, 24.623964).

CIIMAR_BIV4392. (COI: OR841821, go to Genbank) ditto.

CIIMAR_BIV4393. (COI: OR841822, go to Genbank) ditto.

CIIMAR_BIV4394. (COI: OR841823, go to Genbank) ditto.

CIIMAR_BIV4395. (COI: OR841824, go to Genbank) ditto.

CIIMAR_BIV4396. (COI: OR841825, go to Genbank) ditto.

Poland

CIIMAR_BIV3560. (COI: OR841826, go to Genbank) Blizna River (Neman Basin), Poland (53.923814, 23.050983).

CIIMAR_BIV3561. (COI: OR841827, go to Genbank) ditto.

CIIMAR_BIV3562. (COI: OR841828, go to Genbank) ditto.

CIIMAR_BIV3563. (COI: OR841829, go to Genbank) ditto.

CIIMAR_BIV3564. (COI: OR841830, go to Genbank) ditto.

CIIMAR_BIV3554. (COI: OR841836, go to Genbank) Brzeźniczanka River (Oder Basin), Poland (51.778672, 15.271731).

CIIMAR_BIV3520. (COI: OR841842, go to Genbank) Bug River (Vistula Basin), Poland (52.363106, 22.871444).

CIIMAR_BIV3521. (COI: OR841843, go to Genbank) ditto.

CIIMAR_BIV3522. (COI: OR841844, go to Genbank) ditto.

CIIMAR_BIV3523. (COI: OR841845, go to Genbank) ditto.

CIIMAR_BIV3524. (COI: OR841846, go to Genbank) ditto.

Mp_4376_UcF3. (COI: KF515713, go to Genbank) Czarna Hancza River/Lake (Neman Basin), Poland (53.970883, 23.303617).

Mp_4376_UcF4. (COI: KF515714, go to Genbank) ditto.

Mp_4376_UcF5. (COI: KF515715, go to Genbank) ditto.

Mp_4376_UcF2. (COI: KF515719, go to Genbank) Drawa River/Lake (Oder Basin), Poland (53.495583, 15.809933).

CIIMAR_BIV3530. (COI: OR841837, go to Genbank) Slupia River (Slupia Basin), Poland (54.357033, 17.153969).

CIIMAR_BIV3531. (COI: OR841838, go to Genbank) ditto.

CIIMAR_BIV3532. (COI: OR841839, go to Genbank) ditto.

CIIMAR_BIV3533. (COI: OR841840, go to Genbank) ditto.

CIIMAR_BIV3534. (COI: OR841841, go to Genbank) ditto.

TUM_PSP01. (COI: OR841831, go to Genbank) Szeszupa River (Neman Basin), Poland (54.2576, 22.89388).

TUM_PSP02. (COI: OR841832, go to Genbank) ditto.

TUM_PSP03. (COI: OR841833, go to Genbank) ditto.

TUM_PSP04. (COI: OR841834, go to Genbank) ditto.

TUM_PSP05. (COI: OR841835, go to Genbank) ditto.

Mp_4376_UcF1. (COI: KF515718, go to Genbank) Szeszupa River/Lake (Neman Basin), Poland (54.271467, 22.9117).

Mp_4376_UcF6. (COI: KF515716, go to Genbank) Warkocz River/Lake (Vistula Basin), Poland (50.840236, 20.757725).

Mp_4376_UcF7. (COI: KF515717, go to Genbank) ditto.

Romania

CIIMAR_BIV8437. (COI: OR841853, go to Genbank) Caraș River (Danube Basin), Romania (45.080957, 21.548143).

CIIMAR_BIV8438. (COI: OR841854, go to Genbank) ditto.

CIIMAR_BIV8439. (COI: OR841855, go to Genbank) ditto.

CIIMAR_BIV8440. (COI: OR841856, go to Genbank) ditto.

CIIMAR_BIV8441. (COI: OR841857, go to Genbank) ditto.

CIIMAR_BIV8734. (COI: OR841858, go to Genbank) Prut River (Danube Basin), Romania (46.24386, 28.110237).

CIIMAR_BIV8736. (COI: OR841859, go to Genbank) ditto.

CIIMAR_BIV8738. (COI: OR841860, go to Genbank) ditto.

TUM_RSM01. (COI: OR841847, go to Genbank) Someș River (Danube Basin), Romania (47.631908, 23.377935).

TUM_RSM02. (COI: OR841848, go to Genbank) ditto.

TUM_RSM03. (COI: OR841849, go to Genbank) ditto.

TUM_RSM04. (COI: OR841850, go to Genbank) ditto.

TUM_RSM05. (COI: OR841851, go to Genbank) ditto.

CIIMAR_BIV4171. (COI: OR841852, go to Genbank) Talna River (Danube Basin), Romania (47.888876, 23.230259).

Russia

RMBH_biv_304_10. (COI: OR841866, go to Genbank) Iren River (Volga Basin), Russia (57.416828, 56.949358).

RMBH_biv_304_9. (COI: OR841867, go to Genbank) ditto.

RMBH_biv_280_1. (COI: OR841863, SRA: SRR26935083, go to Genbank) Khopyor River (Don Basin), Russia (51.487836, 42.593142).

RMBH_biv_280_10. (COI: OR841864, go to Genbank) ditto.

RMBH_biv_280_6. (COI: OR841865, go to Genbank) ditto.

RMBH_biv_196_5. (COI: OR841861, go to Genbank) Khopyor River (Don Basin), Russia (51.580074, 43.182045).

RMBH_biv_196_6. (COI: OR841862, go to Genbank) ditto.

RMBH_biv_304_5. (16S: MK574407, 28S: MK574548, COI: MK574271, go to Genbank) Russia: European part, Perm Region, Iren River (57.4167°N, 56.9496°E), Volga Basin, Caspian Sea drainage.

RMBH_biv_304_7. (16S: MK574408, 28S: MK574549, COI: MK574272, go to Genbank) ditto.

RMBH_biv_304_8. (16S: MK574409, 28S: MK574550, COI: MK574273, go to Genbank) ditto.

RMBH_biv_305_1. (16S: MK574410, 28S: MK574551, COI: MK574274, go to Genbank) Russia: European part, Samara Region, Samara River (53.0068°N, 50.7768°E), Volga Basin, Caspian Sea drainage.

RMBH_biv_305_2. (16S: MK574411, 28S: MK574552, COI: MK574275, go to Genbank) ditto.

RMBH_biv_305_3. (16S: MK574412, 28S: MK574553, COI: MK574276, go to Genbank) ditto.

RMBH_biv_196_1. (16S: MK574401, 28S: MK574542, COI: MK574265, go to Genbank) Russia: European part, Saratov Region, Khopyor River (51.5630°N, 43.1530°E), Don Basin, Azov Sea drainage.

RMBH_biv_196_2. (16S: MK574402, 28S: MK574543, COI: MK574266, go to Genbank) ditto.

RMBH_biv_196_3. (16S: MK574403, 28S: MK574544, COI: MK574267, go to Genbank) ditto.

RMBH_biv_280_3. (16S: MK574404, 28S: MK574545, COI: MK574268, go to Genbank) Russia: European part, Voronezh Region, Khopyor River (51.4878°N, 42.5931°E), Don Basin, Azov Sea drainage.

RMBH_biv_280_4. (16S: MK574405, 28S: MK574546, COI: MK574269, go to Genbank) ditto.

RMBH_biv_280_5. (16S: MK574406, 28S: MK574547, COI: MK574270, go to Genbank) ditto.

Mp_3904_Cr1. (16S: MH544931, ITS1: MH544886, go to Genbank) Russia: River Ivitza, Volga basin, Rameshkovsky District, Tver Oblast.

Mp_3904_Cr11. (16S: MH544928, 5.8S: MH544889, go to Genbank) ditto.

Mp_3904_Cr17. (16S: MH544927, 5.8S: MH544890, ITS1: MH544890, go to Genbank) ditto.

Mp_3904_Cr24. (16S: MH544926, ITS1: MH544891, go to Genbank) ditto.

Mp_3904_Cr27. (16S: MH544917, ITS1: MH544883, go to Genbank) ditto.

Mp_3904_Cr3. (16S: MH544930, 5.8S: MH544887, ITS1: MH544887, go to Genbank) ditto.

Mp_3904_Cr32. (16S: MH544918, 5.8S: MH544884, COI: MH539794, ITS1: MH544884, go to Genbank) ditto.

Mp_3904_Cr33. (16S: MH544916, 5.8S: MH544885, COI: MH539797, ITS1: MH544885, go to Genbank) ditto.

Mp_3904_Cr34. (16S: MH544925, ITS1: MH544892, go to Genbank) ditto.

Mp_3904_Cr41. (16S: MH544924, 5.8S: MH544893, ITS1: MH544893, go to Genbank) ditto.

Mp_3904_Cr48. (16S: MH544915, COI: MH539796, go to Genbank) ditto.

Mp_3904_Cr49. (16S: MH544923, 5.8S: MH544894, ITS1: MH544894, go to Genbank) ditto.

Mp_3904_Cr51. (16S: MH544922, 5.8S: MH544895, ITS1: MH544895, go to Genbank) ditto.

Mp_3904_Cr52. (16S: MH544921, 5.8S: MH544896, ITS1: MH544896, go to Genbank) ditto.

Mp_3904_Cr53. (16S: MH544920, 5.8S: MH544897, ITS1: MH544897, go to Genbank) ditto.

Mp_3904_Cr7. (16S: MH544929, ITS1: MH544888, go to Genbank) ditto.

Mp_3904_Cr8. (16S: MH544919, 5.8S: MH544882, COI: MH539795, ITS1: MH544882, go to Genbank) ditto.

RMBH_biv_305_4. (COI: OR841868, go to Genbank) Samara River (Volga Basin), Russia (53.006977, 50.776563).

RMBH_biv_305_5. (COI: OR841869, go to Genbank) ditto.

RMBH_biv_305_6. (COI: OR841870, go to Genbank) ditto.

RMBH_biv_305_7. (COI: OR841871, go to Genbank) ditto.

RMBH_biv_305_8. (COI: OR841872, go to Genbank) ditto.

RMBH_biv_305_9. (COI: OR841873, go to Genbank) ditto.

Sweden

TUM_F5. (COI: OR841879, go to Genbank) Bråån River (Kävlingeåns Basin), Sweden (55.800696, 13.64148).

TUM_F6. (COI: OR841880, go to Genbank) ditto.

TUM_F7. (COI: OR841881, go to Genbank) ditto.

TUM_F8. (COI: OR841882, go to Genbank) ditto.

TUM_F9. (COI: OR841883, go to Genbank) ditto.

CIIMAR_BIV3410. (COI: OR841875, go to Genbank) Gardvedaan River (Eman Basin), Sweden (57.352863, 15.820028).

CIIMAR_BIV3411. (COI: OR841876, go to Genbank) ditto.

CIIMAR_BIV3412. (COI: OR841877, go to Genbank) ditto.

CIIMAR_BIV3413. (COI: OR841878, go to Genbank) ditto.

CIIMAR_BIV3425. (COI: OR841884, go to Genbank) Storan River (Storan Basin), Sweden (58.021804, 15.652948).

CIIMAR_BIV3426. (COI: OR841885, go to Genbank) ditto.

CIIMAR_BIV3427. (COI: OR841886, SRA: SRR26935081, go to Genbank) ditto.

CIIMAR_BIV3428. (COI: OR841887, go to Genbank) ditto.

CIIMAR_BIV3429. (COI: OR841888, go to Genbank) ditto.

CIIMAR_BIV3449. (COI: OR841889, go to Genbank) Svartaan River (Svartaan Basin), Sweden (58.86938, 17.121288).

CIIMAR_BIV3450. (COI: OR841890, go to Genbank) ditto.

CIIMAR_BIV3451. (COI: OR841891, go to Genbank) ditto.

CIIMAR_BIV3452. (COI: OR841892, go to Genbank) ditto.

CIIMAR_BIV3453. (COI: OR841893, go to Genbank) ditto.

NRM_42648-M099. (16S: DQ060162, COI: DQ060174, go to Genbank) Sweden.

CIIMAR_BIV3409. (16S: MZ519056, 18S: PP693707, 28S: PP693722, COI: OR841874, mtGenome: OR912547, go to Genbank) Sweden, Gardvedaan River (Eman Basin), Sweden (57.352863, 15.820028).

TUM_T10. (COI: OR841894, go to Genbank) Tommarpsån River (Tommarpsån Basin), Sweden (55.554078, 14.149254).

TUM_T11. (COI: OR841895, go to Genbank) ditto.

TUM_T13. (COI: OR841896, go to Genbank) ditto.

TUM_T14. (COI: OR841897, go to Genbank) ditto.

TUM_T15. (COI: OR841898, go to Genbank) ditto.

Ukraine

ZSU-U13. (COI: KY930360, go to Genbank) Desna River (Dnipro Basin), Ukraine (51.907434, 33.215528).

ZSU-U11. (COI: KY930359, go to Genbank) Stalineshty River (Danube Basin), Ukraine (48.25755, 26.543876).

CIIMAR_BIV3510. (COI: OR841899, go to Genbank) Stryi River (Dniester Basin), Ukraine (49.215956, 23.298919).

CIIMAR_BIV3511. (COI: OR841900, go to Genbank) ditto.

CIIMAR_BIV3512. (COI: OR841901, go to Genbank) ditto.

CIIMAR_BIV3513. (COI: OR841902, go to Genbank) ditto.

CIIMAR_BIV3514. (COI: OR841903, go to Genbank) ditto.

CIIMAR_BIV3500. (COI: OR841904, go to Genbank) Styr River (Dnipro Basin), Ukraine (50.171344, 24.896869).

CIIMAR_BIV3501. (COI: OR841905, go to Genbank) ditto.

CIIMAR_BIV3502. (COI: OR841906, go to Genbank) ditto.

CIIMAR_BIV3503. (COI: OR841907, SRA: SRR26935080, go to Genbank) ditto.

CIIMAR_BIV3504. (COI: OR841908, go to Genbank) ditto.