Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Unionini | Unionina | Unio

species Unio crassus Philipsson in Retzius, 1788: specimens

112 available nominal species synonyms (190 total synonyms)

Recent Taxonomic Opinions

Unio crassus — Graf (2007), Graf & Cummings (2007), Manendo et al. (2008), Dhora (2009), Araujo et al. (2009), ... Wu et al. (2025), Gorbunova et al. (2025), Nagel (2025), Campbell & Lydeard (2026), Prie et al. (2026)
Unio crassus gontieri — Graf (2007)
Pleurobema sintoxia — Williams et al. (2008)
Crassiana crassa crassa — Kantor et al. (2010)
Crassiana cyprinorum — Kantor et al. (2010)
Crassiana irenjensis — Kantor et al. (2010)
Crassiana irgizlaica — Kantor et al. (2010)
Crassiana musiva musiva — Kantor et al. (2010)
“Unio crassus” ** — Soroka (2010), Reis et al. (2013), Douda et al. (2014), Froufe et al. (2016), Burzynski et al. (2017), ... Chen et al. (2025), Klymus et al. (2025), Wu et al. (2026), Campbell & Lydeard (2026), Wu et al. (2026)
Unio crassus crassus — Fauna Europaea (2012), Prié & Puillandre (2014), Klishko et al. (2017), Bogatov (2022)
Unio crassus batavus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio mancus mancus — Prie (2012)
Unio mancus moquinianus — Prie (2012)
Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017)
“Unio crassus crassus” ** — Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016)
Unio (Crassiana) irenjensis — Vinarski & Kantor (2016)
Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016)
Unio (Crassiana) musivus — Vinarski & Kantor (2016)
Unio crassus cf. courtillieri — Klishko et al. (2017)
Unio crassus courtillierii [sic] — Araujo et al. (2018)
“Unio crassus” ** — Araujo et al. (2018), Lyubas et al. (2022)
“Unio crassus” ** — Araujo et al. (2018)
Unio c. crassus — Príe et al. (2021)
Unio c. courtillieri — Príe et al. (2021)
Unio cf. courtillieri — Lopes-Lima et al. (2021)
Unio crassus musivus — Bogatov (2022)

Range

Widespread in continental Europe through the Dardanelles to Transcaucasia. (North Eurasia: EU1)

Map of Georeferenced Specimens

specimens map of Unio crassus

MUSSELp Specimen Records

Type Specimens

image of lot MNHN_22098
MNHN_22098
Le Mans (Sarthe)
lectotype
Unio ampullaceus Locard, 1893

image of lot MNHN_22100
MNHN_22100
La Maine à Angers
possible syntype
Unio andegavensis Locard, 1882

image of lot MNHN_22101
MNHN_22101
le Gouet, à Saint-Brieue (Côtes-du-Nord)
lectotype
Unio andeliacus Bourguignat in Locard, 1889

image of lot MNHN_22110
MNHN_22110
Basse Indre (Loire-Inf.)
idiotype
Unio arenarum Bourguignat in Locard, 1882

image of lot MNHN_22118
MNHN_22118
Bahus (Landes)
holotype
Unio aturicus Locard, 1889

image of lot MNHN_22122
MNHN_22122
La Loire, à Balbigny (Loire)
lectotype
Unio balbignyanus Locard, 1889

image of lot MNHN_22127
MNHN_22127
La Bouille, près de Rouen (Seine-Inf.)
lectotype
Unio batavellus Locard, 1885

image of lot MNHN_22132
MNHN_22132
La Marne, à Charenton (Seine)
idiotype
Unio berthelini Bourguignat in Locard, 1882

image of lot MNHN_22133
MNHN_22133
La Loire, à Saint-Gemmes (Maine-et-Loire)
lectotype
Unio besnardianus Bourguignat in Locard, 1889

image of lot MNHN_22134
MNHN_22134
La Saône (Côte-d’Or)
paralectotype
Unio besnardianus Bourguignat in Locard, 1889

image of lot MNHN_22145
MNHN_22145
La Canne, à Goulnot
lectotype
Unio brevierei Locard, 1882

image of lot MNHN_22162
MNHN_22162
[illegible] (Charente)
idiotype
Unio carantoni Bourguignat in Locard, 1882

image of lot MNHN_22171
MNHN_22171
La Marne, à Charenton (Seine)
lectotype
Unio catalaunicus Bourguignat in Locard, 1889

image of lot MNHN_22182
MNHN_22182
La Canne, à Jailly (Nièvre)
lectotype
Unio crassatellus Bourguignat in Locard, 1882

image of lot MNHN-IM-2000-1666
MNHN-IM-2000-1666
Saone
syntype
Unio crassulus Drouët, 1888

image of lot MNHN_22185
MNHN_22185
l’Ornain (Meuse)
idiotype
Unio cyprinorum Bourguignat in Locard, 1882

image of lot MNHN_22192
MNHN_22192
La Loire à Ingrandes
possible syntype
Unio diptychus Bourguignat in Locard, 1889

image of lot MNHN_22215
MNHN_22215
La Moselle
possible syntype
Unio feliciani Bourguignat in Locard, 1882

image of lot MNHN_22217
MNHN_22217
La Marne à Châlons (Marne)
lectotype
Unio financei Locard, 1889

image of lot MNHN_22241
MNHN_22241
La Loire à Nantes (Loire-Infereur)
lectotype
Unio gliscerus Locard, 1893

image of lot MNHN_22254
MNHN_22254
La Loire, à Basse-Indre
syntype
Unio hattemani Bourguignat in Locard, 1889

image of lot MNHN_22255
MNHN_22255
La Loire à Nantes (Loire-Infereur)
syntype
Unio hattemani Bourguignat in Locard, 1889

image of lot MNHN_22256
MNHN_22256
La Loire, à Saint-Gondon (Loiret)
syntype
Unio hattemani Bourguignat in Locard, 1889

image of lot MNHN_22282
MNHN_22282
Ingrandes (Maine-et-Loire)
possible syntype
Unio ingrandiensis Bourguignat in Locard, 1889

image of lot MNHN_22303
MNHN_22303
Condal (Saône-et-Loire)
syntype
Unio lagnisicus Bourguignat in Locard, 1882

image of lot MNHN_22304
MNHN_22304
La Drée (Saône-et-Loire)
syntype
Unio lagnisicus Bourguignat in Locard, 1882

image of lot MNHN_22305
MNHN_22305
le Seine au nord d. Rouen
syntype
Unio lancelevei Locard, 1893

image of lot MNHN_22306
MNHN_22306
le Reyran (Var)
lectotype
Unio lemotheuxi Bourguignat in Locard, 1889

image of lot MNHN_22310
MNHN_22310
La Loire à Ingrandes
idiotype
Unio ligericus Bourguignat in Locard, 1882

image of lot MNHN_22316
MNHN_22316
l’Albane (Côte-d’Or)
idiotype
Unio locardianus Bourguignat in Locard, 1882

image of lot MNHN_22342
MNHN_22342
La Loire à Nantes (Loire-Infereur)
idiotype
Unio mariae Bourguignat in Locard, 1889

image of lot MNHN_22347
MNHN_22347
la Somme à Abbeville
lectotype
Unio materniacus Locard, 1889

image of lot MNHN_22355
MNHN_22355
La Marne à Lagny (Seine-et-Marne)
lectotype
Unio matronicus Bourguignat in Locard, 1882

image of lot MNHN_22360
MNHN_22360
La Loire à Balbigny (Loire)
lectotype
Unio melantatus Locard, 1889

image of lot MNHN_22438
MNHN_22438
La Canne, dans la Nièvre
lectotype
Unio oxyrhynchus Bourguignat in Locard, 1882

image of lot MNHN_22460
MNHN_22460
Passavant (Haute-Saône)
idiotype
Unio pilloti Bourguignat in Locard, 1882

image of lot MNHN_22483
MNHN_22483
Gigny (Jura)
lectotype
Unio rayi Bourguignat in Locard, 1882

image of lot MNHN_22523
MNHN_22523
La Loire à Orléans
lectotype
Unio seneauxi Bourguignat in Locard, 1889

image of lot MNHN_22528
MNHN_22528
le Mandrezey, à Saulcy-sur-Meurthe (Vosges)
idiotype
Unio septentrionalis Bourguignat in Locard, 1882

image of lot MNHN_22529
MNHN_22529
La Marne à Lagny (Seine-et-Marne)
lectotype
Unio sequanicus Bourguignat in Locard, 1882

image of lot MNHN_22559
MNHN_22559
La Saône, à Auxonne (Côte-d’Or)
syntype
Unio subamnicus Locard, 1889

image of lot MNHN_22569
MNHN_22569
La Loire à Ingrandes
syntype
Unio surraulti Bourguignat in Locard, 1889

image of lot MNHN_22570
MNHN_22570
La Loire à Nantes (Loire-Infereur)
syntype
Unio surraulti Bourguignat in Locard, 1889

MNHN_22099. Le Mans (Sarthe) — lectotype Unio ampullaceus Locard, 1893

MNHN_22123. La Loire, à Balbigny (Loire) — lectotype Unio balbignyanus Locard, 1889

MNHN_MP_2453. La Loire, à Balbigny (Loire) / la Canne, Nievre — lectotype Unio balbignyanus Locard, 1889

MNHN_22124. Saint-Germain-des Fossés — lectotype Unio balbignyanus Locard, 1889

MNHN_22125. le Mandrezey, à Saulcy-sur-Meurthe (Vosges) — lectotype Unio balbignyanus Locard, 1889

MNHN_22126. La Loire à Villerest (Loire) — lectotype Unio balbignyanus Locard, 1889

MNHN_22128. a Grosne, à La Ferté — lectotype Unio batavellus Locard, 1885

MNHN_22147. Marais de Dampierre — lectotype Unio brevierei Locard, 1882

MNHN_22146. La Canne à Saint-Saulge — lectotype Unio brevierei Locard, 1882

MNHN_22183. La Drée, à Épinac (Saône-et-Loire) — lectotype Unio crassatellus Bourguignat in Locard, 1882

MNHN_22184. La Drée (Saône-et-Loire) — lectotype Unio crassatellus Bourguignat in Locard, 1882

MNHN_22218. L’Eure, près d’Évreux — idiotype Unio financei Locard, 1889

MNHN_22219. Domsure (Jura) — idiotype Unio financei Locard, 1889

MNHN_22242. La Loire à Nantes (Loire-Infereur) — lectotype Unio gliscerus Locard, 1893

MNHN_22243. La Saône à Lyon [Rhône] — lectotype Unio gliscerus Locard, 1893

MNHN_22244. La Grosne, à la Ferté (Saône-et-Loire) — lectotype Unio gliscerus Locard, 1893

MNHN_22245. La Grosne, à Marnay (Saône-et-Loire) — lectotype Unio gliscerus Locard, 1893

MNHN_22307. La Loire, à Nantes — lectotype Unio lemotheuxi Bourguignat in Locard, 1889

MNHN_22348. la Seine, à Corbeil — lectotype Unio materniacus Locard, 1889

MNHN_22349. la Seine, à Charenton — lectotype Unio materniacus Locard, 1889

MNHN_22350. Pontoise — lectotype Unio materniacus Locard, 1889

MNHN_22351. le Meurthe à Lunéville — lectotype Unio materniacus Locard, 1889

MNHN_22352. l’Oise, à Creil — lectotype Unio materniacus Locard, 1889

MNHN_22353. l’Oise, à Compiègne — lectotype Unio materniacus Locard, 1889

MNHN_22354. l’Ornoise (Meuse) — lectotype Unio materniacus Locard, 1889

MNHN_22356. La Marne à Lagny (Seine-et-Marne) — lectotype Unio matronicus Bourguignat in Locard, 1882

MNHN_22361. La Loire à Balbigny (Loire) — lectotype Unio melantatus Locard, 1889

MNHN_22362. La Loire à Villerest (Loire) — lectotype Unio melantatus Locard, 1889

MNHN_22484. St-Amour (Jura) — lectotype Unio rayi Bourguignat in Locard, 1882

MNHN_22524. La Loire à Nantes (Loire-Infereur) — lectotype Unio seneauxi Bourguignat in Locard, 1889

MNHN_22525. La Grosne, à Marnay (Saône-et-Loire) — lectotype Unio seneauxi Bourguignat in Locard, 1889

MNHN_22530. La Marne à Lagny (Seine-et-Marne) — lectotype Unio sequanicus Bourguignat in Locard, 1882

MNHN_22560. La Marne à Lagny (Seine-et-Marne) — syntype Unio subamnicus Locard, 1889

MNHN_22561. le Grosne, à Marnay (Saône-et-Loire) — syntype Unio subamnicus Locard, 1889

Other Specimens by Country | show by collection

1 Specimens from Bosnia and Herzegovina

INHS_8376. Lasva

5 Specimens from France

INHS_21545. France

INHS_21552. Hungary

INHS_21549. Saône River, Lyon, Rhone, Rhône-Alpes, France

INHS_21550. Saône River, Saone, Rhône-Alpes, France

INHS_21553. Trencsin, Hungary

7 Specimens from Germany

image of lot KON_MP_02
KON_MP_02
Sulzbach River (a small tributary of the Rhine), Sulzbach upstream of Stollhofen (37 km SW of Krlsruhe) 08°04'04"E, 48°45'11"N

INHS_21558. Germany

INHS_21560. Nister River, Nister, Rhineland-Palatinate, Germany

INHS_21561. Oberschlesien [Upper Silesia], Germany

INHS_21557. Oder River, Vizenot Oppehr. [?], Germany

INHS_21559. Ottmachari, Silesia, Germany

INHS_21665. Schleswig- Holstein, Germany

1 Specimens from Romania

INHS_21566. Transylvania, Romania

3 Specimens from Switzerland

INHS_21563. Biwir Creek, Switzerland

INHS_21564. Geneva, Marais de Lionett [Lionett Marsh], Switzerland

INHS_21565. Lake Leman, Villeneuve, Vaud, Switzerland

2 Specimens from Ukraine

INHS_22459. Pzisluch [Sluch] River, Rivne Region, Rivne Oblast, Ukraine

INHS_22460. Ubort River, near Olevsk, Zhytomyr Region, Zhytomyr Oblast, Ukraine

GenBank Vouchers

Georeferenced GenBank Vouchers

Austria

CIIMAR_BIV4658. (COI: OR841742, go to GenBank) Aschach River (Danube Basin), Austria (48.321463, 14.013764).

CIIMAR_BIV4659. (COI: OR841743, go to GenBank) ditto.

CIIMAR_BIV4660. (COI: OR841744, go to GenBank) ditto.

CIIMAR_BIV4661. (COI: OR841745, go to GenBank) ditto.

Belarus

RMBH_biv_1230_1. (COI: ON648805, go to GenBank) Belarus: Ross, Neman Basin (53.0442, 24.4229).

Belgium

TUM_GH02. (COI: OR841750, go to GenBank) Grande Honelles River (Scheldt Basin), Belgium (50.384939, 3.687902).

TUM_GH03. (COI: OR841751, go to GenBank) ditto.

TUM_GH04. (COI: OR841752, go to GenBank) ditto.

TUM_GH05. (COI: OR841753, go to GenBank) ditto.

TUM_ME01. (COI: OR841746, go to GenBank) Mehaigne River (Meuse Basin), Belgium (50.586811, 5.189187).

TUM_ME03. (COI: OR841747, go to GenBank) ditto.

TUM_ME04. (COI: OR841748, go to GenBank) ditto.

TUM_ME05. (COI: OR841749, go to GenBank) ditto.

Bulgaria

CIIMAR_BIV7928. (COI: OR841759, go to GenBank) Dyavolska River (Dyavolska Basin), Bulgaria (42.26879, 27.66478).

CIIMAR_BIV7929. (COI: OR841760, go to GenBank) ditto.

CIIMAR_BIV7930. (COI: OR841761, go to GenBank) ditto.

CIIMAR_BIV7931. (COI: OR841762, go to GenBank) ditto.

CIIMAR_BIV7768. (COI: OR841754, go to GenBank) Osam River (Danube Basin), Bulgaria (43.40044, 25.1551).

CIIMAR_BIV7769. (COI: OR841755, go to GenBank) ditto.

CIIMAR_BIV7770. (COI: OR841756, go to GenBank) ditto.

CIIMAR_BIV7771. (COI: OR841757, go to GenBank) ditto.

CIIMAR_BIV7772. (COI: OR841758, go to GenBank) ditto.

Croatia

CIIMAR_BIV6093. (COI: OR841763, go to GenBank) Kupa River (Danube Basin), Croatia (45.628231, 15.508737).

CIIMAR_BIV6094. (COI: OR841764, go to GenBank) ditto.

CIIMAR_BIV6095. (COI: OR841765, go to GenBank) ditto.

CIIMAR_BIV6096. (COI: OR841766, go to GenBank) ditto.

CIIMAR_BIV6097. (COI: OR841767, go to GenBank) ditto.

Czech Republic

Mp_3653_ACZ1. (COI: KJ526773, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526796, go to GenBank) Chumava River (Elbe Basin), Czechia (49.870833, 14.004722).

Mp_3653_ACZ11. (COI: KJ526782, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526805, go to GenBank) ditto.

Mp_3653_ACZ13. (COI: KJ526783, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526806, go to GenBank) ditto.

Mp_3653_ACZ2. (COI: KJ526774, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526797, go to GenBank) ditto.

Mp_3653_ACZ3. (COI: KJ526775, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526798, go to GenBank) ditto.

Mp_3653_ACZ4. (COI: KJ526776, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526799, go to GenBank) ditto.

Mp_3653_ACZ5. (COI: KJ526777, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526800, go to GenBank) ditto.

Mp_3653_ACZ6. (COI: KJ526778, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526801, go to GenBank) ditto.

Mp_3653_ACZ8. (COI: KJ526780, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526803, go to GenBank) ditto.

Mp_3653_ACZ9. (COI: KJ526781, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526804, go to GenBank) ditto.

CIIMAR_UC07. (COI: OR841768, go to GenBank) Luznice River (Elbe Basin), Czechia (49.311013, 14.502444).

CIIMAR_UC08. (COI: OR841769, go to GenBank) ditto.

CIIMAR_UC09. (COI: OR841770, go to GenBank) ditto.

Mp_3653_BCZ26. (COI: KJ526784, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526807, go to GenBank) Sazava River (Elbe Basin), Czechia (49.856111, 14.700278).

Mp_3653_BCZ27. (COI: KJ526785, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526808, go to GenBank) ditto.

Mp_3653_BCZ28. (COI: KJ526786, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526809, go to GenBank) ditto.

Mp_3653_BCZ29. (COI: KJ526787, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526810, go to GenBank) ditto.

Mp_3653_BCZ30. (COI: KJ526788, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526811, go to GenBank) ditto.

Mp_3653_BCZ32. (COI: KJ526789, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526812, go to GenBank) ditto.

Mp_3653_BCZ33. (COI: KJ526790, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526813, go to GenBank) ditto.

Mp_3653_BCZ34. (COI: KJ526791, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526814, go to GenBank) ditto.

Mp_3653_BCZ35. (COI: KJ526792, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526815, go to GenBank) ditto.

Mp_3653_BCZ37. (COI: KJ526793, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526816, go to GenBank) ditto.

Mp_3653_BCZ38. (COI: KJ526794, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526817, go to GenBank) ditto.

Mp_3653_BCZ40. (COI: KJ526795, ND2, ND3, tRNA-His, tRNA-Ala, tRNA-Ser, tRNA-Glu: KJ526818, go to GenBank) ditto.

Finland

CIIMAR_BIV3480. (COI: OR841771, go to GenBank) Kokemäenjoki River (Kokemäenjoki Basin), Finland (61.56193, 21.66649).

CIIMAR_BIV3481. (COI: OR841772, go to GenBank) ditto.

CIIMAR_BIV3482. (COI: OR841773, go to GenBank) ditto.

CIIMAR_BIV3483. (COI: OR841774, go to GenBank) ditto.

CIIMAR_BIV3484. (COI: OR841775, go to GenBank) ditto.

France

MNHN_IM-2009-17754. (16S: KC703451, 28S: KC703696, COI: KC703897, go to GenBank) Aisne River (Seine Basin), France (49.475847, 4.589161).

MNHN_IM-2009-17755. (16S: KC703452, 28S: KC703697, COI: KC703898, go to GenBank) ditto.

MNHN_IM-2009-17763. (16S: KC703458, 28S: KC703703, COI: KC703900, go to GenBank) ditto.

MNHN_IM-2009-17792. (16S: KC703472, 28S: KC703717, COI: KC703901, go to GenBank) ditto.

MNHN_IM-2009-17796. (16S: KC703476, 28S: KC703721, COI: KC703905, go to GenBank) ditto.

MNHN_IM-2009-22398. (16S: KC703613, 28S: KC703862, COI: KC703961, go to GenBank) ditto.

MNHN_IM-2009-17761. (16S: KC703457, 28S: KC703702, go to GenBank) France: Aisne River.

MNHN_IM-2009-17766. (16S: KC703461, 28S: KC703706, go to GenBank) ditto.

MNHN_IM-2009-17790. (16S: KC703470, 28S: KC703715, go to GenBank) ditto.

MNHN_IM-2009-17805. (16S: KC703485, 28S: KC703730, go to GenBank) ditto.

MNHN_IM-2009-12721. (16S: KC703435, 28S: KC703680, go to GenBank) France: Aubetin River.

MNHN_IM-2009-12617. (16S: KC703382, 28S: KC703641, go to GenBank) France: Boivre River.

MNHN_IM-2009-21230. (16S: KC703538, 28S: KC703787, go to GenBank) France: Ourcq River.

MNHN_IM-2009-21213. (16S: KC703523, 28S: KC703771, COI: KC703926, go to GenBank) La Bruche Canal (Rhine Basin), France (48.579961, 7.678739).

MNHN_IM-2009-12720. (16S: KC703434, 28S: KC703679, COI: KC703895, go to GenBank) Lunain River (Seine Basin), France (48.2812, 2.7954).

MNHN_IM-2009-21219. (16S: KC703529, 28S: KC703777, COI: KC703928, go to GenBank) Ourcq River (Seine Basin), France (49.186301, 3.360753).

MNHN_IM-2009-21221. (16S: KC703530, 28S: KC703779, COI: KC703929, go to GenBank) ditto.

MNHN_IM-2009-21227. (16S: KC703535, 28S: KC703784, COI: KC703930, go to GenBank) ditto.

MNHN_IM-2009-21232. (16S: KC703540, 28S: KC703789, COI: KC703931, go to GenBank) ditto.

MNHN_IM-2009-12677. (16S: KC703403, COI: KC703890, go to GenBank) Petit Gland River (Seine Basin), France (49.9059, 4.1585).

MNHN_IM-2009-12680. (16S: KC703405, 28S: KC703650, COI: KC703891, go to GenBank) ditto.

MNHN_IM-2009-17769. (16S: KC703516, 28S: KC703763, COI: KC703922, go to GenBank) ditto.

MNHN_IM-2009-17842. (16S: KC703508, 28S: KC703756, COI: KC703914, go to GenBank) ditto.

MNHN_IM-2009-17848. (16S: KC703514, 28S: KC703762, COI: KC703920, go to GenBank) ditto.

MNHN_IM-2009-17811. (16S: KC703491, 28S: KC703736, COI: KC703909, go to GenBank) Therain River (Seine Basin), France (49.50631, 1.971917).

MNHN_IM-2009-12564. (16S: KC703351, 28S: KC703622, COI: JX046553, go to GenBank) Vienne River (Loire Basin), France (47.017062, 0.563053).

MNHN_IM-2009-12566. (16S: KC703352, COI: KC703870, go to GenBank) ditto.

MNHN_IM-2009-12595. (16S: KC703369, 28S: KC703634, COI: KC703872, go to GenBank) ditto.

MNHN_IM-2009-12616. (16S: KC703381, COI: KC703878, go to GenBank) ditto.

MNHN_IM-2009-12629. (16S: KC703385, 28S: KC703644, COI: KC703879, go to GenBank) ditto.

MNHN_IM-2009-12630. (16S: KC703386, 28S: KC703645, COI: KC703880, go to GenBank) ditto.

Germany

TUM_AI02. (COI: OR841787, go to GenBank) Aisch River (Rhine Basin), Germany (49.70439, 10.854872).

TUM_AI03. (COI: OR841788, go to GenBank) ditto.

CIIMAR_BIV4202. (COI: OR841789, go to GenBank) Alte Schwentine River (Schlei/Trave Basin), Germany (54.185686, 10.241204).

CIIMAR_BIV4203. (COI: OR841790, go to GenBank) ditto.

CIIMAR_BIV4204. (COI: OR841791, go to GenBank) ditto.

CIIMAR_BIV4205. (COI: OR841792, go to GenBank) ditto.

CIIMAR_BIV4206. (COI: OR841793, go to GenBank) ditto.

CIIMAR_BIV4192. (COI: OR841777, go to GenBank) Bollingstedter Au River (Eider Basin), Germany (54.588631, 9.433135).

CIIMAR_BIV4193. (COI: OR841778, go to GenBank) ditto.

CIIMAR_BIV4194. (COI: OR841779, go to GenBank) ditto.

CIIMAR_BIV4195. (COI: OR841780, go to GenBank) ditto.

CIIMAR_BIV4196. (COI: OR841781, go to GenBank) ditto.

CIIMAR_BIV4197. (COI: OR841782, go to GenBank) Ilmenau River (Elbe Basin), Germany (53.148217, 10.467698).

CIIMAR_BIV4198. (COI: OR841783, go to GenBank) ditto.

CIIMAR_BIV4199. (COI: OR841784, go to GenBank) ditto.

CIIMAR_BIV4200. (COI: OR841785, go to GenBank) ditto.

CIIMAR_BIV4201. (COI: OR841786, go to GenBank) ditto.

TUM_RG03. (COI: OR841776, go to GenBank) Regen River (Danube Basin), Germany (49.211311, 12.218066).

Hungary

TUM_UC02. (COI: OR841794, go to GenBank) Ipoly River (Danube Basin), Hungary (47.930964, 18.771094).

TUM_UC03. (COI: OR841795, go to GenBank) ditto.

TUM_UC04. (COI: OR841796, go to GenBank) ditto.

TUM_UC06. (COI: OR841797, go to GenBank) ditto.

Kazakhstan

RMBH_biv_595_1. (COI: MK603933, go to GenBank) Kazakhstan: Ural River (50.7131°N, 51.3808 °E), Caspian Sea drainage.

RMBH_biv_595_2. (COI: MK603934, go to GenBank) ditto.

RMBH_biv_595_3. (COI: MK603935, go to GenBank) ditto.

RMBH_biv_595_4. (COI: MK603936, go to GenBank) ditto.

Latvia

TUM_DG04. (COI: OR841798, go to GenBank) Düna River (Western Dvina Basin), Latvia (55.89361, 26.875624).

TUM_DG05. (COI: OR841799, go to GenBank) ditto.

TUM_DG06. (COI: OR841800, go to GenBank) ditto.

TUM_DG07. (COI: OR841801, go to GenBank) ditto.

TUM_DG08. (COI: OR841802, go to GenBank) ditto.

TUM_IN01. (COI: OR841803, go to GenBank) Indrica River (Western Dvina Basin), Latvia (55.845895, 27.413246).

TUM_IN02. (COI: OR841804, go to GenBank) ditto.

TUM_IN05. (COI: OR841805, go to GenBank) ditto.

TUM_IN06. (COI: OR841806, go to GenBank) ditto.

TUM_IN07. (COI: OR841807, go to GenBank) ditto.

Lithuania

CIIMAR_BIV4401. (COI: OR841814, go to GenBank) Jūra River (Neman Basin), Lithuania (55.540367, 21.993161).

CIIMAR_BIV4402. (COI: OR841815, go to GenBank) ditto.

CIIMAR_BIV4403. (COI: OR841816, go to GenBank) ditto.

CIIMAR_BIV4404. (COI: OR841817, go to GenBank) ditto.

CIIMAR_BIV4405. (COI: OR841818, go to GenBank) ditto.

CIIMAR_BIV4406. (COI: OR841819, go to GenBank) ditto.

CIIMAR_BIV4461. (COI: OR841808, go to GenBank) Nemunėlis River/Lake (Lielupe Basin), Lithuania (56.151583, 25.139853).

CIIMAR_BIV4462. (COI: OR841809, go to GenBank) ditto.

CIIMAR_BIV4463. (COI: OR841810, go to GenBank) ditto.

CIIMAR_BIV4464. (COI: OR841811, go to GenBank) ditto.

CIIMAR_BIV4465. (COI: OR841812, go to GenBank) ditto.

CIIMAR_BIV4466. (COI: OR841813, go to GenBank) ditto.

CIIMAR_BIV4391. (COI: OR841820, go to GenBank) Ūla River (Neman Basin), Lithuania (54.046133, 24.623964).

CIIMAR_BIV4392. (COI: OR841821, go to GenBank) ditto.

CIIMAR_BIV4393. (COI: OR841822, go to GenBank) ditto.

CIIMAR_BIV4394. (COI: OR841823, go to GenBank) ditto.

CIIMAR_BIV4395. (COI: OR841824, go to GenBank) ditto.

CIIMAR_BIV4396. (COI: OR841825, go to GenBank) ditto.

Poland

CIIMAR_BIV3560. (COI: OR841826, go to GenBank) Blizna River (Neman Basin), Poland (53.923814, 23.050983).

CIIMAR_BIV3561. (COI: OR841827, go to GenBank) ditto.

CIIMAR_BIV3562. (COI: OR841828, go to GenBank) ditto.

CIIMAR_BIV3563. (COI: OR841829, go to GenBank) ditto.

CIIMAR_BIV3564. (COI: OR841830, go to GenBank) ditto.

CIIMAR_BIV3554. (COI: OR841836, go to GenBank) Brzeźniczanka River (Oder Basin), Poland (51.778672, 15.271731).

CIIMAR_BIV3520. (COI: OR841842, go to GenBank) Bug River (Vistula Basin), Poland (52.363106, 22.871444).

CIIMAR_BIV3521. (COI: OR841843, go to GenBank) ditto.

CIIMAR_BIV3522. (COI: OR841844, go to GenBank) ditto.

CIIMAR_BIV3523. (COI: OR841845, go to GenBank) ditto.

CIIMAR_BIV3524. (COI: OR841846, go to GenBank) ditto.

Mp_4376_UcF3. (COI: KF515713, go to GenBank) Czarna Hancza River/Lake (Neman Basin), Poland (53.970883, 23.303617).

Mp_4376_UcF4. (COI: KF515714, go to GenBank) ditto.

Mp_4376_UcF5. (COI: KF515715, go to GenBank) ditto.

Mp_4376_UcF2. (COI: KF515719, go to GenBank) Drawa River/Lake (Oder Basin), Poland (53.495583, 15.809933).

CIIMAR_BIV3530. (COI: OR841837, go to GenBank) Slupia River (Slupia Basin), Poland (54.357033, 17.153969).

CIIMAR_BIV3531. (COI: OR841838, go to GenBank) ditto.

CIIMAR_BIV3532. (COI: OR841839, go to GenBank) ditto.

CIIMAR_BIV3533. (COI: OR841840, go to GenBank) ditto.

CIIMAR_BIV3534. (COI: OR841841, go to GenBank) ditto.

TUM_PSP01. (COI: OR841831, go to GenBank) Szeszupa River (Neman Basin), Poland (54.2576, 22.89388).

TUM_PSP02. (COI: OR841832, go to GenBank) ditto.

TUM_PSP03. (COI: OR841833, go to GenBank) ditto.

TUM_PSP04. (COI: OR841834, go to GenBank) ditto.

TUM_PSP05. (COI: OR841835, go to GenBank) ditto.

Mp_4376_UcF1. (COI: KF515718, go to GenBank) Szeszupa River/Lake (Neman Basin), Poland (54.271467, 22.9117).

Mp_4376_UcF6. (COI: KF515716, go to GenBank) Warkocz River/Lake (Vistula Basin), Poland (50.840236, 20.757725).

Mp_4376_UcF7. (COI: KF515717, go to GenBank) ditto.

Romania

CIIMAR_BIV8437. (COI: OR841853, go to GenBank) Caraș River (Danube Basin), Romania (45.080957, 21.548143).

CIIMAR_BIV8438. (COI: OR841854, go to GenBank) ditto.

CIIMAR_BIV8439. (COI: OR841855, go to GenBank) ditto.

CIIMAR_BIV8440. (COI: OR841856, go to GenBank) ditto.

CIIMAR_BIV8441. (COI: OR841857, go to GenBank) ditto.

CIIMAR_BIV8734. (COI: OR841858, go to GenBank) Prut River (Danube Basin), Romania (46.24386, 28.110237).

CIIMAR_BIV8736. (COI: OR841859, go to GenBank) ditto.

CIIMAR_BIV8738. (COI: OR841860, go to GenBank) ditto.

TUM_RSM01. (COI: OR841847, go to GenBank) Someș River (Danube Basin), Romania (47.631908, 23.377935).

TUM_RSM02. (COI: OR841848, go to GenBank) ditto.

TUM_RSM03. (COI: OR841849, go to GenBank) ditto.

TUM_RSM04. (COI: OR841850, go to GenBank) ditto.

TUM_RSM05. (COI: OR841851, go to GenBank) ditto.

CIIMAR_BIV4171. (COI: OR841852, go to GenBank) Talna River (Danube Basin), Romania (47.888876, 23.230259).

Russia

RMBH_biv_304_10. (COI: OR841866, go to GenBank) Iren River (Volga Basin), Russia (57.416828, 56.949358).

RMBH_biv_304_9. (COI: OR841867, go to GenBank) ditto.

RMBH_biv_280_1. (COI: OR841863, SRA: SRX22628950, go to GenBank) Khopyor River (Don Basin), Russia (51.487836, 42.593142).

RMBH_biv_280_10. (COI: OR841864, go to GenBank) ditto.

RMBH_biv_280_6. (COI: OR841865, go to GenBank) ditto.

RMBH_biv_196_5. (COI: OR841861, go to GenBank) Khopyor River (Don Basin), Russia (51.580074, 43.182045).

RMBH_biv_196_6. (COI: OR841862, go to GenBank) ditto.

RMBH_biv_304_5. (16S: MK574407, 28S: MK574548, COI: MK574271, go to GenBank) Russia: European part, Perm Region, Iren River (57.4167°N, 56.9496°E), Volga Basin, Caspian Sea drainage.

RMBH_biv_304_7. (16S: MK574408, 28S: MK574549, COI: MK574272, go to GenBank) ditto.

RMBH_biv_304_8. (16S: MK574409, 28S: MK574550, COI: MK574273, go to GenBank) ditto.

RMBH_biv_305_1. (16S: MK574410, 28S: MK574551, COI: MK574274, go to GenBank) Russia: European part, Samara Region, Samara River (53.0068°N, 50.7768°E), Volga Basin, Caspian Sea drainage.

RMBH_biv_305_2. (16S: MK574411, 28S: MK574552, COI: MK574275, go to GenBank) ditto.

RMBH_biv_305_3. (16S: MK574412, 28S: MK574553, COI: MK574276, go to GenBank) ditto.

RMBH_biv_196_1. (16S: MK574401, 28S: MK574542, COI: MK574265, go to GenBank) Russia: European part, Saratov Region, Khopyor River (51.5630°N, 43.1530°E), Don Basin, Azov Sea drainage.

RMBH_biv_196_2. (16S: MK574402, 28S: MK574543, COI: MK574266, go to GenBank) ditto.

RMBH_biv_196_3. (16S: MK574403, 28S: MK574544, COI: MK574267, go to GenBank) ditto.

RMBH_biv_280_3. (16S: MK574404, 28S: MK574545, COI: MK574268, go to GenBank) Russia: European part, Voronezh Region, Khopyor River (51.4878°N, 42.5931°E), Don Basin, Azov Sea drainage.

RMBH_biv_280_4. (16S: MK574405, 28S: MK574546, COI: MK574269, go to GenBank) ditto.

RMBH_biv_280_5. (16S: MK574406, 28S: MK574547, COI: MK574270, go to GenBank) ditto.

Mp_3904_Cr1. (16S: MH544931, ITS1: MH544886, go to GenBank) Russia: River Ivitza, Volga basin, Rameshkovsky District, Tver Oblast.

Mp_3904_Cr11. (16S: MH544928, 5.8S: MH544889, go to GenBank) ditto.

Mp_3904_Cr17. (16S: MH544927, ITS1, 5.8S: MH544890, go to GenBank) ditto.

Mp_3904_Cr24. (16S: MH544926, ITS1: MH544891, go to GenBank) ditto.

Mp_3904_Cr27. (16S: MH544917, ITS1: MH544883, go to GenBank) ditto.

Mp_3904_Cr3. (16S: MH544930, ITS1, 5.8S: MH544887, go to GenBank) ditto.

Mp_3904_Cr32. (16S: MH544918, COI: MH539794, ITS1, 5.8S: MH544884, go to GenBank) ditto.

Mp_3904_Cr33. (16S: MH544916, COI: MH539797, ITS1, 5.8S: MH544885, go to GenBank) ditto.

Mp_3904_Cr34. (16S: MH544925, ITS1: MH544892, go to GenBank) ditto.

Mp_3904_Cr41. (16S: MH544924, ITS1, 5.8S: MH544893, go to GenBank) ditto.

Mp_3904_Cr48. (16S: MH544915, COI: MH539796, go to GenBank) ditto.

Mp_3904_Cr49. (16S: MH544923, ITS1, 5.8S: MH544894, go to GenBank) ditto.

Mp_3904_Cr51. (16S: MH544922, ITS1, 5.8S: MH544895, go to GenBank) ditto.

Mp_3904_Cr52. (16S: MH544921, ITS1, 5.8S: MH544896, go to GenBank) ditto.

Mp_3904_Cr53. (16S: MH544920, ITS1, 5.8S: MH544897, go to GenBank) ditto.

Mp_3904_Cr7. (16S: MH544929, ITS1: MH544888, go to GenBank) ditto.

Mp_3904_Cr8. (16S: MH544919, COI: MH539795, ITS1, 5.8S: MH544882, go to GenBank) ditto.

RMBH_biv_305_4. (COI: OR841868, go to GenBank) Samara River (Volga Basin), Russia (53.006977, 50.776563).

RMBH_biv_305_5. (COI: OR841869, go to GenBank) ditto.

RMBH_biv_305_6. (COI: OR841870, go to GenBank) ditto.

RMBH_biv_305_7. (COI: OR841871, go to GenBank) ditto.

RMBH_biv_305_8. (COI: OR841872, go to GenBank) ditto.

RMBH_biv_305_9. (COI: OR841873, go to GenBank) ditto.

Sweden

TUM_F5. (COI: OR841879, go to GenBank) Bråån River (Kävlingeåns Basin), Sweden (55.800696, 13.64148).

TUM_F6. (COI: OR841880, go to GenBank) ditto.

TUM_F7. (COI: OR841881, go to GenBank) ditto.

TUM_F8. (COI: OR841882, go to GenBank) ditto.

TUM_F9. (COI: OR841883, go to GenBank) ditto.

CIIMAR_BIV3410. (COI: OR841875, go to GenBank) Gardvedaan River (Eman Basin), Sweden (57.352863, 15.820028).

CIIMAR_BIV3411. (COI: OR841876, go to GenBank) ditto.

CIIMAR_BIV3412. (COI: OR841877, go to GenBank) ditto.

CIIMAR_BIV3413. (COI: OR841878, go to GenBank) ditto.

CIIMAR_BIV3425. (COI: OR841884, go to GenBank) Storan River (Storan Basin), Sweden (58.021804, 15.652948).

CIIMAR_BIV3426. (COI: OR841885, go to GenBank) ditto.

CIIMAR_BIV3427. (COI: OR841886, SRA: SRX22628952, go to GenBank) ditto.

CIIMAR_BIV3428. (COI: OR841887, go to GenBank) ditto.

CIIMAR_BIV3429. (COI: OR841888, go to GenBank) ditto.

CIIMAR_BIV3449. (COI: OR841889, go to GenBank) Svartaan River (Svartaan Basin), Sweden (58.86938, 17.121288).

CIIMAR_BIV3450. (COI: OR841890, go to GenBank) ditto.

CIIMAR_BIV3451. (COI: OR841891, go to GenBank) ditto.

CIIMAR_BIV3452. (COI: OR841892, go to GenBank) ditto.

CIIMAR_BIV3453. (COI: OR841893, go to GenBank) ditto.

NRM_42648-M099. (16S: DQ060162, COI: DQ060174, go to GenBank) Sweden.

CIIMAR_BIV3409. (16S: MZ519056, 18S: PP693707, 28S: MZ519293, 28S: PP693722, COI: MZ512216, COI: OR841874, mtGenome: OR912547, go to GenBank) Sweden, Gardvedaan River (Eman Basin), Sweden (57.352863, 15.820028).

TUM_T10. (COI: OR841894, go to GenBank) Tommarpsån River (Tommarpsån Basin), Sweden (55.554078, 14.149254).

TUM_T11. (COI: OR841895, go to GenBank) ditto.

TUM_T13. (COI: OR841896, go to GenBank) ditto.

TUM_T14. (COI: OR841897, go to GenBank) ditto.

TUM_T15. (COI: OR841898, go to GenBank) ditto.

Ukraine

ZSU-U13. (COI: KY930360, go to GenBank) Desna River (Dnipro Basin), Ukraine (51.907434, 33.215528).

ZSU-U11. (COI: KY930359, go to GenBank) Stalineshty River (Danube Basin), Ukraine (48.25755, 26.543876).

CIIMAR_BIV3510. (COI: OR841899, go to GenBank) Stryi River (Dniester Basin), Ukraine (49.215956, 23.298919).

CIIMAR_BIV3511. (COI: OR841900, go to GenBank) ditto.

CIIMAR_BIV3512. (COI: OR841901, go to GenBank) ditto.

CIIMAR_BIV3513. (COI: OR841902, go to GenBank) ditto.

CIIMAR_BIV3514. (COI: OR841903, go to GenBank) ditto.

CIIMAR_BIV3500. (COI: OR841904, go to GenBank) Styr River (Dnipro Basin), Ukraine (50.171344, 24.896869).

CIIMAR_BIV3501. (COI: OR841905, go to GenBank) ditto.

CIIMAR_BIV3502. (COI: OR841906, go to GenBank) ditto.

CIIMAR_BIV3503. (COI: OR841907, SRA: SRX22628953, go to GenBank) ditto.

CIIMAR_BIV3504. (COI: OR841908, go to GenBank) ditto.

GenBank Vouchers Not Yet Georeferenced

Biv_Suc1D. (28S: OQ351306, go to GenBank)

Biv_SucD. (28S: OQ351307, go to GenBank)

C3Gf. (COI: OL872298, go to GenBank)

MNCN_N1177. (16S: KX400176, go to GenBank)

MNCN_N426. (16S: KX400194, go to GenBank)

MNCN_N432. (16S: KX400200, go to GenBank)

MNCN_N438. (16S: KX400206, go to GenBank)

MNCN_N440. (16S: KX400208, go to GenBank)

MNCN_N59. (16S: KX400210, go to GenBank)

MNCN_N65. (16S: KX400212, go to GenBank)

MNCN_N66. (16S: KX400213, go to GenBank)

MNCN_N74. (16S: KX400214, go to GenBank)

MNCN_N75. (16S: KX400215, go to GenBank)

Mp_4335_C10. (COI: KJ525921, go to GenBank)

Mp_4335_C11. (COI: KJ525922, go to GenBank)

Mp_4335_C2. (COI: KJ525913, go to GenBank)

Mp_4335_C4. (COI: KJ525915, go to GenBank)

Mp_4335_C6. (COI: KJ525917, go to GenBank)

Mp_4335_C7. (COI: KJ525918, go to GenBank)

Mp_4335_C8. (COI: KJ525919, go to GenBank)

Mp_4335_C9. (COI: KJ525920, go to GenBank)

Mp_4335_I11. (5.8S, ITS1: KJ525946, go to GenBank)

Mp_4335_I14. (5.8S, ITS1: KJ525949, go to GenBank)

Mp_4335_I15. (5.8S, ITS1: KJ525950, go to GenBank)

Mp_4335_I20. (5.8S, ITS1: KJ525955, go to GenBank)

Mp_4335_I23. (5.8S, ITS1: KJ525958, go to GenBank)

Mp_4335_I24. (5.8S, ITS1: KJ525959, go to GenBank)

Mp_4335_I25. (5.8S, ITS1: KJ525960, go to GenBank)

Mp_4335_I26. (5.8S, ITS1: KJ525961, go to GenBank)

Mp_4335_I27. (5.8S, ITS1: KJ525962, go to GenBank)

Mp_4335_I28. (5.8S, ITS1: KJ525963, go to GenBank)

Mp_4335_I29. (5.8S, ITS1: KJ525964, go to GenBank)

Mp_4335_I7. (5.8S, ITS1: KJ525942, go to GenBank)

Mp_4335_I8. (5.8S, ITS1: KJ525943, go to GenBank)

Mp_4335_N4. (ND1: KJ525931, go to GenBank)

Mp_4335_N5. (ND1: KJ525932, go to GenBank)

Mp_4335_N6. (ND1: KJ525933, go to GenBank)

Mp_4335_N7. (ND1: KJ525934, go to GenBank)

Mp_4335_N8. (ND1: KJ525935, go to GenBank)

NRM_41778-M024. (18S, ITS1: DQ060188, go to GenBank)

NRM_42769-M074. (18S, ITS1: DQ060187, go to GenBank)

P3Nf. (COI: OL872299, go to GenBank)