species Lanceolaria grayii (Griffith & Pidgeon, 1833): synonymy

Unio grayii Griffith & Pidgeon, 1833

‘Lea’ Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): 601, pl. 21, fig. 3.

Nodularia (sec. Lanceolaria) grayana — Simpson (1900), Simpson (1914)
Unio (Nodularia) grayanus — Dautzenberg & Fischer (1905)
Lanceolaria grayana — Haas (1911), Liu et al. (1979), Wu et al. (2019)
Lanceolaria grayi — Dang et al. (1980), Do et al. (2018)
Lanceolaria grayii — IUCN (2012), He & Zhuang (2013), Bogatov & Prozorova (2017), Zieritz et al. (2018), Wu et al. (2018), ... Nagel (2025), Campbell & Lydeard (2026), Wu et al. (2026), Wu et al. (2026), Li et al. (2026)
“Lanceolaria grayii” ** — Lopes-Lima et al. (2017), Bolotov et al. (2017), Bolotov et al. (2017), Huang et al. (2019), Jeratthitikul et al. (2022), Campbell & Lydeard (2026)
“Lanceolaria grayii” ** — Sano et al. (2017), Sano et al. (2020), Hata et al. (2022), Campbell & Lydeard (2026)

Unio grayanus Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 66 [178], pl. 9, fig. 26.
HOLOTYPE USNM_84162, China.

Nodularia (sec. Lanceolaria) grayana — Simpson (1900), Simpson (1914)
Unio (Nodularia) grayanus — Dautzenberg & Fischer (1905)
Lanceolaria grayana — Haas (1911), Haas (1924), Zhadin (1938), Zhadin (1965), Haas (1969), ... Kim et al. (2021), Zieritz et al. (2021), Wu et al. (2021), Dai et al. (2024), Xiang et al. (2025)
Lanceolaria (s.s.) grayana — Thiele (1934)
Lanceolaria grayana grayana — Haas (1969)
Lanceolaria grayii — He & Zhuang (2013), Graf & Cummings (2021), Konopleva et al. (2023), Dai et al. (2025), Wu et al. (2026)
Lanceolaria glayana [sic] — Wang et al. (2016)
Lanceolaria grayi — Do et al. (2018)

Nodularia cylindrica Simpson, 1900

(correct name for Nodularia cylindracea Simpson, 1900)
Schrenck, 1867. Reise Forsch. Amurlande 2: 694, pl. 27, figs. 1-3 [as ‘Unio grayanus Lea’].
Simpson, 1900. Proc. Acad. Nat. Sci. 52: 84 [as ‘cylindracea’].
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 807.
Simpson, 1914. Descr. Cat. Naiades: 958.

Nodularia (sec. Cylindrica) cylindrica — Simpson (1900), Simpson (1914)
Nodularia cylindracea [sic] — Simpson (1900)
Lanceolaria cylindrica — Haas (1911), Zhadin (1938), Zhadin (1965), Haas (1969), Bába (2000), ... Soliman et al. (2016), Bogatov & Prozorova (2017), Zieritz et al. (2018), Graf & Cummings (2021), Liu et al. (2022)
Lanceolaria (sec. Pericylindrica) cylindrica — Thiele (1934)
Lanceolaria maacki — Moskvicheva (1973), Kantor & Sysoev (2005), Vinarski & Kantor (2016)
Lanceolaria (Pericylindrica) maacki — Zatravkin & Bogatov (1987)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)

Unio (Nodularia) gracillimus Rolle, 1904

Rolle, 1904. Nachr. Bl. Dtsch. Malak. Ges. 36: 26, pl. 4, figs. b-c.
SYNTYPE SMF_3570, Zw. Hanoi u Hoa Bimo, Tonkin.

Unio (Nodularia) grayanus — Dautzenberg & Fischer (1905)
Lanceolaria grayana — Haas (1911)
Nodularia (sec. Lanceolaria) grayana — Simpson (1914)
Lanceolaria grayana grayana — Haas (1969)
Lanceolaria grayii — He & Zhuang (2013), Nagel (2025)
Lanceolaria grayi — Do et al. (2018)

Unio (Nodularia) grayanus ‘Lea’ of Authors

(reference to Unio grayanus Lea, 1834)
‘Lea’ of Authors.
Dautzenberg & Fischer, 1905. J. de Conch. 53: 202.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 44, pl. 4b-c.

Lanceolaria grayi — Dang et al. (1980)
Lanceolaria grayana — Kondo (2008)

Lanceolaria eucylindrica Lin, 1962

Lin, 1962. Acta Zool. Sinica 14(2): 249, fig. 14.

Lanceolaria eucylindrica — Wu et al. (1999), Wu et al. (2000), Prozorova et al. (2005), Hu (2005), Graf & Cummings (2007), ... Huang et al. (2019), Liu et al. (2020), Graf & Cummings (2021), Liu et al. (2022), Campbell & Lydeard (2026)
Lanceolaria grayii — Wu et al. (2026)

Lanceolaria maacki Moskvicheva, 1973

‘Lea’ Schrenck, 1867. Reise Forsch. Amurlande 2: 694, pl. 27, figs. 1-3 [as as ‘Unio grayanus Lea’].
Moskvicheva, 1973. Zool. Zhur. 52(10): 1465, figs. 2 е-ж.

Lanceolaria maacki — Moskvicheva (1973), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria (Pericylindrica) maacki — Zatravkin (1983), Zatravkin & Bogatov (1987)
Lanceolaria cylindrica — Graf (2007), IUCN (2011), He & Zhuang (2013)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)
Lanceolaria maacki maacki — Bogatov (2022)

Lanceolaria ussuriensis Moskvicheva, 1973

(new name for Unio grayanus var. decurvata Schrenck, 1867)
Moskvicheva, 1973. Zool. Zhur. 52(10): 1465, figs. 2 з-и.

Lanceolaria ussuriensis — Moskvicheva (1973), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria (Pericylindrica) ussuriensis — Zatravkin (1983), Zatravkin & Bogatov (1987)
Lanceolaria cylindrica — Graf (2007), IUCN (2011), He & Zhuang (2013)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)
Lanceolaria maacki ussuriensis — Bogatov (2022)

Lanceolaria chankensis Moskvicheva, 1973

Moskvicheva, 1973. Zool. Zhur. 52(10): 1466, figs. к-л.

Lanceolaria chankensis — Moskvicheva (1973), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017), Bogatov (2022)
Lanceolaria (Pericylindrica) chankensis — Zatravkin (1983), Zatravkin & Bogatov (1987)
Lanceolaria cylindrica — Graf (2007), IUCN (2011), He & Zhuang (2013)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)

Lanceolaria (Pericylindrica) bogatovi Zatravkin & Starobogatov, 1984

Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1785, figs. 1-4.

Lanceolaria (Pericylindrica) sp. n. — Zatravkin (1983)
Lanceolaria (Pericylindrica) bogatovi — Zatravkin & Bogatov (1987)
Lanceolaria bogatovi — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Lanceolaria cylindrica — Graf (2007), IUCN (2011), He & Zhuang (2013)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)
Lanceolaria maacki bogatovi — Bogatov (2022)

Lanceolaria yueyingae He & Zhuang, 2013

He & Zhuang, 2013. Freshwater Bivalves China: 76, fig. 173.

Lanceolaria yueyingae — He & Zhuang (2013), WoRMS (2017), Zieritz et al. (2018), Liu et al. (2020), Liu et al. (2022)
Lanceolaria eucylindrica — Bogatov & Prozorova (2017)
Lanceolaria grayii — Wu et al. (2026)

Unio grayensis Jay, 1835

(error for Unio grayanus Lea, 1834)
Jay, 1835. Cat. Rec. Shells: 17.

Unio grayanus Schrenck, 1867

(reference to Unio grayanus Lea, 1834)
‘Lea’ Schrenck, 1867. Reise Forsch. Amurlande 2: 694, pl. 27, figs. 1-3.

Nodularia (sec. Cylindrica) cylindrica — Simpson (1900), Simpson (1914)
Nodularia cylindracea [sic] — Simpson (1900)
Lanceolaria cylindrica — Haas (1911), Haas (1969), IUCN (2011), He & Zhuang (2013)
Lanceolaria maacki — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010)
Lanceolaria (Pericylindrica) maacki — Zatravkin & Bogatov (1987)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020), Wu et al. (2026)

Unio grayanus var. decurvata Schrenck, 1867

(renamed Lanceolaria ussuriensis Moskvicheva, 1973; homonym of Unio decurvatus Rossmässler, 1835)
Schrenck, 1867. Reise Forsch. Amurlande 2: 694, pl. 27, figs. 1-3.

Lanceolaria ussuriensis — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010)
Lanceolaria (Pericylindrica) ussuriensis — Zatravkin & Bogatov (1987)
Lanceolaria cylindrica — He & Zhuang (2013)
Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Nodularia cylindracea Simpson, 1900

(incorrect original spelling of Nodularia cylindrica Simpson, 1900)
Simpson, 1900. Proc. Acad. Nat. Sci. 52: 84.

Lanceolaria grayii — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Lanceolaria glayana Wang et al., 2016

(error for Unio grayanus Lea, 1834)
Wang et al., 2016. mtDNA 27(2): 846.