Serb, J.M., J.E. Buhay & C. Lydeard. 2003. Molecular systematics of the North American freshwater bivalve genus Quadrula (Unionidae: Ambleminae) based on mitochondrial ND1 sequences. Molecular Phylogenetics and Evolution 28: 1-11. Available electronically.
(click the figure number to see the cladogram)
+-- Lampsilis_siliquoidea
+--------------------+
| +-- Lampsilis_ornata
|
| +-------------------- Obliquaria_reflexa
| |
| +-------------------- Amblema_plicata
| |
| +-------------------- Reginaia_rotulata
| |
--+ +-------------------- Potamilus_alatus
| |
| | +-- Elliptoideus_sloatianus
| +-----------------+
| | +-- Plectomerus_dombeyanus
| |
| | +----- Medionidus_conradicus
| +--------------+
| | | +-- Dromus_dromas
+--+ +--+
| +-- Cyprogenia_aberti
|
| +-------- Hemistena_lata
+-----------+
| | +----- Elliptio_complanata
| +--+
| | +-- Fusconaia_burkei
| +--+
| +-- Fusconaia_flava
|
| +----------------- Megalonaias_nervosa
| |
| | +-------- Theliderma_cylindrica
| | +-----+
+--+ | | +----- Theliderma_intermedia
| | +--+
| | | +-- Theliderma_sparsa
| | +--+
| | +-- Theliderma_metanevra
+--+
| +----- Quadrula_verrucosa
| +-----+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_nobilis
+--+
| +-------- Pustulosa_kieneriana
| |
| +-------- Pustulosa_infucata
+--+
| +-- Pustulosa_pustulosa
| +--+
| | +-- Pustulosa_succissa
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina
+-- Lampsilis_siliquoidea
+-----------------------------------------+
| +-- Lampsilis_ornata
|
--+ +----------------------------------------- Potamilus_alatus
| |
| | +----- Medionidus_conradicus
+--+ +--------------------------------+
| | | +-- Dromus_dromas
| | +--+
| | +-- Cyprogenia_aberti
+--+
| +----------------------------------- Obliquaria_reflexa
| |
| | +-------------------------------- Amblema_plicata
+--+ |
| | +----- Reginaia_rotulata
| | +-----------------------+
+--+ | | +-- Elliptoideus_sloatianus
| | +--+
| | +-- Plectomerus_dombeyanus
| |
+--+ +-------- Hemistena_lata
| +-----------------+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Fusconaia_burkei
| | +--+
+--+ +-- Fusconaia_flava
|
| +----------------------- Megalonaias_nervosa
| |
| | +-- Quadrula_verrucosa
+--+ +-----------------+
| | +-- Quadrula_nobilis
| |
+--+ +----------------- Quadrula_quadrula
| |
| | +-------- Theliderma_cylindrica
+--+ +-----+
| | | +----- Theliderma_sparsa
| | +--+
| | | +-- Theliderma_intermedia
+--+ +--+
| +-- Theliderma_metanevra
|
| +----------- Pustulosa_kieneriana
| |
+--+ +-------- Pustulosa_infucata
| |
+--+ +-- Pustulosa_succissa
| +--+
| | +-- Pustulosa_pustulosa
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina