Bivalvia | ... | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula quadrula

nominal species Obliquaria (Quadrula) quadrula Rafinesque, 1820

Taxonomic Opinions

= Quadrula quadrula (Rafinesque, 1820) — Campbell & Lydeard, 2026: 289.

= Quadrula quadrula — Xiang et al., 2025: Table 2.

= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2025: Tables 1-2.

= Quadrula quadrula (Rafinesque, 1820) — Freeman et al., 2025: 247, Table 2.

= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2025: Table S3.

= Quadrula quadrula (Rafinesque, 1820) — FMCS, 2025: [6].

= Quadrula quadrula (Rafinesque, 1820) — Cummings et al., 2025: 454, pl. 92 top, middle left & right, Tables 6, 16.

= Quadrula quadrula — Hou et al., 2025: Table S3.

= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2025: Table 1.

= Quadrula quadrula — Wu et al., 2024: 7, Table 2.

= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2024: Table 1.

= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2024: Table 3.

= Quadrula quadrula — Wu et al., 2024: Table S3.

= Quadrula quadrula (Rafinesque, 1820) — Keogh et al., 2024: 2, Table S1.

= Quadrula quadrula — Hou et al., 2024: Table 2.

= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2023: 157, Table 2.

= Quadrula quadrula (Rafinesque, 1820) — Neemuchwala et al., 2023: 2, fig. 1e, Table 1.

= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2023: 134, Table 1.

= Quadrula quadrula (Rafinesque, 1820) — Hopper et al., 2023.

= Quadrula quadrula — Hauck et al., 2023: fig. 7.

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2023: Tables 2-3.

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2022: Table S2.

= Quadrula quadrula (Rafinesque, 1820) — Jeratthitikul et al., 2022: Table S1.

= Quadrula quadrula (Rafinesque, 1820) — Pavan-Kumar et al., 2022: Table S1.

= Quadrula quadrula — Mejia-Trujillo & Smith, 2022: 1856.

= Quadrula quadrula (Rafinesque, 1820) — Hnytka et al., 2022: 50, figs. 41-42.

= Quadrula quadrula (Rafinesque) — Hata et al., 2022: 2636.

= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2021: 20.

= Quadrula quadrula — Porto-Hannes et al., 2021: Table S2.

= Quadrula quadrula — Zieritz et al., 2021: 1091.

= Quadrula quadrula (Rafinesque, 1820) — FMCS, 2021: [6].

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2021: attached Tables 1-2.

= Quadrula quadrula — Lopes-Lima et al., 2020: Table 1.

= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2020: S22.

= Quadrula quadrula — Fukata & Iigo, 2020: 1216.

= Quadrula quadrula — Wu et al., 2020: 196.

= Quadrula quadrula — Inoue et al., 2020: 460.

= Quadrula quadrula — Smith et al., 2020: 507.

= Quadrula quadrula (Rafinesque, 1820) — Smith et al., 2020: 64.

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2020: 54.

= Quadrula quadrula — Chen et al., 2020: 2800.

= Quadrula quadrula — Teiga-Teixeira et al., 2020: 2978.

= Quadrula quadrula — Gasparini et al., 2020: Table S1.

= Quadrula quadrula — Gasparini et al., 2020: Table S1.

= Quadrula quadrula (Rafinesque, 1820) — Huang et al., 2019: Table S4.

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2019: 29.

= Quadrula quadrula (Rafinesque, 1820) — Lopes-Lima et al., 2019: [18], Table 7.

= Quadrula quadrula — Froufe et al., 2019: 185.

= Quadrula quadrula — Smith et al., 2019: 337, fig. 2, Table S1.

= Quadrula quadrula — Guerra et al., 2019: Table S1.

= Quadrula quadrula — Soroka & Burzynski, 2018: 242.

= Quadrula quadrula (Rafinesque, 1820) — Mathias et al., 2018: 207.

= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2018: Table S1.

= Quadrula quadrula — Huang et al., 2018: 304.

= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2018: 7.

= Quadrula quadrula — Johnson et al., 2018: 4.

= Quadrula quadrula — Wang et al., 2018: 611.

= Quadrula quadrula — Chase et al., 2018: 108.

= Quadrula quadrula (Rafinesque, 1820) — Burzynski & Soroka, 2018: 3.

= Quadrula quadrula — Aunins et al., 2018: 1125.

= Quadrula quadrula — Wang et al., 2018: 140.

= Quadrula quadrula — Marshall et al., 2018: 54.

= Quadrula quadrula — Cha et al., 2018: 946.

= Quadrula quadrula — Wu et al., 2018: 732.

= Quadrula quadrula (Rafinesque, 1820) — Lopes-Lima et al., 2017: 178.

= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2017: [16], Table S1.

= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2017: Table S1.

= Quadrula quadrula — Guerra et al., 2017: 8.

= Quadrula quadrula — Burzynski et al., 2017: 606.

= Quadrula quadrula (Rafinesque, 1820) — Williams et al., 2017: 43.

= Quadrula quadrula — Zhou et al., 2017: 235.

= Quadrula quadrula (Rafinesque, 1820) — Amaro et al., 2017: 743.

= Quadrula quadrula — Lopes-Lima et al., 2017: 250.

= Quadrula quadrula — Wen et al., 2017: 9.

= Quadrula quadrula (Rafinesque, 1820) — Amaro et al., 2017: 743.

= Quadrula quadrula — Wei et al., 2016: 1.

= Quadrula quadrula — Xue et al., 2016: 2699.

= Quadrula quadrula — Zhou et al., 2016: 4616.

= Quadrula quadrula — Wu et al., 2016: 4521.

= Quadrula quadrula — Mitchell et al., 2016: 5.

= Quadrula quadrula — Fonseca et al., 2016: 955.

= Quadrula quadrula — Feng et al., 2016: 445.

= Quadrula quadrula (Rafinesque, 1820) — Pfeiffer & Graf, 2015: 310.

= Quadrula quadrula — Froufe et al., 2015: 3572.

= Quadrula quadrula — Kongim et al., 2015: 103.

= Quadrula quadrula — Zieritz et al., 2015: 82.

= Quadrula quadrula (Rafinesque, 1820) — Paterson et al., 2015: 1123.

= Quadrula quadrula — Inoue et al., 2014: 540.

= Quadrula quadrula — Soroka & Burzynski, 2014: 1679.

= Q. quadrula — Sheng et al., 2014: 324.

= Quadrula quadrula — Inoue et al., 2013: 6.

= Quadrula quadrula (Rafinesque, 1820) — Pfeiffer & Graf, 2013: 251.

= Quadrula quadrula — Reis et al., 2013: 22.

= Quadrula quadrula — Huang et al., 2013: 5.

= Quadrula quadrula — Reis et al., 2013: 22.

= Quadrula quadrula (Rafinesque, 1820) — Campbell & Lydeard, 2012: 15.

= Quadrula quadrula — Campbell & Lydeard, 2012: 23.

= Quadrula quadrula quadrula (Rafinesque, 1820) — ITIS, 2012: [web].

= Quadrula quadrula (Rafinesque, 1820) — Szumowski et al., 2012: 72.

= Quadrula quadrula — Haag, 2012: 69.

= Quadrula quadrula (Rafinesque, 1820) — McMurray et al., 2012: 79.

= Quadrula quadrula — Chen et al., 2012: 2420.

= Quadrula quadrula — Boyer et al., 2011: 186.

= Quadrula quadrula — Breton et al., 2011: 1653.

= Quadrula quadrula (Rafinesque, 1820) — Whelan et al., 2011: 506.

= Quadrula quadrula — Plazzi et al., 2011: Table S5.

= Quadrula quadrula Rafinesque, 1820 — Bogan, 2010: [web].

= Quadrula quadrula — Soroka & Burzynski, 2010: 470.

= Quadrula quadrula — Soroka, 2010: 191.

= Quadrula quadrula — Cummings & Graf, 2010: 376, 377.

= Quadrula quadrula — Soroka, 2010: 156.

= Quadrula quadrula — Breton et al., 2009: 1576.

= Quadrula quadrula (Rafinesque, 1820) — Watters et al., 2009: 284, figs. 185-186.

= Quadrula quadrula — Jadhav & Jamkhedkar, 2009: 43.

= Quadrula quadrula — Hoeh et al., 2009: 310.

= Quadrula quadrula (Rafinesque, 1820) — Williams et al., 2008: 678.

= Quadrula quadrula — Chapman et al., 2008: 11.

= Quadrula quadrula (Rafinesque, 1820) — Chapman et al., 2008: 306.

= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2007: 308.

= Quadrula quadrula — Zhou et al., 2007: 1029.

= Quadrula quadrula (Rafinesque, 1820) — Tiemann et al., 2007: 111.

= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2006: 349.

= Quadrula quadrula (Rafinesque, 1820) — Hoke, 2005: 30, 32, Tables 1-2.

= Quadrula quadrula (Rafinesque) — Jones et al., 2005: 91, Appendix 1.

= Quadrula quadrula — Curole & Kocher, 2005: 384.

= Quadrula quadrula (Rafinesque, 1820) — Campbell et al., 2005: 155.

= Quadrula quadrula — Krebs, 2004: 1703.

= Quadrula quadrula — Serb et al., 2003: 10.

= Quadrula quadrula — Krebs et al., 2003: 311.

= Quadrula quadrula (Rafinesque, 1820) — Roe & Hoeh, 2003: 110.

= Quadrula quadrula (Rafinesque, 1820) — Cicerello & Schuster, 2003: 48, text fig..

= Quadrula quadrula — Sietman, 2003: 60, text fig..

= Quadrula quadrula — Serb et al., 2003: 10.

= Quadrula quadrula (Raf., 1820) — Graf, 2002: 191, Table 3.

= Quadrula quadrula — Curole & Kocher, 2002: 1324.

= Quadrula quadrula (Rafinesque, 1820) — Hoeh et al., 2002: 127.

= Quadrula quadrula (Rafinesque, 1820) — Hoeh et al., 2001: 260.

= Quadrula quadrula (Raf., 1820) — Graf & Ó Foighil, 2000: 160.

= Quadrula quadrula — Turner et al., 2000: 781.

= Quadrula quadrula (Rafinesque) — Lydeard et al., 2000: 148.

= Quadrula quadrula (Rafinesque, 1820) — Bogan & Hoeh, 2000: 162.

= Quadrula quedrula [sic] (Rafinesque, 1820) — Contreras-Arquieta, 2000: 44.

= Quadrula quadrula (Rafinesque) — Lydeard et al., 2000: 148.

= Quadrula quadrula (Rafinesque) — Lydeard et al., 1999: 130.

= Quadrula quadrula (Rafinesque, 1820) — Parmalee & Bogan, 1998: 220, pl. 108.

= Quadrula quadrula (Rafinesque, 1820) — Turgeon et al., 1998: 37.

= Quadrula quadrula (Rafinesque) — Johnson et al., 1998: 322.

= Quadrula quadrula — Berg et al., 1998: 1365.

= Quadrula quadrula Rafinesque — Strayer & Jirka, 1997: 85, pl. 27, fig. 8.

= Quadrula quadrula — Rosenberg et al., 1997: 303.

= Quadrula quadrula (Rafinesque, 1820) — Cummings & Mayer, 1997: 137.

= Quadrula quadrula (Rafinesque, 1820) — Howells et al., 1996: 124, text fig..

= Quadrula quadrula (Rafinesque, 1820) — IUCN, 1996: [web].

= Quadrula quadrula — Lydeard et al., 1996: 1595.

= Quadrula quadrula — Rosenberg et al., 1994: 112.

= Quadrula quadrula (Rafinesque) — Williams et al., 1993: 14.

= Quadrula quadrula (Rafinesque, 1820) — Vidrine, 1993: 53, pl. 6, figs. I-Q.

= Quadrula quadrula (Rafinesque, 1820) — Cummings & Mayer, 1992: 30, text fig..

= Quadrula quadrula (Rafinesque, 1820) — Turgeon et al., 1988: 33.

= Quadrula quadrula (Rafinesque, 1820) — Oesch, 1984: 85, text figs..

= Quadrula quadrula (Rafinesque) — Davis, 1984: 631.

= Quadrula quadrula (Raf.) — Davis, 1983: 197, fig. 1H.

= Quadrula quadrula (Raf.) — Davis & Fuller, 1981: 219.

= Quadrula quadrula (Rafinesque, 1820) — R.I. Johnson, 1980: 90, List 1, sp. 11; Tables 2-5.

= Quadrula quadrula (Rafinesque) — Mackie et al., 1980: 7.

= Quadrula quadrula — Gordon et al., 1979: 34, Table 3.

= Quadrula quadrula (Raf.) — Mathiak, 1979: 48, pl. 7-C.

= Quadrula quadrula (Rafinesque, 1820) — Habe, 1977: 119.

= Quadrula quadrula (Rafinesque, 1820) — Burch, 1975: 10.

= Obliquaria quadrula Rafinesque, 1820 — Burch, 1975: 171.

= Quadrula quadrula (Rafinesque, 1820) — Clarke, 1973: 32, pl. 1, figs. 3-4.

= Quadrula (s.s.) quadrula quadrula (Rafinesque, 1820) — Haas, 1969: 289.

= Quadrula (s.s.) quadrula (Rafinesque, 1820) — Haas, 1969: N437, fig. D34:1.

= Quadrula quadrula (Rafinesque, 1820) — Murray & Leonard, 1962: 50, pl. 8, figs. 1-4.

= Quadrula quadrula (Rafinesque, 1820) — Modell, 1942: 187.

= Quadrula quadrula (Rafinesque, 1820) — Thiele, 1934: 826, [1260].

= Quadrula (s.s.) gigantea [sic] (Raf.) — Haas, 1930: 327.

= Quadrula (s.s.) quadrula (Rafinesque, 1820) — F.C. Baker, 1928: 84, pl. 46, fig. 4; pl. 48, figs. 2-4.

= Quadrula (s.s.) quadrula (Rafinesque, 1820) — Frierson, 1927: 47, sp. 207.

= Quadrula quadrula (Rafinesque) — Ortmann, 1925: 334.

= Quadrula quadrula (Rafinesque) — Ortmann, 1924: 16.

= Quadrula quadrula (Rafinesque, 1820) — Ortmann & Walker, 1922: 16.

= Quadrula quadrula Rafinesque — Utterback, 1915: 135, pl. 9, fig. 19; pl. 18, figs. 45a-f.

= Quadrula quadrula Raf. — Vanatta, 1915: 556.

+ — Simpson, 1914: 731.

+ — de Gregorio, 1914: 37.

+ — Simpson, 1900: 744.

= Unio (sec. Quadrula) quadrulus (Rafinesque, 1820) — Paetel, 1890: 165.

+ — Lea, 1870: 92.

= Unio quadratus [sic] (Rafinesque, 1820) — Reeve, 1864: pl. 6, sp. 24.

= Unio quadrulus Rafinesque — Küster, 1861: 184, pl. 58, fig. 1.

= Unio (Theliderma) quadrulus Raf. — Conrad, 1853: 256, 269.

+ — Lea, 1852: 58.

= Unio quadrulus (Rafinesque, 1820) — Conrad, 1840: 112, pl. 62.

+ — Lea, 1836: 59.

= Unio quadrulus Raf. — Ferussac, 1835: 27.

= Obliquaria (Quadrula) quadrula — Rafinesque, 1820: 307, sp. 35.