MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Thu 19 March 2026
Bivalvia | ... | Unionidae | Unioninae | Unionini | Oxynaiina | Nodularia douglasiae
nominal species Nodularia denserugata Haas, 1910
Nodularia denserugata Haas, 1910 = Nodularia douglasiae
Haas, 1910. Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.
Taxonomic Opinions
+ Nodularia douglasiae — Wu et al., 2024: 13, Tables S1, S3.
= Ptychorhynchus denserugatum (Haas, 1910) — Graf & Cummings, 2021: 21.
= Ptychorhynchus denserugata (Haas, 1910) — Zieritz et al., 2018: online Table 1.
= Ptychorhynchus denserugata (Haas, 1910) — He & Zhuang, 2013: 94, fig. 223.
= Nodularia denserugata Haas, 1910 — Bogan, 2010: [23].
= Nodularia denserugata Haas, 1910 — Graf & Cummings, 2007: 304.
= Middendorffinaia (Sujfununio) denserugata (Haas, 1910) — Moskvicheva & Starobogatov, 1973: 30.
= Unio denserugatus (Haas, 1910) — Haas, 1969: 59.
= Nodularia (s.s.) denserugata Haas, 1910 — Simpson, 1914: 966.
= Nodularia (s.s.) denserugata Haas, 1910 — Haas, 1911: 83, pl. 8, figs. 3-4.
