MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Fri 13 February 2026
Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae | Pseudunio marocanus
nominal species Margaritana marocana Pallary, 1918
Margaritana marocana Pallary, 1918 = Pseudunio marocanus
Pallary, 1918. Bull. Soc. Hist. Nat. Afr. Nord., Alger 9: 117.
Pallary, 1920. unknown publication: 157, pl. 5, figs. 1-2.
Type(s): SYNTYPES BMNH_1937-12-30-13034-13035, l’oued Fès [Maroc].
Taxonomic Opinions
= Pseudunio marocanus (Pallary, 1918) — Dai et al., 2025: Table 2.
= Pseudunio marocanus (Pallary, 1918) — Chen et al., 2025: Table S3.
= Pseudunio marocanus — Chen et al., 2025: Table 1.
= Pseudunio marocanus (Pallary, 1918) — Dai et al., 2024: Table 3.
= Pseudunio marocana — Nakamura et al., 2024: 4.
= Pseudunio marocanus — Wu et al., 2023: Table 2.
= Pseudunio marocanus (Pallary, 1918) — Wu et al., 2022: Table S2.
= Pseudunio marocanus (Pallary, 1918) — Jeratthitikul et al., 2022: Table S1.
= Pseudunio marocanus (Pallary, 1918) — Graf & Cummings, 2021: 25.
= Pseudunio marocanus Pallary, 1918 — Príe et al., 2021: 2934, Table 1.
= Pseudunio marocanus — Zieritz et al., 2021: 1092.
= Pseudunio marocanus — Wu et al., 2021: attached Table 2.
= Pseudunio marocanus — Riccardi et al., 2020: 750.
= Margaritifera marocana — Fukata & Iigo, 2020: 1216.
= Pseudunio marocana — Wu et al., 2020: 195.
= Margaritifera marocana — Sano et al., 2020: 99.
= Margaritifera marocana (Pallary, 1918) — Huang et al., 2019: Table S4.
= Pseudunio marocanus — Froufe et al., 2019: 185.
= Pseudunio marocanus — Gomes-dos-Santos et al., 2019: 1418.
= Pseudunio marocanus (Pallary, 1918) — Lyubas et al., 2019: 8, Table 3.
= Pseudunio marocanus (Pallary, 1918) — Gomes-dos-Santos et al., 2019: 421.
= Pseudunio marocana (Pallary, 1918) — Vikhrev et al., 2019: 1, fig. 5, Table 3.
= Pseudunio marocanus — Guerra et al., 2019: Table S1.
= Margaritifera marocana — Vikhrev et al., 2018: 73, appendix 5.
= Margaritifera marocana — Huang et al., 2018: 302.
= Pseudunio marocanus — Lopes-Lima et al., 2018: 103, Table 3.
= Margaritifera marocana (Pallary, 1918) — Burzynski & Soroka, 2018: 3.
= Margaritifera marocana — Cha et al., 2018: 946.
= Margaritifera marocana (Pallary, 1918) — Bolotov et al., 2017: [16], Table S1.
= Margaritifera marocana (Pallary, 1918) — Bolotov et al., 2017: Table S1.
= Margaritifera marocana (Pallary, 1918) — Lopes-Lima et al., 2017: 249.
= Margaritifera marocana — Araujo et al., 2017: 280.
= Margaritifera (Pseudunio) marocana — Bolotov et al., 2016: 107, 111.
= Margaritifera marocana (Pallary, 1918) — Araujo et al., 2016: 290.
= Margaritifera marocana — Bolotov et al., 2015: 7.
= Margaritifera marocana — Takeuchi et al., 2015: 220.
= Margaritifera marocana — Inoue et al., 2013: 287.
= Margaritifera marocana (Pallary, 1918) — Graf & Cummings, 2011: 24.
= Margaritifera marocana (Pallary, 1918) — IUCN, 2010: [web].
= Margaritifera marocana Pallary, 1918 — Bogan, 2010: [20].
= Margaritifera marocana (Pallary, 1918) — Araujo et al., 2009: 95, fig. 1.
= Pseudunio auricularia marocana (Pallary, 1918) — Graf & Cummings, 2007: 95.
= Pseudunio marocana (Pallary, 1918) — D.G. Smith, 2001: 40, 42.
+ Margaritifera auricularia (Spengler, 1793) — Daget, 1998: 16.
+ Margaritifera auricularia (Spengler, 1793) — Ziuganov et al., 1994: 59.
+ Margaritifera auricularia (Spengler, 1793) — Mandahl-Barth, 1988: 35, 40.
= Margaritifera (Pseudunio) auricularia marocana (Pallary, 1918) — Van Damme, 1984: 52, fig. 70.
= Margaritifera (Pseudunio) auricularia marocana (Pallary, 1918) — Haas, 1969: 13.
= Margaritifera auricularia maroccana [sic] (Pallary, 1918) — Haas, 1940: 119.
