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Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio pictorum (Linnaeus, 1758): specimens
     100 available nominal species synonyms (158 total synonyms)

Recent Taxonomic Opinions

Unio pictorum — Graf (2007), Graf & Cummings (2007), Taylor et al. (2007), Manendo et al. (2008), Dhora (2009), ... Wu et al. (2024), Dai et al. (2024), Hou et al. (2024), Dai et al. (2025), Chen et al. (2025)
Unio pictorum pictorum — Kantor et al. (2010), Fauna Europaea (2012), Lopes-Lima et al. (2017), Bogatov (2022)
Unio rostratus rostratus — Kantor et al. (2010)
Unio limosus limosus — Kantor et al. (2010)
Unio limosus graniger — Kantor et al. (2010)
Unio pictorum ponderosus — Kantor et al. (2010)
Unio rostratus gentilis — Kantor et al. (2010)
Unio annulatus — Kantor et al. (2010)
Unio protractus — Kantor et al. (2010)
Unio limosus behningi — Kantor et al. (2010)
Unio pictorum rostratus — Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio pictorum platyrhynchus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio pictorum platyrhynchoideus [sic] — Fauna Europaea (2012)
Unio pictorum latirostris — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio pictorum praeposterus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio pictorum proechistus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio pictorum schrenkianus — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio pictorum gaudioni — Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio elongatulus bourgeticus — Fauna Europaea (2012)
Unio mancus bourgeticus — Prie (2012)
Unio pictorum ascanius — Fauna Europaea (2012)
Unio pictorum deshayesii — Prié & Puillandre (2014)
Unio pictorum platyrhinchoideus [sic] — Prié & Puillandre (2014)
Unio cf. pictorum — Nishihara et al. (2016)
Unio (s.s.) pictorum — Vinarski & Kantor (2016)
Unio (s.s.) rostrata — Vinarski & Kantor (2016)
Unio (s.s.) limosus — Vinarski & Kantor (2016)
Unio (s.s.) protractus — Vinarski & Kantor (2016)
— Vinarski & Kantor (2016)
Unio longirostris — Bogatov & Prozorova (2017)
Unio pictorum platyrinchoideus — Lopes-Lima et al. (2017)
Unio tumidus — Bogatov et al. (2018), Anisimova et al. (2018)
Unio elongatulus — Marrone et al. (2019)
Unio pictorum limosus — Bogatov (2022)
Unio tumidus longirostris — Bogatov (2022)
Unio pictorum protractus — Bogatov (2022)

Range

Widespread in Europe, from the Atlantic to the Volga and Black Sea. Introduced to the Amur Basin, eastern Asia.

Map of Georeferenced Specimens

INHS_21547. Gallenstein, France

INHS_19857. Lubrzanka Creek, 6 km S Kielce, Świętokrzyskie Voivodeship, Poland

INHS_19858. Lubrzanka Creek, 6 km S Kielce, Świętokrzyskie Voivodeship, Poland

INHS_78822. Nidda River, Frankfurt; L3005 bridge, Darmstadt, Hesse, Germany

INHS_21427. Nieuwe Merwede, S of Dordrecht, "Kop van Land", South Holland, Netherlands

INHS_21548. Oude Maas, Rhoon, Rotterdam Municipality, South Holland, Netherlands

INHS_21666. Schleswig- Holstein, Germany

INHS_21554. Uleser, Hungary

INHS_21668. Varenholz/ Lippe, North Rhine-Westphalia, Germany

INHS_21430. Waal River, W of Haaften, Gelderland, Netherlands

Austria

INHS_18534. Donaukanal (Danube Canal), Vienna, between Salztor bridge & Augarten bridge, Vienna, Austria

England

INHS_23641. New Bedford River, Sutton Gault, 11 km from Ely, Cambridgeshire, East of England, England

France

MNHN_22144. l’Aure, à Bayeux (Calvados)

MNHN_22456. Auxonne (Côte-d’Or)

INHS_21546. Côney River, Selles Dept., Haut Saone, Franche-Comté, France

MNHN_22455. fort de la Vitriolerie, à Lyon

MNHN_22309. fossés du fort de la Vitriolerie, à Lyon

MNHN_22210. La Bonde-Gendret près Troyes (Aude)

MNHN_22381. La Canne, à Pontillard (Nièvre)

MNHN_22488. La Drée (Saône-et-Loire)

MNHN_22211. La Fère (Aisne)

MNHN_22236. La Grosne, à la Ferté (Saône-et-Loire)

MNHN_22487. La Grosne, à Marnay (Saône-et-Loire)

MNHN_22150. La Loire à Balbigny (Loire)

MNHN_22149. La Loire, à Balbigny (Loire)

MNHN_22161. La Loire, à Villerest (Loire)

MNHN_22491. La Marne à Chelles (Seine-et-Marne)

MNHN_22490. La Marne à Lagny (Seine-et-Marne)

MNHN_22489. La Saône (Saône-et-Loire)

MNHN_22324. La Saône à Auxonne

MNHN_22540. La Saône à Neuville-sur-Saône (Rhône)

MNHN_22323. La Saône à Saint-Jean-de-Losne

MNHN_22320. La Saône à Tournus

MNHN_22235. La Saône, à Asnière

MNHN_22325. la Saône, à Gray (Haute-Saône)

MNHN_22539. la Saône, à Saint-Germain-au-Mont-d’Or

MNHN_22322. La Seine à Rouen (Seine-Inférieure)

MNHN_22212. le canal du Midi, Villefranche

MNHN_22160. le canal du Nivernais, à Bazolles

MNHN_22319. Les fossés du fort de la Vitriolerie, à Lyon

MNHN_22486. Vers (Jura)

MNHN_22151. Vienne (Isère)

Germany

Ukraine

INHS_22461. 12 km S Novovolynsk, Volyn Region, Volyn Oblast, Ukraine

Georeferenced Genbank Vouchers

France

MNHN_IM-2009-22390. (16S: KC703606, 28S: KC703855, COI: KC703954, go to Genbank) France: Ain River.

MNHN_IM-2009-17757. (16S: KC703453, 28S: KC703698, go to Genbank) France: Aisne River.

MNHN_IM-2009-17758. (16S: KC703454, 28S: KC703699, go to Genbank) ditto.

MNHN_IM-2009-17764. (16S: KC703459, 28S: KC703704, COI: JX046627, go to Genbank) ditto.

MNHN_IM-2009-17798. (16S: KC703478, 28S: KC703723, COI: JX046634, go to Genbank) ditto.

MNHN_IM-2009-17799. (16S: KC703479, 28S: KC703724, COI: JX046635, go to Genbank) ditto.

MNHN_IM-2009-17801. (16S: KC703481, 28S: KC703726, COI: JX046636, go to Genbank) ditto.

MNHN_IM-2009-17802. (16S: KC703482, 28S: KC703727, COI: JX046637, go to Genbank) ditto.

MNHN_IM-2009-21250. (16S: KC703554, 28S: KC703803, COI: JX046674, go to Genbank) France: Armance River.

MNHN_IM-2009-21212. (16S: KC703522, 28S: KC703770, go to Genbank) France: Canal Bruche Eckbolsheim.

MNHN_IM-2009-21297. (16S: KC703591, 28S: KC703840, COI: JX046695, go to Genbank) France: Cher River.

MNHN_IM-2009-21301. (16S: KC703595, 28S: KC703844, COI: JX046699, go to Genbank) ditto.

MNHN_IM-2009-12722. (16S: KC703436, 28S: KC703681, go to Genbank) France: Essonne River.

MNHN_IM-2009-21285. (16S: KC703584, 28S: KC703833, COI: JX046688, go to Genbank) France: Eure River.

MNHN_IM-2009-21286. (16S: KC703585, 28S: KC703834, COI: JX046689, go to Genbank) ditto.

MNHN_IM-2009-21287. (16S: KC703586, 28S: KC703835, COI: JX046690, go to Genbank) ditto.

MNHN_IM-2009-12600. (16S: KC703372, 28S: KC703636, go to Genbank) France: Garonne River.

MNHN_IM-2009-21214. (16S: KC703524, 28S: KC703772, COI: JX046659, go to Genbank) France: Gave de Pau River.

MNHN_IM-2009-21215. (16S: KC703525, 28S: KC703773, COI: JX046660, go to Genbank) ditto.

MNHN_IM-2009-21216. (16S: KC703526, 28S: KC703774, COI: JX046661, go to Genbank) ditto.

MNHN_IM-2009-21266. (16S: KC703568, 28S: KC703817, go to Genbank) France: Grand Melda River.

MNHN_IM-2009-12703. (16S: KC703419, 28S: KC703664, COI: JX046587, go to Genbank) France: Lake Bourget.

MNHN_IM-2009-12705. (16S: KC703421, 28S: KC703666, COI: JX046589, go to Genbank) ditto.

MNHN_IM-2009-17835. (16S: KC703501, 28S: KC703749, COI: JX046654, go to Genbank) France: Lake Brindos.

MNHN_IM-2009-17836. (16S: KC703502, 28S: KC703750, COI: JX046655, go to Genbank) ditto.

MNHN_IM-2009-17837. (16S: KC703503, 28S: KC703751, COI: JX046656, go to Genbank) ditto.

MNHN_IM-2009-17838. (16S: KC703504, 28S: KC703752, COI: JX046657, go to Genbank) ditto.

MNHN_IM-2009-17839. (16S: KC703505, 28S: KC703753, COI: JX046658, go to Genbank) ditto.

MNHN_IM-2009-12588. (16S: KC703366, 28S: KC703632, COI: JX046566, go to Genbank) France: Lary River.

MNHN_IM-2009-12589. (16S: KC703367, 28S: KC703633, COI: JX046567, go to Genbank) ditto.

MNHN_IM-2009-12573. (16S: KC703357, 28S: KC703626, COI: JX046558, go to Genbank) France: Meuse River.

MNHN_IM-2009-12574. (16S: KC703358, COI: JX046559, go to Genbank) ditto.

MNHN_IM-2009-12576. (16S: KC703360, 28S: KC703628, COI: JX046561, go to Genbank) ditto.

MNHN_IM-2009-12577. (16S: KC703361, COI: JX046562, go to Genbank) ditto.

MNHN_IM-2009-12685. (16S: KC703409, 28S: KC703654, go to Genbank) France: Oise River.

MNHN_IM-2009-12688. (16S: KC703410, 28S: KC703655, COI: JX046581, go to Genbank) ditto.

MNHN_IM-2009-21236. (16S: KC703542, 28S: KC703791, COI: JX046671, go to Genbank) France: Ourcq River.

MNHN_IM-2009-12691. (16S: KC703411, 28S: KC703656, go to Genbank) France: Petit Gland River.

MNHN_IM-2009-12693. (16S: KC703412, 28S: KC703657, go to Genbank) ditto.

MNHN_IM-2009-12695. (16S: KC703413, 28S: KC703658, COI: JX046582, go to Genbank) France: Seine River.

MNHN_IM-2009-12696. (16S: KC703414, 28S: KC703659, COI: JX046583, go to Genbank) ditto.

MNHN_IM-2009-17771. (16S: KC703463, 28S: KC703708, COI: JX046628, go to Genbank) France: Somme River.

MNHN_IM-2009-17772. (16S: KC703464, 28S: KC703709, COI: JX046629, go to Genbank) ditto.

MNHN_IM-2009-22396. (16S: KC703611, 28S: KC703860, COI: KC703959, go to Genbank) France: Vire River.

MNHN_IM-2009-22397. (16S: KC703612, 28S: KC703861, COI: KC703960, go to Genbank) ditto.

Germany

SMF_341808. (16S: OK055688, 28S: OK077671, COI: OK050296, go to Genbank) Braubach, Dörnigheim (50.1426, 8.8247).

SMF_354468. (16S: OK055690, 28S: OK077673, COI: OK050298, go to Genbank) Horloff, Oberflorstadt (50.3291, 8.871).

Moldova

RMBH_biv_776_1. (COI: MZ165267, go to Genbank) Moldova: Ryshkansky district, Prut River.

RMBH_biv_776_2. (COI: MZ165268, go to Genbank) ditto.

RMBH_biv_776_3. (COI: MZ165269, go to Genbank) ditto.

Russia

INREC-UP1. (COI: KY930361, go to Genbank) Kenon Lake, Russia.

INREC-UP11. (COI: KY930367, go to Genbank) ditto.

INREC-UP2. (COI: KY930362, go to Genbank) ditto.

INREC-UP2b. (COI: KY930363, go to Genbank) ditto.

INREC-UP5. (COI: KY930364, go to Genbank) ditto.

INREC-UP6. (COI: KY930365, go to Genbank) ditto.

INREC-UP6a. (COI: KY930368, go to Genbank) ditto.

INREC-UP9. (COI: KY930366, go to Genbank) ditto.

RMBH_biv_555. (COI: MZ165229, go to Genbank) Russia: Adygeya Region, Krasnodarskoe Reservoir.

RMBH_biv_592_1. (COI: MZ165240, go to Genbank) Russia: Astrakhan Region, Volga River.

RMBH_biv_584_2. (COI: MZ165237, go to Genbank) Russia: Bashkortostan Region, Belaya River.

RMBH_biv_588_2. (COI: MZ165239, go to Genbank) ditto.

Mp_4084_3HH. (16S: MK574387, 28S: MK574528, COI: MK034162, go to Genbank) Russia: European part, Arkhangelsk Region, Tsiglomenka River (64.4561°N, 40.5813°E), Northern Dvina Basin, White Sea drainage.

RMBH_biv_197_1. (16S: MK574389, 28S: MK574530, COI: MK574250, go to Genbank) Russia: European part, Novgorod Region, Volkhov River (58.5197°N, 31.2776°E), Ladoga Lake basin, Baltic Sea drainage.

RMBH_biv_197_2. (16S: MK574390, 28S: MK574531, COI: MK574251, go to Genbank) ditto.

RMBH_biv_196_7. (16S: MK574388, 28S: MK574529, COI: MK574249, go to Genbank) Russia: European part, Saratov Region, Khopyor River (51.5630° N, 43.1530° E), Don Basin, Azov Sea drainage.

RMBH_biv_282_1. (16S: MK574391, 28S: MK574532, COI: MK574252, go to Genbank) ditto.

RMBH_biv_728_1. (COI: MZ165248, go to Genbank) Russia: Krasnodar Region, Afips River (Kuban River basin).

RMBH_biv_728_2. (COI: MZ165249, go to Genbank) ditto.

RMBH_biv_728_3. (COI: MZ165250, go to Genbank) ditto.

RMBH_biv_728_4. (COI: MZ165251, go to Genbank) ditto.

RMBH_biv_728_5. (COI: MZ165252, go to Genbank) ditto.

RMBH_biv_725_2. (COI: MZ165245, go to Genbank) Russia: Krasnodar Region, Beysuzhek Levyi River.

RMBH_biv_725_3. (COI: MZ165246, go to Genbank) ditto.

RMBH_biv_556_2. (COI: MZ165231, go to Genbank) Russia: Krasnodar Region, Kirpili River.

Mp_3904_U11. (16S: MH544959, ITS1: MH544852, go to Genbank) Russia: River Ivitza, Volga basin, Rameshkovsky District, Tver Oblast.

Mp_3904_U17. (16S: MH544949, ITS1: MH544845, go to Genbank) ditto.

Mp_3904_U18. (16S: MH544943, 5.8S: MH544839, COI: MH539806, ITS1: MH544839, go to Genbank) ditto.

Mp_3904_U24. (16S: MH544950, ITS1: MH544844, go to Genbank) ditto.

Mp_3904_U25. (16S: MH544951, ITS1: MH544846, go to Genbank) ditto.

Mp_3904_U30. (16S: MH544944, COI: MH539805, ITS1: MH544840, go to Genbank) ditto.

Mp_3904_U37. (16S: MH544957, 5.8S: MH544848, ITS1: MH544848, go to Genbank) ditto.

Mp_3904_U40. (16S: MH544952, ITS1: MH544843, go to Genbank) ditto.

Mp_3904_U41. (16S: MH544946, 5.8S: MH544841, COI: MH539804, ITS1: MH544841, go to Genbank) ditto.

Mp_3904_U5. (16S: MH544945, COI: MH539807, ITS1: MH544838, go to Genbank) ditto.

Mp_3904_U62. (16S: MH544961, ITS1: MH544857, go to Genbank) ditto.

Mp_3904_U64. (16S: MH544962, ITS1: MH544856, go to Genbank) ditto.

Mp_3904_U7. (16S: MH544947, ITS1: MH544842, go to Genbank) ditto.

Mp_3904_U72. (16S: MH544958, 5.8S: MH544851, ITS1: MH544851, go to Genbank) ditto.

Mp_3904_U8. (16S: MH544948, ITS1: MH544847, go to Genbank) ditto.

RMBH_biv_562. (COI: ON648757, go to Genbank) Russia: Staryi Donets Lake, Don Basin (48.3406, 40.1096).

RMBH_biv_770. (COI: ON648760, go to Genbank) Russia: Tikhaya Sosna, Don Basin (50.8387, 39.0757).

RMBH_biv_1104_1. (COI: MZ165272, go to Genbank) Russia: Tyumen Region, Tura River.

RMBH_biv_1104_2. (COI: MZ165273, go to Genbank) ditto.

RMBH_biv_592_3. (COI: ON648759, go to Genbank) Russia: Volga (47.2532, 47.0418).

RMBH_biv_772_1. (COI: MZ165260, go to Genbank) Russia: Voronezh Region, Don River.

RMBH_biv_772_2. (COI: MZ165261, go to Genbank) ditto.

RMBH_biv_772_3. (COI: MZ165262, go to Genbank) ditto.

RMBH_biv_774_3. (COI: MZ165265, go to Genbank) Russia: Voronezh Region, Khoper River.

Sweden

NRM_42649-M064. (16S: DQ060163, COI: DQ060175, go to Genbank) Sweden.

Turkey

GCMAE_SS10_I_U1. (16S: MZ518924, 28S: MZ519161, COI: MZ511064, go to Genbank) Turkey: Atnos: Susurluk Basin (39.615322, 28.066988).

GCMAE_SS10_I_U2. (16S: MZ518925, 28S: MZ519162, COI: MZ511065, go to Genbank) ditto.

GCMAE_SS10_I_U3. (16S: MZ518926, 28S: MZ519163, COI: MZ511066, go to Genbank) ditto.

GCMAE_SS10_I_U4. (COI: MZ511067, go to Genbank) ditto.

GCMAE_SS10_I_U5. (COI: MZ511068, go to Genbank) ditto.

GCMAE_BK001. (16S: MZ518999, 28S: MZ519236, COI: MZ511112, go to Genbank) Turkey: Filyos: Filyos Basin (41.577354, 32.048551).

GCMAE_BK002. (COI: MZ511113, go to Genbank) ditto.

GCMAE_BK003. (16S: MZ519000, 28S: MZ519237, COI: MZ511114, go to Genbank) ditto.

GCMAE_BK004. (16S: MZ519001, 28S: MZ519238, COI: MZ511115, go to Genbank) ditto.

GCMAE_BK005. (COI: MZ511116, go to Genbank) ditto.

CIIMAR_BIV2174. (16S: MZ518881, 28S: MZ519118, COI: MZ511036, go to Genbank) Turkey: Gördes : Gediz Basin (38.669587, 27.979596).

CIIMAR_BIV2175. (16S: MZ518882, 28S: MZ519119, COI: MZ511037, go to Genbank) ditto.

CIIMAR_BIV2176. (16S: MZ518883, 28S: MZ519120, COI: MZ511038, go to Genbank) ditto.

CIIMAR_BIV2201. (16S: MZ518890, 28S: MZ519127, COI: MZ511042, go to Genbank) Turkey: Karamenderes: Karamenderes Basin (39.992947, 26.208696).

CIIMAR_BIV2202. (COI: MZ511043, go to Genbank) ditto.

GCMAE_M28_20. (16S: MZ518956, 28S: MZ519193, COI: MZ511076, go to Genbank) Turkey: Kuleli: Maritsa Basin (41.369225, 26.958031).

CIIMAR_BIV2260. (16S: MZ518897, 28S: MZ519134, COI: MZ511046, go to Genbank) Turkey: Kuş : Susurluk Basin (40.225219, 28.052855).

CIIMAR_BIV2261. (16S: MZ518898, 28S: MZ519135, COI: MZ511047, go to Genbank) ditto.

CIIMAR_BIV2262. (16S: MZ518899, 28S: MZ519136, COI: MZ511048, go to Genbank) ditto.

Mp_4217_BIV2258. (COI: MZ511044, go to Genbank) ditto.

Mp_4217_BIV2259. (COI: MZ511045, go to Genbank) ditto.

GCMAE_SS24_U1. (16S: MZ518936, 28S: MZ519173, COI: MZ511075, go to Genbank) Turkey: Madra: Susurluk Basin (39.728344, 27.674884).

GCMAE_M48/46_1. (16S: MZ518965, 28S: MZ519202, COI: MZ511077, go to Genbank) Turkey: Maritsa: Maritsa Basin (41.626782, 26.581009).

CIIMAR_BIV2317. (16S: MZ518906, 28S: MZ519143, COI: MZ511056, go to Genbank) Turkey: Sapanca inlet: Sapanca Basin (40.716928, 30.141743).

CIIMAR_BIV2318. (16S: MZ518907, 28S: MZ519144, COI: MZ511057, go to Genbank) ditto.

CIIMAR_BIV2319. (16S: MZ518908, 28S: MZ519145, COI: MZ511058, go to Genbank) ditto.

Mp_4217_BIV2315. (COI: MZ511054, go to Genbank) ditto.

Mp_4217_BIV2316. (COI: MZ511055, go to Genbank) ditto.

GCMAE_SS22_1. (COI: MZ511069, go to Genbank) Turkey: Simav: Susurluk Basin (39.917702, 28.166751).

GCMAE_SS22_2. (16S: MZ518927, 28S: MZ519164, COI: MZ511070, go to Genbank) ditto.

GCMAE_SS22_3. (16S: MZ518928, 28S: MZ519165, COI: MZ511071, go to Genbank) ditto.

GCMAE_SS22_4. (16S: MZ518929, 28S: MZ519166, COI: MZ511072, go to Genbank) ditto.

GCMAE_SS22_6. (COI: MZ511073, go to Genbank) ditto.

GCMAE_SS22_9. (COI: MZ511074, go to Genbank) ditto.

CIIMAR_BIV2296. (16S: MZ518903, 28S: MZ519140, COI: MZ511051, go to Genbank) Turkey: Uluabat : Susurluk Basin (40.136279, 28.674396).

CIIMAR_BIV2297. (16S: MZ518904, 28S: MZ519141, COI: MZ511052, go to Genbank) ditto.

CIIMAR_BIV2298. (16S: MZ518905, 28S: MZ519142, COI: MZ511053, go to Genbank) ditto.

Mp_4217_BIV2294. (COI: MZ511049, go to Genbank) ditto.

Mp_4217_BIV2295. (COI: MZ511050, go to Genbank) ditto.

Ukraine

ZSU-U1. (COI: KY930353, go to Genbank) Shoska River, Ukraine.

ZSU-U5. (COI: KY930355, go to Genbank) Snov River, Ukraine.

ZSU-U2. (COI: KY930354, go to Genbank) South Bug, Ukraine.

ZSU-U6. (COI: KY930356, go to Genbank) ditto.