MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Wed 04 February 2026
Bivalvia | ... | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula quadrula
nominal species Obliquaria (Quadrula) quadrula Rafinesque, 1820
Obliquaria (Quadrula) quadrula Rafinesque, 1820 = Quadrula quadrula
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 307 [also as ‘Unio quadrula’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 51.
Type(s): NEOTYPE ANSP_20224 (by Johnson & Baker, 1973: 167), Salt River.
Taxonomic Opinions
= Quadrula quadrula — Xiang et al., 2025
= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2025
= Quadrula quadrula (Rafinesque, 1820) — Freeman et al., 2025
= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2025
= Quadrula quadrula (Rafinesque, 1820) — FMCS, 2025
= Quadrula quadrula (Rafinesque, 1820) — Cummings et al., 2025
= Quadrula quadrula — Hou et al., 2025
= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2025
= Quadrula quadrula — Wu et al., 2024
= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2024
= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2024
= Quadrula quadrula — Wu et al., 2024
= Quadrula quadrula (Rafinesque, 1820) — Keogh et al., 2024
= Quadrula quadrula — Hou et al., 2024
= Quadrula quadrula (Rafinesque, 1820) — Dai et al., 2023
= Quadrula quadrula (Rafinesque, 1820) — Neemuchwala et al., 2023
= Quadrula quadrula (Rafinesque, 1820) — Chen et al., 2023
= Quadrula quadrula (Rafinesque, 1820) — Hopper et al., 2023
= Quadrula quadrula — Hauck et al., 2023
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2023
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2022
= Quadrula quadrula (Rafinesque, 1820) — Jeratthitikul et al., 2022
= Quadrula quadrula (Rafinesque, 1820) — Pavan-Kumar et al., 2022
= Quadrula quadrula — Mejia-Trujillo & Smith, 2022
= Quadrula quadrula (Rafinesque, 1820) — Hnytka et al., 2022
= Quadrula quadrula (Rafinesque) — Hata et al., 2022
= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2021
= Quadrula quadrula — Porto-Hannes et al., 2021
= Quadrula quadrula — Zieritz et al., 2021
= Quadrula quadrula (Rafinesque, 1820) — FMCS, 2021
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2021
= Quadrula quadrula — Lopes-Lima et al., 2020
= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2020
= Quadrula quadrula — Fukata & Iigo, 2020
= Quadrula quadrula — Wu et al., 2020
= Quadrula quadrula — Inoue et al., 2020
= Quadrula quadrula — Smith et al., 2020
= Quadrula quadrula (Rafinesque, 1820) — Smith et al., 2020
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2020
= Quadrula quadrula — Chen et al., 2020
= Quadrula quadrula — Teiga-Teixeira et al., 2020
= Quadrula quadrula — Gasparini et al., 2020
= Quadrula quadrula — Gasparini et al., 2020
= Quadrula quadrula (Rafinesque, 1820) — Huang et al., 2019
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2019
= Quadrula quadrula (Rafinesque, 1820) — Lopes-Lima et al., 2019
= Quadrula quadrula — Froufe et al., 2019
= Quadrula quadrula — Smith et al., 2019
= Quadrula quadrula — Guerra et al., 2019
= Quadrula quadrula — Soroka & Burzynski, 2018
= Quadrula quadrula (Rafinesque, 1820) — Mathias et al., 2018
= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2018
= Quadrula quadrula — Huang et al., 2018
= Quadrula quadrula (Rafinesque, 1820) — Wu et al., 2018
= Quadrula quadrula — Johnson et al., 2018
= Quadrula quadrula — Wang et al., 2018
= Quadrula quadrula — Chase et al., 2018
= Quadrula quadrula (Rafinesque, 1820) — Burzynski & Soroka, 2018
= Quadrula quadrula — Aunins et al., 2018
= Quadrula quadrula — Wang et al., 2018
= Quadrula quadrula — Marshall et al., 2018
= Quadrula quadrula — Cha et al., 2018
= Quadrula quadrula — Wu et al., 2018
= Quadrula quadrula (Rafinesque, 1820) — Lopes-Lima et al., 2017
= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2017
= Quadrula quadrula (Rafinesque, 1820) — Bolotov et al., 2017
= Quadrula quadrula — Guerra et al., 2017
= Quadrula quadrula — Burzynski et al., 2017
= Quadrula quadrula (Rafinesque, 1820) — Williams et al., 2017
= Quadrula quadrula — Zhou et al., 2017
= Quadrula quadrula (Rafinesque, 1820) — Amaro et al., 2017
= Quadrula quadrula — Lopes-Lima et al., 2017
= Quadrula quadrula — Wen et al., 2017
= Quadrula quadrula (Rafinesque, 1820) — Amaro et al., 2017
= Quadrula quadrula — Wei et al., 2016
= Quadrula quadrula — Xue et al., 2016
= Quadrula quadrula — Zhou et al., 2016
= Quadrula quadrula — Wu et al., 2016
= Quadrula quadrula — Mitchell et al., 2016
= Quadrula quadrula — Fonseca et al., 2016
= Quadrula quadrula — Feng et al., 2016
= Quadrula quadrula (Rafinesque, 1820) — Pfeiffer & Graf, 2015
= Quadrula quadrula — Froufe et al., 2015
= Quadrula quadrula — Kongim et al., 2015
= Quadrula quadrula — Zieritz et al., 2015
= Quadrula quadrula (Rafinesque, 1820) — Paterson et al., 2015
= Quadrula quadrula — Inoue et al., 2014
= Quadrula quadrula — Soroka & Burzynski, 2014
= Q. quadrula — Sheng et al., 2014
= Quadrula quadrula — Inoue et al., 2013
= Quadrula quadrula (Rafinesque, 1820) — Pfeiffer & Graf, 2013
= Quadrula quadrula — Reis et al., 2013
= Quadrula quadrula — Huang et al., 2013
= Quadrula quadrula — Reis et al., 2013
= Quadrula quadrula (Rafinesque, 1820) — Campbell & Lydeard, 2012
= Quadrula quadrula — Campbell & Lydeard, 2012
= Quadrula quadrula quadrula (Rafinesque, 1820) — ITIS, 2012
= Quadrula quadrula (Rafinesque, 1820) — Szumowski et al., 2012
= Quadrula quadrula — Haag, 2012
= Quadrula quadrula (Rafinesque, 1820) — McMurray et al., 2012
= Quadrula quadrula — Chen et al., 2012
= Quadrula quadrula — Boyer et al., 2011
= Quadrula quadrula — Breton et al., 2011
= Quadrula quadrula (Rafinesque, 1820) — Whelan et al., 2011
= Quadrula quadrula — Plazzi et al., 2011
= Quadrula quadrula Rafinesque, 1820 — Bogan, 2010
= Quadrula quadrula — Soroka & Burzynski, 2010
= Quadrula quadrula — Soroka, 2010
= Quadrula quadrula — Cummings & Graf, 2010
= Quadrula quadrula — Soroka, 2010
= Quadrula quadrula — Breton et al., 2009
= Quadrula quadrula (Rafinesque, 1820) — Watters et al., 2009
= Quadrula quadrula — Jadhav & Jamkhedkar, 2009
= Quadrula quadrula — Hoeh et al., 2009
= Quadrula quadrula (Rafinesque, 1820) — Williams et al., 2008
= Quadrula quadrula — Chapman et al., 2008
= Quadrula quadrula (Rafinesque, 1820) — Chapman et al., 2008
= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2007
= Quadrula quadrula — Zhou et al., 2007
= Quadrula quadrula (Rafinesque, 1820) — Tiemann et al., 2007
= Quadrula quadrula (Rafinesque, 1820) — Graf & Cummings, 2006
= Quadrula quadrula (Rafinesque, 1820) — Hoke, 2005
= Quadrula quadrula (Rafinesque) — Jones et al., 2005
= Quadrula quadrula — Curole & Kocher, 2005
= Quadrula quadrula (Rafinesque, 1820) — Campbell et al., 2005
= Quadrula quadrula — Krebs, 2004
= Quadrula quadrula — Serb et al., 2003
= Quadrula quadrula — Krebs et al., 2003
= Quadrula quadrula (Rafinesque, 1820) — Roe & Hoeh, 2003
= Quadrula quadrula (Rafinesque, 1820) — Cicerello & Schuster, 2003
= Quadrula quadrula — Sietman, 2003
= Quadrula quadrula — Serb et al., 2003
= Quadrula quadrula (Raf., 1820) — Graf, 2002
= Quadrula quadrula — Curole & Kocher, 2002
= Quadrula quadrula (Rafinesque, 1820) — Hoeh et al., 2002
= Quadrula quadrula (Rafinesque, 1820) — Hoeh et al., 2001
= Quadrula quadrula (Raf., 1820) — Graf & Ó Foighil, 2000
= Quadrula quadrula — Turner et al., 2000
= Quadrula quadrula (Rafinesque) — Lydeard et al., 2000
= Quadrula quadrula (Rafinesque, 1820) — Bogan & Hoeh, 2000
= Quadrula quedrula [sic] (Rafinesque, 1820) — Contreras-Arquieta, 2000
= Quadrula quadrula (Rafinesque) — Lydeard et al., 2000
= Quadrula quadrula (Rafinesque) — Lydeard et al., 1999
= Quadrula quadrula (Rafinesque, 1820) — Parmalee & Bogan, 1998
= Quadrula quadrula (Rafinesque, 1820) — Turgeon et al., 1998
= Quadrula quadrula (Rafinesque) — Johnson et al., 1998
= Quadrula quadrula — Berg et al., 1998
= Quadrula quadrula Rafinesque — Strayer & Jirka, 1997
= Quadrula quadrula — Rosenberg et al., 1997
= Quadrula quadrula (Rafinesque, 1820) — Cummings & Mayer, 1997
= Quadrula quadrula (Rafinesque, 1820) — Howells et al., 1996
= Quadrula quadrula (Rafinesque, 1820) — IUCN, 1996
= Quadrula quadrula — Lydeard et al., 1996
= Quadrula quadrula — Rosenberg et al., 1994
= Quadrula quadrula (Rafinesque) — Williams et al., 1993
= Quadrula quadrula (Rafinesque, 1820) — Vidrine, 1993
= Quadrula quadrula (Rafinesque, 1820) — Cummings & Mayer, 1992
= Quadrula quadrula (Rafinesque, 1820) — Turgeon et al., 1988
= Quadrula quadrula (Rafinesque, 1820) — Oesch, 1984
= Quadrula quadrula (Rafinesque) — Davis, 1984
= Quadrula quadrula (Raf.) — Davis, 1983
= Quadrula quadrula (Raf.) — Davis & Fuller, 1981
= Quadrula quadrula (Rafinesque, 1820) — R.I. Johnson, 1980
= Quadrula quadrula (Rafinesque) — Mackie et al., 1980
= Quadrula quadrula — Gordon et al., 1979
= Quadrula quadrula (Raf.) — Mathiak, 1979
= Quadrula quadrula (Rafinesque, 1820) — Habe, 1977
= Quadrula quadrula (Rafinesque, 1820) — Burch, 1975
= Obliquaria quadrula Rafinesque, 1820 — Burch, 1975
= Quadrula quadrula (Rafinesque, 1820) — Clarke, 1973
= Quadrula (s.s.) quadrula quadrula (Rafinesque, 1820) — Haas, 1969
= Quadrula (s.s.) quadrula (Rafinesque, 1820) — Haas, 1969
= Quadrula quadrula (Rafinesque, 1820) — Murray & Leonard, 1962
= Quadrula quadrula (Rafinesque, 1820) — Modell, 1942
= Quadrula quadrula (Rafinesque, 1820) — Thiele, 1934
= Quadrula (s.s.) gigantea [sic] (Raf.) — Haas, 1930
= Quadrula (s.s.) quadrula (Rafinesque, 1820) — F.C. Baker, 1928
= Quadrula (s.s.) quadrula (Rafinesque, 1820) — Frierson, 1927
= Quadrula quadrula (Rafinesque) — Ortmann, 1925
= Quadrula quadrula (Rafinesque) — Ortmann, 1924
= Quadrula quadrula (Rafinesque, 1820) — Ortmann & Walker, 1922
= Quadrula quadrula Rafinesque — Utterback, 1915
= Quadrula quadrula Raf. — Vanatta, 1915
+ — Simpson, 1914
+ — Simpson, 1900
= Unio (sec. Quadrula) quadrulus (Rafinesque, 1820) — Paetel, 1890
+ — Lea, 1870
= Unio quadratus [sic] (Rafinesque, 1820) — Reeve, 1864
= Unio quadrulus Rafinesque — Küster, 1861
= Unio (Theliderma) quadrulus Raf. — Conrad, 1853
+ — Lea, 1852
= Unio quadrulus (Rafinesque, 1820) — Conrad, 1840
+ — Lea, 1836
= Unio quadrulus Raf. — Ferussac, 1835
= Obliquaria (Quadrula) quadrula — Rafinesque, 1820
