MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Lanceolariini | Lanceolaria
species Lanceolaria grayii (Griffith & Pidgeon, 1833): specimens
11 available nominal species synonyms (16 total synonyms)
Recent Taxonomic Opinions
Lanceolaria grayana — Graf (2007), Graf & Cummings (2007), Kondo (2008), Bogan (2010), Ouyang et al. (2011), ... Kim et al. (2021), Zieritz et al. (2021), Wu et al. (2021), Dai et al. (2024), Xiang et al. (2025)
Lanceolaria cylindrica — Graf (2007), Graf & Cummings (2007), Bogan (2010), IUCN (2011), He & Zhuang (2013), Soliman et al. (2016), Bogatov & Prozorova (2017), Zieritz et al. (2018), Graf & Cummings (2021), Liu et al. (2022)
Lanceolaria eucylindrica — Graf & Cummings (2007), Bogan (2010), Xiong et al. (2012), He & Zhuang (2013), Bogatov & Prozorova (2017), Zieritz et al. (2018), Wu et al. (2018), Huang et al. (2019), Liu et al. (2020), Graf & Cummings (2021), Liu et al. (2022), Campbell & Lydeard (2026)
Lanceolaria bogatovi — Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria chankensis — Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017), Bogatov (2022)
Lanceolaria maacki — Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria ussuriensis — Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria grayii — IUCN (2012), He & Zhuang (2013), Bogatov & Prozorova (2017), Zieritz et al. (2018), Wu et al. (2018), ... Dai et al. (2025), Chen et al. (2025), Campbell & Lydeard (2026), Wu et al. (2026), Wu et al. (2026)
Lanceolaria yueyingae — He & Zhuang (2013), WoRMS (2017), Zieritz et al. (2018), Liu et al. (2020), Liu et al. (2022)
Lanceolaria glayana [sic] — Wang et al. (2016)
“Lanceolaria grayii” ** — Lopes-Lima et al. (2017), Bolotov et al. (2017), Bolotov et al. (2017), Huang et al. (2019), Jeratthitikul et al. (2022), Campbell & Lydeard (2026)
“Lanceolaria grayii” ** — Sano et al. (2017), Sano et al. (2020), Hata et al. (2022), Campbell & Lydeard (2026)
Lanceolaria grayi — Do et al. (2018)
Lanceolaria maacki maacki — Bogatov (2022)
Lanceolaria maacki bogatovi — Bogatov (2022)
Lanceolaria maacki ussuriensis — Bogatov (2022)
Range
Amur Basin of Russia to southern China (East Asia: AS2 AS4 AS5)
Map of Georeferenced Specimens
MUSSELp Specimen Records
Type Specimens
SMF_3570
Zw. Hanoi u Hoa Bimo, Tonkin [Road From Hanoi To Hoa-Bimo, China, North Pacific Ocean]
paratype
Unio gracillimus Rolle, 1904
USNM_84162
China
figured type
Unio grayanus Lea, 1834
SMF_3571
zwischen Hanoi [Hà Nội] und Hoa Bimo [= Hòa Bình?], Tonkin [Vietnam]
paratypes
Unio gracillimus Rolle, 1904
Other Specimens by Country | show by collection
42 Specimens from [blank]
MNHN_MP_3817
[locality illegible - Asia]
MNHN_MP_3887
[locality illegible - Asia]
MNHN_MP_3888
[locality illegible - Asia]
MNHN_MP_3905
[locality illegible - Asia]
SMF_14347
A.d. Kaiserkanal [Jīng Háng Dà Yùnhé] und seinen Seen zwischen Tsiningtschau [Jǐníng] und Hantschwang [Hanzhuangzhen (Wēishān Xiàn, Jǐníng Shì)], Schantung [Shāndōng] [China Shāndōng]
NCSM_27935
Asia, Chang (Yangtze) Basin, China, Kiangsu (Jiangsu), Wu Xian Co. Dong Shan, Guang Ming, Tai Hu Lake
NCSM_84955
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam, Mien Nui va trung du (Northeastern), Phú Thọ Co. Lao Cai market, vendor 2, [point estimated Quang Trung]. 21.93333° N, 104.6306° E
NCSM_84921
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam, Mien Nui va trung du (Northeastern), Phú Thọ Co. Viet Tri market, [point estimated 2.91 air miles WNW center Phú Thọ]. 21.32429° N, 105.4027° E
NCSM_84945
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam, Mien nui va trung du (Northeastern), Yen Ba Co. Lao Cai market, vendor 1, [point estimated center Dai Buc]. 21.86583° N, 104.6325° E
NCSM_84992
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam, Mien nui va trung du (Northeastern), Yen Bai Co. [Unnamed body of water], Luc Yen District, bought in maret in Ha Giang, [point estimated on Sông Chảy 3.81 air miles WSW center Yên Thế], 22.09051° N, 104.7114° E
NCSM_84926
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam, Mien nui va trung du (Northeastern), Yen Bai Co. Yen Bai market, Ha Hoa district, [point estimated 3.57 air miles NE center Hop Minh]. 21.71696° N, 104.9075° E
NCSM_85262
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam,Đồng Bằng Sông Hồng (northern). Hanoi Co. Cầu Giấy district , [4.17 air miles S center Phú Thường]. 21.02431° N, 105.7977° E
NCSM_85265
Asia, Hong Basin (Red Basin)/Hong R. (Red R.) Vietnam,Đồng Bằng Sông Hồng (northern). Hanoi Co. Cầu Giấy district of Hanoi, [3.86 air miles S center Phú Thường]. 21.02669° N, 105.7984° E
NCSM_33481
Asia, Jiangsu Minor Coastal (misc. creek tribs) China, Kiangsu (Jiangsu), Wu Xian Co. Tai Hu [Tai-Hu] Lake, Dong Shan, Guang Ming, [point estimated 12.0 air miles SSW center Mudu]. 31.09417° N, 120.4178° E
NCSM_27185
Asia, Philipine Sea, Honshû, Okayama Prefecture Co. Small drain of 'Gion-yôsui', in front of JA Takashima, Takashima, Okayama City, Okayama Prefecture, western Honshû. 34.68917° N, 133.9644° E
NCSM_47209
Asia, Vietnam Minor Coastal (misc. creek tribs) Vietnam, Bắc Trung Bộ (North Central), Hà Tinh Co. [Point estimated center] Hồng Lĩnh, market. 18.53137° N, 105.7234° E
SMF_24004
Bächlein nördlich Shih-Tze-Shan bei Tan-Chia-Chiao, Distrikt Ta-Yeh [Dàyě], Hupeh [Húběi] [China Húběi Shěng]
SMF_14351
China
SMF_14354
China
SMF_14358
China
SMF_14357
Hoihow [= Hǎikǒu], Hainan [China Hǎinán]
SMF_14355
Hunan [Húnán Shěng], Mittel-China [China Húnán Shěng]
ANSP_125761
Kachet, Hainan, Fukien Prov., China
SMF_14352
Kanäle des Tai-hu-Sees [Tài Hú], 80 km westlich von Shanghai [Shànghǎi] [China Jiāngsū]
SMF_14359
Kiangsi [= Jiāngxī], Chi-an-hsien [= Ji'an?], in kleinen Flüssen, Jangtse-Gebiet [China Jiāngxī]
SMF_14360
Ningpo [Níngbō shì] [China Zhèjiāng]
SMF_14353
Nordfluss [= Běi Jiāng], Kanton, China [China Guǎngdōng (Kanton)]
MCZ_109289
ponds just outside Peiping, China
SMF_168055
Route de Hanoi [Hà Nội] à Hoa [= Hòa Bình?], Tonkin [Vietnam]
BMNH_57-8-3-35
Shang Kae
SMF_14362
Soochow (= Sutschau) [Sūzhōu Shì], Provinz Nganhwei [Ānhuī], China [China Ānhuī]
USNM_349149
Suifu, Szechuan, China
SMF_24003
Teich bei Hwei-Lung-Pu [Huilongpuzhen?] und Altwasser [des Weishui/Weǐshuǐ Hé?] bei Yü-Tan-Chiao, Distrikt Ning-Hsiang [Níngxiāng Xiàn], Provinz Hunan [Húnán Shěng] [China Húnán Shěng]
SMF_14361
Tscho bei Jentschoufu [Yǎnzhōu Shì] [China Shāndōng]
BMNH_1902-3-22-1-2
Yen Bay, Roton Fleuss
USNM_360430
Yenting, Chekiang Prov., China
50 Specimens from China
BMNH_73-6-9-6
China
FMNH_142714
China
FMNH_8371
China
ANSP_125469
China
MNHN_MP_3819
China
ANSP_41666
China
ANSP_41735
China
UMMZ_108082
China
UMMZ_184031
China
USNM_17395
China
MCZ_6612
China
USNM_31451
China
USNM_472396
China
ANSP_48434
China
BMNH_MP_0678
China
BMNH_MP_0679
China
BMNH_MP_0680
China
BMNH_MP_0685
China
BMNH_MP_0688
China
UMMZ_23060
China
USNM_73069
China
ANSP_195985
China
BMNH_37-6-10-309-37
Chine
MNHN_MP_3813
Chine
MNHN_MP_3814
Chine
MNHN_MP_3826
Chine
MNHN_MP_3827
Chine
MNHN_MP_3828
Chine
MNHN_MP_3831
Chine
MNHN_MP_3833
Chine
MNHN_MP_PARIS3
Chine
MNHN_MP_3836
Chine
MNHN_MP_3830
Chine Centrale
MNHN_MP_PARIS1
Fleuve Yangt-Se, Chine
BMNH_MP_0682
Foo Chow
SMF_14350
Hunan, Mittelchina
USNM_333839
Kien-nin-fu, China
UMMZ_41016
lake in summer palace grounds, Peking, China
MNHN_MP_3812
Nord de la Chine
ANSP_184640
Peping, China
MCZ_MP_349
Prov. Nanking, China
BMNH_20020551
Shores of Poyang Lake, China
USNM_334268
Soochow (canals) China
ANSP_165016
Soochow, China
INHS_20152
Tai Hu Lake, Dong Shan, Guang Ming, Wu Xian, Jiangsu, China
UMMZ_108087
Wau, 100 mi upriver from Nanking, China
UMMZ_41018
Wau, China
ANSP_211319
Yangtse River, Kiangyinku [Kiangsu or Jiangsu Province], China
UMMZ_108088
Yangtze Kiang River, China
UMMZ_41017
Yantze River, Nanking, China
1 Specimens from Thailand
MNHN_MP_3961
Indo-Chine
5 Specimens from unknown
MNHN_MP_3890
[no locality available - Asia]
MNHN_MP_3901
[no locality available - Asia]
BMNH_MP_0683
[no locality available - Asia]
BMNH_MP_0686
[no locality available - Asia]
BMNH_MP_0729
[no locality available - Asia]
28 Specimens from Vietnam
MNHN_MP_3963
Bas Luc Nam, Village de Van Ien, Tonkin
MNHN_MP_3902
Fleuve Rouge, pres Hanoi
MNHN_MP_3965
Hanoi
MNHN_MP_3904
Hanoi, Tonkin
MCZ_MP_351
Hanoi, Tonkin
USNM_110044
Hanoi, Tonquin
MNHN_MP_3959
Lang Son, Tonkin
FMNH_115905
Luc Nam Tonkin, French Indo-China
MNHN_MP_3958
Luc Nam, Song Luc Nam, Tonkin
MNHN_MP_3001
Minh Le, Vietnam
UMMZ_108083
River Muong Kong, Hanoi, Tonkin, French Indo China
USNM_339729
Road From Hanoi To Hoa-Bimo, Tonkin
UMMZ_108084
Route de Hanoi, Hoa Bimo, Tonkin, French Indo China
SMF_14356
Sept-Pagodes, Tonkin
MCZ_79602
Song Luc Nam, Tonkin
ANSP_145262
Song Luc Nam, Tonkin
UMMZ_60377
Song Luc Nam, Tonkin, French Indo China
MCZ_102060
Song Luc Nam, Tonkin, French Indo-China
MCZ_MP_352
Song-Ky-Kong, Cao Bang, Tonkin
MNHN_MP_3899
That Ke, Tonkin
UMMZ_108086
That Khe, Tonkin, French Indo China
FMNH_70923
Tonkin
FMNH_21743
Tonkin
MNHN_MP_3889
Tonkin
MNHN_MP_3896
Tonkin
MNHN_MP_3898
Tonkin
UMMZ_108085
Tonkin, French Indo China
FMNH_21744
Vietnam Locale: Song Luc Nam, Tonkin
GenBank Vouchers
Georeferenced GenBank Vouchers
China
16_NCU_XPWU_SU099. (28S: MG595536, COI: MG463009, go to GenBank) China: Jiangxi, Gan River.
16_NCU_XPWU_SU101. (28S: MG595538, COI: MG463008, go to GenBank) ditto.
16_NCU_XPWU_SU093. (28S: MG595530, COI: MG463002, go to GenBank) China: Jiangxi, Qinglan Lake.
16_NCU_XPWU_SU094. (28S: MG595531, COI: MG463005, go to GenBank) ditto.
16_NCU_XPWU_SU095. (28S: MG595532, COI: MG463004, go to GenBank) ditto.
16_NCU_XPWU_SU096. (28S: MG595533, COI: MG463003, go to GenBank) ditto.
16_NCU_XPWU_SU100. (28S: MG595537, COI: MG463010, go to GenBank) ditto.
NCUBM_20180100. (ND1: MG933777, go to GenBank) Gan River (Jiangxi Province, N: 28.57° - 28.86°; E: 115.57° - 116.00°).
NCUBM_20180101. (ND1: MG933778, go to GenBank) ditto.
NCUBM_20180102. (ND1: MG933779, go to GenBank) ditto.
NCUBM_20180235. (COI: MG933721, go to GenBank) Poyang Lake (Jiangxi Province, N: 29.03° - 29.17°; E: 116.20° - 116.26°), China.
Mp_2657_Lgr. (16S: AF389408, go to GenBank) Qing Lan Lake, Jiangxi, China.
Mp_2735_Lgra1. (16S: AY297463, go to GenBank) ditto.
Mp_2735_Lgra2. (16S: AY297464, go to GenBank) ditto.
Russia
RMBH_biv_502_1. (16S: MK574384, 28S: MK574525, COI: MK574246, go to GenBank) Russia: Far East, Primorye Region, Komissarovka River (44.8255°N, 132.0456°E), Khanka Lake basin, Okhotsk Sea drainage.
RMBH_biv_502_2. (16S: MK574385, 28S: MK574526, COI: MK574247, go to GenBank) ditto.
RMBH_biv_502_3. (16S: MK574386, 28S: MK574527, COI: MK574248, go to GenBank) ditto.
GenBank Vouchers Not Yet Georeferenced
CIIMAR_BIV4717. (COI: MT020643, go to GenBank)
Mp_4493_154LG. (COI: KJ434525, ND1: KJ434577, go to GenBank)
Mp_PARK_C2. (16S: GQ451847, COI: GQ451861, Cytb: GQ451874, go to GenBank)
Mp_PARK_C22_1. (16S: GQ451848, Cytb: GQ451875, go to GenBank)
