Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Publication Zhadin (1938): taxonomic opinions
     publication summary

Complete Citation

Zhadin, V.I. 1938. Sem. Unionidae. Faune URSS, Mollusques 4(1). 169 pp.

Taxonomic Opinions

Anodonta (Haasiella) arcaeformis (Heude, 1877) = Anemina arcaeformis

Anodonta (Haasiella) arcaeformis var. suifunica Lindholm, 1925 = Buldowskia suifunica

Anodonta (Haasiella) euscaphys (Heude, 1879) = Anemina arcaeformis

Anodonta (Pseudanodonta) complanata ‘Ziegl.’ Rossmässler, 1835 = Pseudanodonta complanata

Anodonta (Pseudanodonta) complanata var. borealis Kobelt, 1891 = Pseudanodonta complanata

Anodonta (Pseudanodonta) complanata var. middendorffi Siemaschko, 1848 = Pseudanodonta complanata

Anodonta (Pseudanodonta) complanata var. nordenskiöldi Boettger, 1880 [sic] = Pseudanodonta complanata

Anodonta (Pseudanodonta) rossmaessleri Bourguignat, 1881 = Pseudanodonta complanata

Anodonta (Pteranodon) woodiana (Lea, 1834) = Sinanodonta woodiana

Anodonta (Pteranodon) woodiana var. elliptica (Heude, 1878) = Sinanodonta elliptica

Anodonta (s.s.) anatina (Linnaeus, 1758) = Anodonta anatina

Anodonta (s.s.) anatina var. convexa Drouet, 1888 = Anodonta anatina

Anodonta (s.s.) anatina var. lenae Zhadin, 1938 = Anodonta anatina

Anodonta (s.s.) anatina var. petshorica Zhadin, 1938 = Anodonta anatina

Anodonta (s.s.) anatina var. viridiflava Drouet, 1893 = Anodonta anatina

Anodonta (s.s.) beringiana Middendorff, 1851 = Beringiana beringiana

Anodonta (s.s.) beringiana var. suifunensis Zhadin, 1938 = Buldowskia suifunica

Anodonta (s.s.) beringiana var. taranetzi Zhadin, 1938 = Beringiana beringiana

Anodonta (s.s.) cellensis [sic] (Gmelin, 1791) = Anodonta cygnea

Anodonta (s.s.) cellensis [sic] var. cariosa Küster, 1876 [sic] = Anodonta cygnea

Anodonta (s.s.) cellensis [sic] var. maganica Servain, 1882 = Anodonta cygnea

Anodonta (s.s.) cellensis [sic] var. selengensis Dybowski, 1913 = Anodonta anatina

Anodonta (s.s.) cellensis [sic] var. sorensis [sic] Dybowski, W., 1913 = Anodonta anatina

Anodonta (s.s.) cellensis [sic] var. tehernaica [sic] Bourguignat, 1881 = Anodonta anatina
+ Anodonta tchernaica Bourguignat, 1881

Anodonta (s.s.) cygnea (Linnaeus, 1758) = Anodonta cygnea

Anodonta (s.s.) cyrea Drouet, 1881 = Anodonta anatina

Anodonta (s.s.) cyrea var. armenica Ovtschinnikov, 1935 = Anodonta anatina

Anodonta (s.s.) cyrea var. bactriana Rolle, 1897 = Anodonta anatina

Anodonta (s.s.) cyrea var. georgiana Drouet, 1881 = Anodonta anatina

Anodonta (s.s.) cyrea var. lenkoranensis Drouet, 1881 = Anodonta anatina

Anodonta (s.s.) cyrea var. samarkadensis [sic] Kobelt, 1913 = Anodonta anatina

Anodonta (s.s.) cyrea var. sogdiana Kobelt, 1896 = Anodonta anatina

Anodonta (s.s.) piscinalis Nilsson, 1823 = Anodonta anatina

Anodonta (s.s.) piscinalis var. crimeana Bourguignat, 1881 = Anodonta anatina

Anodonta (s.s.) piscinalis var. letourneuxi Bourguignat, 1870 = Anodonta anatina

Anodonta (s.s.) piscinalis var. ostiaria Drouet, 1881 = Anodonta anatina

Anodonta (s.s.) piscinalis var. ponderosa Pfeiffer, C., 1825 = Anodonta anatina

Anodonta (s.s.) piscinalis var. seisanensis Kobelt, 1912 = Anodonta anatina

Anodonta (s.s.) piscinalis var. sorica Dybowski, 1913 = Anodonta anatina

Anodonta (s.s.) piscinalis var. volgensis Zhadin, 1938 = Anodonta anatina

Anodonta (s.s.) sedakovi [sic] Siemaschko, 1848 = Anodonta anatina

Anodonta (s.s.) sedakowi var. nova Dybowski, 1913 = Anodonta anatina

Cristaria plicata (Leach, 1815)

Cristaria plicata var. herculea Middendorff, 1848 = Cristaria plicata

Lanceolaria cylindrica Simpson, 1900

Lanceolaria grayana (Lea, 1834) = Lanceolaria grayii

Margaritana dahurica (Middendorff, 1850) = Margaritifera dahurica

Margaritana margaritifera (Linnaeus, 1758) = Margaritifera margaritifera

Margaritana middendorffi Rosén, 1926 = Margaritifera middendorffi
+ Unio (Alasm.) complanatus Middendorff, 1851

Margaritana mongolica (Middendorff, 1851) = Middendorffinaia mongolica

Margaritana sachalinensis Zhadin, 1938 = Margaritifera laevis

Psilunio rothi (Bourguignat, 1863) = Potomida semirugata

Psilunio rothi var. armeniacus (Kobelt, 1912) = Unio mardinensis

Psilunio rothi var. komarovi [sic] (Boettger, O., 1880) = Unio mardinensis

Unio (Nodularia) continentalis (Haas, 1910) = Inversidens pantoensis

Unio (Nodularia) douglasiae Gray in Griffith & Pidgeon, 1834 = Nodularia douglasiae

Unio (Nodularia) douglasiae var. schrenki [sic] Westerlund, 1897 = Nodularia douglasiae
+ Unio abbreviatus Westerlund, 1897

Unio (s.s.) ater Nilsson, 1822 = Unio crassus

Unio (s.s.) crassus Retzius, 1788 = Unio crassus

Unio (s.s.) crassus consentancus [sic] ‘Ziegler’ Rossmässler, 1836 = Unio nanus

Unio (s.s.) crassus kungurensis Kobelt, 1912 = Unio crassus

Unio (s.s.) crassus var. chlebnikovi [sic] Kobelt, 1912 = Unio crassus
+ Unio kungurensis sylwensis Kobelt, 1912
+ Unio kungurensis irenjensis Kobelt, 1912

Unio (s.s.) crassus var. ishmensis Zhadin, 1938 = Unio crassus

Unio (s.s.) crassus var. okae Kobelt, 1911 = Unio crassus

Unio (s.s.) crassus var. pseudolitoralis Clessin, 1875 = Unio crassus

Unio (s.s.) crassus var. roseni Kobelt, 1912 = Unio crassus

Unio (s.s.) crassus var. stepanovi [sic] Drouet, 1881 = Unio gontierii

Unio (s.s.) hueti Bourguignat, 1855 = Unio hueti

Unio (s.s.) lindholmi Zhadin, 1938 = Unio pictorum

Unio (s.s.) mingrelicus Drouet, 1881 = Unio gontierii

Unio (s.s.) mingrelicus var. gregorii Kobelt, 1912 = Unio gontierii

Unio (s.s.) mingrelicus var. stevenianiformis Zhadin, 1938 = Unio gontierii

Unio (s.s.) pictorum (Linnaeus, 1758) = Unio pictorum

Unio (s.s.) pictorum defectivus Zhadin, 1938 = Unio pictorum

Unio (s.s.) pictorum gentilis Haas in Hesse, 1911 = Unio pictorum

Unio (s.s.) pictorum var. annulatus Kobelt, 1912 = Unio pictorum

Unio (s.s.) pictorum var. okensis Zhadin, 1938 = Unio pictorum
+ Unio pictorum var. okae Kobelt & Haas, 1910

Unio (s.s.) pictorum var. platyrhynchus Rossmässler, 1835 = Unio pictorum

Unio (s.s.) pictorum var. ponderosus Spitzi in Rossmässler, 1838 = Unio pictorum

Unio (s.s.) pictorum var. pygmaeus Zhadin, 1938 = Unio pictorum

Unio (s.s.) pictorum var. schrenkianus Clessin, 1880 = Unio pictorum

Unio (s.s.) sieversi Drouet, 1881 = Unio gontierii

Unio (s.s.) sieversi var. araxenus Drouet, 1881 = Unio mardinensis

Unio (s.s.) sieversi var. colchicus Drouet, 1881 = Unio gontierii

Unio (s.s.) sieversi var. kutaisiana [sic] Kobelt, 1886 = Unio gontierii

Unio (s.s.) sieversi var. raddei Drouet, 1881 = Unio gontierii

Unio (s.s.) stevenianus Drouet, 1881 = Unio gontierii

Unio (s.s.) tumidus Retzius, 1788 = Unio tumidus

Unio (s.s.) tumidus limicola Mörch, 1864 = Unio tumidus

Unio (s.s.) tumidus var. bashkiricus Zhadin, 1938 = Unio tumidus

Unio (s.s.) tumidus var. borysthenicus Kobelt, 1879 = Unio tumidus

Unio (s.s.) tumidus var. conus Spengler, 1793 = Unio tumidus

Unio (s.s.) tumidus var. fridmani Zhadin, 1938 = Unio tumidus

Unio (s.s.) tumidus var. gerstfeldtianus Clessin, 1880 = Unio tumidus

Unio (s.s.) tumidus var. ilikensis [sic] Kobelt, 1912 = Unio tumidus

Unio (s.s.) tumidus var. kobeltianus Zhadin, 1938 = Unio tumidus
+ Unio tumidus var. okae Kobelt & Haas, 1910

Unio (s.s.) tumidus var. moltshanovi Zhadin, 1938 = Unio tumidus